468 lines
58 KiB
XML
468 lines
58 KiB
XML
<document id="0F0553E9526B25CD7E5326B9D8325489" ID-CLB-Dataset="7105" ID-DOI="10.12782/specdiv.26.7" ID-GBIF-Dataset="5dc29412-7908-479d-93e6-1ecc88fdc1fd" ID-ISSN="2189-7301" ID-Zenodo-Dep="4585295" ID-ZooBank="368822BA-78A5-44BC-9C15-2DCB77047D7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614947948901" checkinUser="felipe" docAuthor="Arai, Mikihito & Fujita, Toshihiko" docDate="2021" docId="A0103577FF8DFFB8FC09D2F27EA806E7" docLanguage="en" docName="SpeciesDiversity.26.1.7-21.pdf" docOrigin="Species Diversity 26 (1)" docSource="http://dx.doi.org/10.12782/specdiv.26.7" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Fromia labeosa Arai & Fujita 2021, sp. nov." docType="treatment" docUuid="71366E37-AEFD-4E10-B456-7DB14E8BAA00" docUuidSource="ZooBank" docVersion="6" lastPageNumber="15" masterDocId="5C294D0FFF8BFFB0FFE7D1017D2A0208" masterDocTitle="Sea Stars of Families Ophidiasteridae and Goniasteridae (Echinodermata: Asteroidea) from the Mesophotic Zone of the Ogasawara Islands, Including Two New Species" masterLastPageNumber="21" masterPageNumber="7" pageNumber="13" updateTime="1698932841729" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="47A5EB2E45219B3778923A19A8C3E728">
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<mods:title id="186D53BB69418DF2BA806C4260889949">Sea Stars of Families Ophidiasteridae and Goniasteridae (Echinodermata: Asteroidea) from the Mesophotic Zone of the Ogasawara Islands, Including Two New Species</mods:title>
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<mods:name id="E73A160F62232E1C9D316DC1CAF76D21" type="personal">
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<mods:roleTerm id="C4F936131EB3A6CC828D136B343F944E">Author</mods:roleTerm>
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<mods:namePart id="BF3D47C789DDF19E6DA8CBCB766C6148">Arai, Mikihito</mods:namePart>
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<mods:affiliation id="16432AE9DCA09C95CED57B2DCB73CEB3">Department of Biological Sciences, Graduate School of Science, the University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 8654, Japan E-mail: arai @ kahaku. go. jp & National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan & Corresponding author</mods:affiliation>
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<mods:nameIdentifier id="F28A2E950B85B9388F87ACFB2C1F2620" type="email">arai@kahaku.go.jp</mods:nameIdentifier>
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<mods:roleTerm id="177194F243A5451F5CB5F58F6B38A2F2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CEE227761FD22F7D501F621E500DCABC">Fujita, Toshihiko</mods:namePart>
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<mods:affiliation id="59599B8D1DA23F94E522396CAD9A75E8">Department of Biological Sciences, Graduate School of Science, the University of Tokyo, 7 - 3 - 1 Hongo, Bunkyo-ku, Tokyo 113 - 8654, Japan E-mail: arai @ kahaku. go. jp & National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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<mods:nameIdentifier id="ACFD76825A5F170308EE7648D5F82416" type="email">arai@kahaku.go.jp</mods:nameIdentifier>
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<mods:titleInfo id="519C51DB7D7FD6AE3272EFCFA8AA3DB8">
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<mods:title id="1B2970D7ED479EF8D0F887959BBF0296">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="75FC14E9C0169691927F9BB0CBB405A4">
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<mods:date id="12FBAF491088C4C616A1D9EF4A6BA51B">2021</mods:date>
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<mods:number id="6B97F168F352F41A6B53BD7CB86E3F92">2021-01-01</mods:number>
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<mods:number id="D4E68A758DB887B30181870088CF5F6F">26</mods:number>
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<mods:number id="A09315E748F2CD2650D5B5FA1DA84FDB">1</mods:number>
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<mods:start id="94641F2020CD29A54A66A2152BFC7E10">7</mods:start>
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<mods:url id="01E551B638EC11725CF3CA1151B464E2">http://dx.doi.org/10.12782/specdiv.26.7</mods:url>
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<mods:classification id="3FED198D888DC084C538B0717488384A">journal article</mods:classification>
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<treatment id="A0103577FF8DFFB8FC09D2F27EA806E7" ID-DOI="http://doi.org/10.5281/zenodo.4734688" ID-GBIF-Taxon="181851164" ID-Zenodo-Dep="4734688" LSID="urn:lsid:zoobank.org:act:71366E37-AEFD-4E10-B456-7DB14E8BAA00" httpUri="http://treatment.plazi.org/id/A0103577FF8DFFB8FC09D2F27EA806E7" lastPageId="8" lastPageNumber="15" pageId="6" pageNumber="13">
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<subSubSection id="60A3D7EAFF8DFFB6FC09D2F279F50603" box="[1006,1247,1011,1035]" pageId="6" pageNumber="13" type="nomenclature">
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<paragraph id="28068461FF8DFFB6FC09D2F279F50603" blockId="6.[1006,1247,1011,1035]" box="[1006,1247,1011,1035]" pageId="6" pageNumber="13">
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<heading id="734E330DFF8DFFB6FC09D2F279F50603" box="[1006,1247,1011,1035]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="13" reason="2">
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<taxonomicName id="EFB9FFE2FF8DFFB6FC09D2F279A70603" ID-CoL="848TZ" authority="Arai & Fujita, 2021" authorityName="Arai & Fujita" authorityYear="2021" box="[1006,1165,1011,1035]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="6" pageNumber="13" phylum="Echinodermata" rank="species" species="labeosa" status="sp. nov.">
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<emphasis id="1ACD5873FF8DFFB6FC09D2F279A70603" bold="true" box="[1006,1165,1011,1035]" italics="true" pageId="6" pageNumber="13">Fromia labeosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="01FEE508FF8DFFB6FB74D2F579F50603" box="[1171,1247,1012,1035]" pageId="6" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="60A3D7EAFF8DFFB6FC8FD515784E0624" box="[872,1380,1044,1068]" pageId="6" pageNumber="13" type="vernacular_names">
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<paragraph id="28068461FF8DFFB6FC8FD515784E0624" blockId="6.[872,1380,1044,1101]" box="[872,1380,1044,1068]" pageId="6" pageNumber="13">[New Japanese name: Ogasawara-juzuberi-hitode]</paragraph>
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</subSubSection>
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<subSubSection id="60A3D7EAFF8DFFB6FBCCD535798B0644" box="[1067,1185,1076,1101]" pageId="6" pageNumber="13" type="description">
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<paragraph id="28068461FF8DFFB6FBCCD535798B0644" blockId="6.[872,1380,1044,1101]" box="[1067,1185,1076,1101]" pageId="6" pageNumber="13">
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(
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<figureCitation id="B08298E4FF8DFFB6FBD5D53579A80645" box="[1074,1154,1076,1101]" captionStart="Fig" captionStartId="2.[822,853,1874,1896]" captionTargetBox="[188,1402,207,2036]" captionTargetId="figure-6@2.[187,1400,205,2029]" captionTargetPageId="2" captionText="Fig. 2. Abactinal view of the live specimens (A–D) and ethanol-preserved specimen (E). A, Bathyferdina caelator sp. nov., NSMT E-8265, holotype; B, Fromia eusticha, NSMT E-10595; C, Fromia labeosa sp. nov., NSMT E-9297, holotype; D, Ogmaster capella, NSMT E-9312; E, Tamaria tenella, NSMT E-9277. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.4585299" httpUri="https://zenodo.org/record/4585299/files/figure.png" pageId="6" pageNumber="13">Figs 2C</figureCitation>
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,
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<figureCitation id="B08298E4FF8DFFB6FB6BD53579B30644" box="[1164,1177,1076,1100]" captionStart="Fig" captionStartId="7.[157,188,1539,1561]" captionTargetBox="[188,1398,206,1499]" captionTargetId="figure-228@7.[187,1399,205,1500]" captionTargetPageId="7" captionText="Fig. 5. Fromia labeosa sp. nov., NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5mm (A, C), 1mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines." figureDoi="http://doi.org/10.5281/zenodo.4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="6" pageNumber="13">5</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="60A3D7EAFF8DFFB6FCD2D57779820685" box="[821,1192,1141,1166]" pageId="6" pageNumber="13" type="reference_group">
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<paragraph id="28068461FF8DFFB6FCD2D57779820685" blockId="6.[821,1192,1141,1166]" box="[821,1192,1141,1166]" pageId="6" pageNumber="13">
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<treatmentCitationGroup id="08A9A34FFF8DFFB6FCD2D57779820685" box="[821,1192,1141,1166]" pageId="6" pageNumber="13">
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<emphasis id="1ACD5873FF8DFFB6FCD2D5777E570685" box="[821,893,1142,1165]" italics="true" pageId="6" pageNumber="13">Fromia</emphasis>
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||
sp.:
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||
<treatmentCitation id="A918A270FF8DFFB6FC4ED57779590686" author="Arai, M. & Tanaka, Y. & Miyazaki, T. & Fujita, T." box="[937,1139,1142,1166]" page="194" pageId="6" pageNumber="13" year="2018">
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<bibRefCitation id="4C28F990FF8DFFB6FC4ED57779590686" author="Arai, M. & Tanaka, Y. & Miyazaki, T. & Fujita, T." box="[937,1139,1142,1166]" pageId="6" pageNumber="13" pagination="191 - 203" refId="ref10506" refString="Arai, M., Tanaka, Y., Miyazaki, T., and Fujita, T. 2018. Valvatida (Echinodermata: Asteroidea) of the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science 52: 191 - 203. [In Japanese]" type="journal article" year="2018">Arai et al. 2018: 194</bibRefCitation>
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</treatmentCitation>
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||
, 196.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="60A3D7EAFF8DFFB6FCA9D5B7781D0779" pageId="6" pageNumber="13" type="materials_examined">
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<paragraph id="28068461FF8DFFB6FCA9D5B7781D0779" blockId="6.[821,1432,1206,2076]" pageId="6" pageNumber="13">
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||
<emphasis id="1ACD5873FF8DFFB6FCA9D5B7793606C6" bold="true" box="[846,1052,1206,1230]" pageId="6" pageNumber="13">Material examined.</emphasis>
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||
<materialsCitation id="98D18E3CFF8DFFB6FBC3D5B679FB06E7" ID-GBIF-Occurrence="3097624304" collectionCode="NSMT" elevation="60" location="East of Ototo-jima Island" pageId="6" pageNumber="13" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="F7023AC3FF8DFFB6FBC3D5B679AF06C7" box="[1060,1157,1207,1231]" pageId="6" pageNumber="13" type="holotype">Holotype</typeStatus>
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||
:
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||
<collectionCode id="4EA81CA4FF8DFFB6FB68D5B679FE06C7" box="[1167,1236,1207,1231]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="13">NSMT</collectionCode>
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E-9297, KY-16-14,
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||
<location id="2D66D2BAFF8DFFB6FCD2D5D9796D06E7" LSID="urn:lsid:plazi:treatment:A0103577FF8DFFB8FC09D2F27EA806E7:2D66D2BAFF8DFFB6FCD2D5D9796D06E7" box="[821,1095,1239,1263]" name="East of Ototo-jima Island" pageId="6" pageNumber="13">East of Ototo-jima Island</location>
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||
,
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<quantity id="EF412984FF8DFFB6FBB2D5D679FB06E7" box="[1109,1233,1239,1263]" metricMagnitude="1" metricUnit="m" metricValue="5.975" metricValueMax="6.29" metricValueMin="5.66" pageId="6" pageNumber="13" unit="m" value="59.75" valueMax="62.9" valueMin="56.6">
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<elevation id="A3946352FF8DFFB6FBB2D5D679FB06E7" box="[1109,1233,1239,1263]" metricMagnitude="1" metricUnit="m" metricValue="5.975" metricValueMax="6.29" metricValueMin="5.66" pageId="6" pageNumber="13" unit="m" value="59.75" valueMax="62.9" valueMin="56.6">56.6–62.9 m</elevation>
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</quantity>
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</materialsCitation>
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.
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||
<typeStatus id="F7023AC3FF8DFFB6FB07D5D9786E06F8" box="[1248,1348,1240,1264]" pageId="6" pageNumber="13" type="paratype">Paratypes</typeStatus>
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:
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||
<collectionCode id="4EA81CA4FF8DFFB6FAB4D5D678B206E7" box="[1363,1432,1239,1263]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="13">NSMT</collectionCode>
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||
E-9293, KY-08-25, West of Nishi-jima Island,
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||
<quantity id="EF412984FF8DFFB6FAC3D5F978B90707" box="[1316,1427,1272,1296]" metricMagnitude="2" metricUnit="m" metricValue="1.28" metricValueMax="1.29" metricValueMin="1.27" pageId="6" pageNumber="13" unit="m" value="128.0" valueMax="129.0" valueMin="127.0">127–129 m</quantity>
|
||
;
|
||
<collectionCode id="4EA81CA4FF8DFFB6FCD2D4197E500738" box="[821,890,1304,1328]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="13">NSMT</collectionCode>
|
||
E-9294 and E-9295, KY-16-06, Northwest of Ototojima Island,
|
||
<quantity id="EF412984FF8DFFB6FC5ED43879020758" box="[953,1064,1337,1361]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="1.3699999999999999" metricValueMin="1.35" pageId="6" pageNumber="13" unit="m" value="136.0" valueMax="137.0" valueMin="135.0">135–137 m</quantity>
|
||
;
|
||
<collectionCode id="4EA81CA4FF8DFFB6FBD0D43879560759" box="[1079,1148,1337,1361]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="13">NSMT</collectionCode>
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||
E-9296, KY-16-09, East of Chichi-jima Island,
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||
<quantity id="EF412984FF8DFFB6FC19D45879530779" box="[1022,1145,1369,1393]" metricMagnitude="1" metricUnit="m" metricValue="9.254999999999999" metricValueMax="9.459999999999999" metricValueMin="9.05" pageId="6" pageNumber="13" unit="m" value="92.55" valueMax="94.6" valueMin="90.5">90.5–94.6 m</quantity>
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||
; 1 individual each.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF8DFFB6FCA9D47B791D0531" pageId="6" pageNumber="13" type="diagnosis">
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<paragraph id="28068461FF8DFFB6FCA9D47B791D0531" blockId="6.[821,1432,1206,2076]" pageId="6" pageNumber="13">
|
||
<emphasis id="1ACD5873FF8DFFB6FCA9D47B7E94079A" bold="true" box="[846,958,1402,1426]" pageId="6" pageNumber="13">Diagnosis.</emphasis>
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||
A species of the genus
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||
<taxonomicName id="EFB9FFE2FF8DFFB6FB25D47B78200799" box="[1218,1290,1402,1425]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="6" pageNumber="13" phylum="Echinodermata" rank="genus">
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||
<emphasis id="1ACD5873FF8DFFB6FB25D47B78200799" box="[1218,1290,1402,1425]" italics="true" pageId="6" pageNumber="13">Fromia</emphasis>
|
||
</taxonomicName>
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||
with a small disc and slender, slightly arched arms. At R=
|
||
<quantity id="EF412984FF8DFFB6FADCD49B78B907BA" box="[1339,1427,1434,1458]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="6" pageNumber="13" unit="mm" value="31.3">31.3 mm</quantity>
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||
, abactinal plates arranged in five longitudinal rows at base of arms. Superomarginal plates only slightly convex, not alternating but regularly decreasing in size toward the tip of arms. Coarse granules enlarged at center of plates. Papulae single and confined on the abactinal surface. Adambulacral plates bear three (rarely two or four) furrow spines and five to nine thicker subambulacral granules graduating in size toward those on neighboring actinal/marginal plates. Oral plates bear five oral spines on the margin and four to six suboral spines on the rest of the plate. Large, elliptical pedicellariae on many actinal plates with their major axis oblique to the ambulacra.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF8DFFB8FCA9D6407C4B0759" lastPageId="8" lastPageNumber="15" pageId="6" pageNumber="13" type="description">
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<paragraph id="28068461FF8DFFB6FCA9D640790405D3" blockId="6.[821,1432,1206,2076]" pageId="6" pageNumber="13">
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||
<emphasis id="1ACD5873FF8DFFB6FCA9D640797F0551" bold="true" box="[846,1109,1857,1881]" pageId="6" pageNumber="13">
|
||
Description of
|
||
<typeStatus id="F7023AC3FF8DFFB6FC15D640797B0551" box="[1010,1105,1857,1881]" pageId="6" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
R=
|
||
<quantity id="EF412984FF8DFFB6FB64D64079F60551" box="[1155,1244,1857,1881]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="6" pageNumber="13" unit="mm" value="31.3">31.3 mm</quantity>
|
||
, r=
|
||
<quantity id="EF412984FF8DFFB6FAE0D640787E0551" box="[1287,1364,1857,1881]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="13" unit="mm" value="8.6">8.6 mm</quantity>
|
||
, R/r= 3.6, width of arm is 9.0 mm at base,
|
||
<quantity id="EF412984FF8DFFB6FB26D66378210571" box="[1217,1291,1890,1914]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="6" pageNumber="13" unit="mm" value="5.4">5.4 mm</quantity>
|
||
at half of R, and
|
||
<quantity id="EF412984FF8DFFB6FC84D6837E870592" box="[867,941,1922,1946]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="13" unit="mm" value="2.9">2.9 mm</quantity>
|
||
at 1/10 R from the tip. Body is slightly arched at both sides, and the abactinal interradial areas are sunken. Arms taper to blunt tips.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8DFFB7FCA9D6E57CF60593" blockId="6.[821,1432,1206,2076]" lastBlockId="7.[156,767,1695,2077]" lastPageId="7" lastPageNumber="14" pageId="6" pageNumber="13">
|
||
Abactinal plates are polygonal, weakly lobate, arranged in staggered longitudinal series where there are five rows at the base of arms (
|
||
<figureCitation id="B08298E4FF8CFFB7FECCD79E7C5004BF" box="[299,378,1695,1719]" captionStart="Fig" captionStartId="7.[157,188,1539,1561]" captionTargetBox="[188,1398,206,1499]" captionTargetId="figure-228@7.[187,1399,205,1500]" captionTargetPageId="7" captionText="Fig. 5. Fromia labeosa sp. nov., NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5mm (A, C), 1mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines." figureDoi="http://doi.org/10.5281/zenodo.4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="7" pageNumber="14">Fig. 5C</figureCitation>
|
||
). Hemispherical glassy bosses (crystal bodies) occur on the abactinal plates. Madreporite is single, circular, and about
|
||
<quantity id="EF412984FF8CFFB7FE86D7E17C8104FF" box="[353,427,1760,1784]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="14" unit="mm" value="0.9">0.9 mm</quantity>
|
||
in diameter, located at about one half r from the anal aperture. Gyri do not regularly radiate from the center of the madreporite. The anal aperture is single and surrounded by six slightly larger ossicles. Terminal plates are conical, truncated at the tip. Several tubercles encircle the top of terminal plates.
|
||
</paragraph>
|
||
<caption id="7CC6D4E9FF8CFFB7FF7AD7027E040464" ID-DOI="http://doi.org/10.5281/zenodo.4585305" ID-Zenodo-Dep="4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="7" pageNumber="14" startId="7.[157,188,1539,1561]" targetBox="[188,1398,206,1499]" targetPageId="7">
|
||
<paragraph id="28068461FF8CFFB7FF7AD7027E040464" blockId="7.[156,1432,1538,1644]" pageId="7" pageNumber="14">
|
||
Fig. 5.
|
||
<taxonomicName id="EFB9FFE2FF8CFFB7FF17D7027C520410" authority="Arai & Fujita, 2021" authorityName="Arai & Fujita" authorityYear="2021" box="[240,376,1539,1560]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="14" phylum="Echinodermata" rank="species" species="labeosa" status="sp. nov.">
|
||
<emphasis id="1ACD5873FF8CFFB7FF17D7027C520410" box="[240,376,1539,1560]" italics="true" pageId="7" pageNumber="14">Fromia labeosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01FEE508FF8CFFB7FE67D7027CE20410" box="[384,456,1539,1560]" pageId="7" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5 mm (A, C), 1 mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="28068461FF8CFFB7FF52D6A27EBC04F0" blockId="7.[156,767,1695,2077]" lastBlockId="7.[820,1432,1695,2077]" pageId="7" pageNumber="14">
|
||
Superomarginal and inferomarginal plates are block-like, rectangular, only slightly convex and not alternating but regularly decreasing in size toward the tip of arms. There are sixteen superomarginal and seventeen inferomarginal plates on one side of the ray when skin and granules are removed. The second to fourth superomarginal plates average
|
||
<quantity id="EF412984FF8CFFB7FAAAD7C178BD04DF" box="[1357,1431,1728,1752]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="14" unit="mm" value="1.7">1.7 mm</quantity>
|
||
in length.
|
||
</paragraph>
|
||
<paragraph id="28068461FF8CFFB7FCA9D60178BE0572" blockId="7.[820,1432,1695,2077]" pageId="7" pageNumber="14">Actinal plates are polygonal, without lobes, and leave no space in between. No glassy bosses occur on actinal plates. These are arranged in three longitudinal series at the base of a denuded arm with a few odd plates on the interradial area.</paragraph>
|
||
<paragraph id="28068461FF8CFFB8FCA9D6827F5E008D" blockId="7.[820,1432,1695,2077]" lastBlockId="8.[156,771,589,2076]" lastPageId="8" lastPageNumber="15" pageId="7" pageNumber="14">
|
||
Adambulacral plates bear three (exceptionally two or four), slender, flattened, cylindrical and bluntly pointed furrow spines and five to nine thicker, more sharply pointed subambulacral granules (
|
||
<figureCitation id="B08298E4FF8CFFB7FBD4D6E5795405F4" box="[1075,1150,2020,2044]" captionStart="Fig" captionStartId="7.[157,188,1539,1561]" captionTargetBox="[188,1398,206,1499]" captionTargetId="figure-228@7.[187,1399,205,1500]" captionTargetPageId="7" captionText="Fig. 5. Fromia labeosa sp. nov., NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5mm (A, C), 1mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines." figureDoi="http://doi.org/10.5281/zenodo.4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="7" pageNumber="14">Fig. 5F</figureCitation>
|
||
). The latter are not in regular rows parallel to the ambulacrum, and are grading in size to the granules on neighboring actinal or marginal plates. Tubefeet are biserial with a stout terminal disc.
|
||
</paragraph>
|
||
<caption id="7CC6D4E9FF83FFB8FF7BD1BF7E3002DC" box="[156,794,190,212]" pageId="8" pageNumber="15">
|
||
<paragraph id="28068461FF83FFB8FF7BD1BF7E3002DC" blockId="8.[156,794,190,212]" box="[156,794,190,212]" pageId="8" pageNumber="15">
|
||
Table 2. Variation in numerical characters of
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FDADD1BE7FFA02DB" authority="Arai & Fujita, 2021" authorityName="Arai & Fujita" authorityYear="2021" box="[586,720,190,212]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa" status="sp. nov.">
|
||
<emphasis id="1ACD5873FF83FFB8FDADD1BE7FFA02DB" box="[586,720,190,212]" italics="true" pageId="8" pageNumber="15">Fromia labeosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01FEE508FF83FFB8FD31D1BE7E3002DC" box="[726,794,191,212]" pageId="8" pageNumber="15" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="28068461FF83FFB8FEF4D1F37860000F" pageId="8" pageNumber="15">
|
||
<table id="5AB976C1FF83004FFF56D1F37854000F" box="[177,1406,242,519]" gridcols="6" gridrows="9" pageId="8" pageNumber="15">
|
||
<tr id="96898623FF83004FFF56D1F37854030D" box="[177,1406,242,261]" gridrow="0" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D1F37CC1030D" box="[177,491,242,261]" gridcol="0" gridrow="0" pageId="8" pageNumber="15">Catalogue Number</th>
|
||
<th id="D558EF5FFF83004FFDA0D1F37FE9030D" box="[583,707,242,261]" gridcol="1" gridrow="0" pageId="8" pageNumber="15">NSMT E-9297</th>
|
||
<th id="D558EF5FFF83004FFD11D1F37E5B030D" box="[758,881,242,261]" gridcol="2" gridrow="0" pageId="8" pageNumber="15">NSMT E-9293</th>
|
||
<th id="D558EF5FFF83004FFC42D1F3790A030D" box="[933,1056,242,261]" gridcol="3" gridrow="0" pageId="8" pageNumber="15">NSMT E-9294</th>
|
||
<th id="D558EF5FFF83004FFBB4D1F379E5030D" box="[1107,1231,242,261]" gridcol="4" gridrow="0" pageId="8" pageNumber="15">NSMT E-9295</th>
|
||
<th id="D558EF5FFF83004FFAE5D1F37854030D" box="[1282,1406,242,261]" gridcol="5" gridrow="0" pageId="8" pageNumber="15">NSMT E-9296</th>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D01A78540327" box="[177,1406,283,303]" gridrow="1" pageId="8" pageNumber="15" rowspan-0="1">
|
||
<td id="D558EF5FFF83004FFDA0D01A7FE90327" box="[583,707,283,303]" gridcol="1" gridrow="1" pageId="8" pageNumber="15">Holotype</td>
|
||
<td id="D558EF5FFF83004FFD11D01A7E5B0327" box="[758,881,283,303]" gridcol="2" gridrow="1" pageId="8" pageNumber="15">Paratype</td>
|
||
<td id="D558EF5FFF83004FFC42D01A790A0327" box="[933,1056,283,303]" gridcol="3" gridrow="1" pageId="8" pageNumber="15">Paratype</td>
|
||
<td id="D558EF5FFF83004FFBB4D01A79E50327" box="[1107,1231,283,303]" gridcol="4" gridrow="1" pageId="8" pageNumber="15">Paratype</td>
|
||
<td id="D558EF5FFF83004FFAE5D01A78540327" box="[1282,1406,283,303]" gridcol="5" gridrow="1" pageId="8" pageNumber="15">Paratype</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D03B78540346" box="[177,1406,314,334]" gridrow="2" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D03B7CC10346" box="[177,491,314,334]" gridcol="0" gridrow="2" pageId="8" pageNumber="15">R (mm)</th>
|
||
<td id="D558EF5FFF83004FFDA0D03B7FE90346" box="[583,707,314,334]" gridcol="1" gridrow="2" pageId="8" pageNumber="15">31.3</td>
|
||
<td id="D558EF5FFF83004FFD11D03B7E5B0346" box="[758,881,314,334]" gridcol="2" gridrow="2" pageId="8" pageNumber="15">11.7</td>
|
||
<td id="D558EF5FFF83004FFC42D03B790A0346" box="[933,1056,314,334]" gridcol="3" gridrow="2" pageId="8" pageNumber="15">3.9</td>
|
||
<td id="D558EF5FFF83004FFBB4D03B79E50346" box="[1107,1231,314,334]" gridcol="4" gridrow="2" pageId="8" pageNumber="15">25.5</td>
|
||
<td id="D558EF5FFF83004FFAE5D03B78540346" box="[1282,1406,314,334]" gridcol="5" gridrow="2" pageId="8" pageNumber="15">11.6</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D05878540364" box="[177,1406,345,364]" gridrow="3" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D0587CC10364" box="[177,491,345,364]" gridcol="0" gridrow="3" pageId="8" pageNumber="15">R/r</th>
|
||
<td id="D558EF5FFF83004FFDA0D0587FE90364" box="[583,707,345,364]" gridcol="1" gridrow="3" pageId="8" pageNumber="15">3.6</td>
|
||
<td id="D558EF5FFF83004FFD11D0587E5B0364" box="[758,881,345,364]" gridcol="2" gridrow="3" pageId="8" pageNumber="15">3.5</td>
|
||
<td id="D558EF5FFF83004FFC42D058790A0364" box="[933,1056,345,364]" gridcol="3" gridrow="3" pageId="8" pageNumber="15">1.9</td>
|
||
<td id="D558EF5FFF83004FFBB4D05879E50364" box="[1107,1231,345,364]" gridcol="4" gridrow="3" pageId="8" pageNumber="15">4.4</td>
|
||
<td id="D558EF5FFF83004FFAE5D05878540364" box="[1282,1406,345,364]" gridcol="5" gridrow="3" pageId="8" pageNumber="15">2.5</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D07978540384" box="[177,1406,376,396]" gridrow="4" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D0797CC10384" box="[177,491,376,396]" gridcol="0" gridrow="4" pageId="8" pageNumber="15">Furrow spines per plate</th>
|
||
<td id="D558EF5FFF83004FFDA0D0797FE90384" box="[583,707,376,396]" gridcol="1" gridrow="4" pageId="8" pageNumber="15">3 (rarely 2–4)</td>
|
||
<td id="D558EF5FFF83004FFD11D0797E5B0384" box="[758,881,376,396]" gridcol="2" gridrow="4" pageId="8" pageNumber="15">3–4</td>
|
||
<td id="D558EF5FFF83004FFC42D079790A0384" box="[933,1056,376,396]" gridcol="3" gridrow="4" pageId="8" pageNumber="15">3–4</td>
|
||
<td id="D558EF5FFF83004FFBB4D07979E50384" box="[1107,1231,376,396]" gridcol="4" gridrow="4" pageId="8" pageNumber="15">3 (rarely 2–4)</td>
|
||
<td id="D558EF5FFF83004FFAE5D07978540384" box="[1282,1406,376,396]" gridcol="5" gridrow="4" pageId="8" pageNumber="15">2–3</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D096785403A3" box="[177,1406,407,427]" gridrow="5" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D0967CC103A3" box="[177,491,407,427]" gridcol="0" gridrow="5" pageId="8" pageNumber="15">Subambulacral spines per plate</th>
|
||
<td id="D558EF5FFF83004FFDA0D0967FE903A3" box="[583,707,407,427]" gridcol="1" gridrow="5" pageId="8" pageNumber="15">5–9</td>
|
||
<td id="D558EF5FFF83004FFD11D0967E5B03A3" box="[758,881,407,427]" gridcol="2" gridrow="5" pageId="8" pageNumber="15">7–9</td>
|
||
<td id="D558EF5FFF83004FFC42D096790A03A3" box="[933,1056,407,427]" gridcol="3" gridrow="5" pageId="8" pageNumber="15">7–8</td>
|
||
<td id="D558EF5FFF83004FFBB4D09679E503A3" box="[1107,1231,407,427]" gridcol="4" gridrow="5" pageId="8" pageNumber="15">5–8</td>
|
||
<td id="D558EF5FFF83004FFAE5D096785403A3" box="[1282,1406,407,427]" gridcol="5" gridrow="5" pageId="8" pageNumber="15">5–7</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D0B4785403C1" box="[177,1406,437,457]" gridrow="6" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D0B47CC103C1" box="[177,491,437,457]" gridcol="0" gridrow="6" pageId="8" pageNumber="15">Granules/mm2 on abactinal plates</th>
|
||
<td id="D558EF5FFF83004FFDA0D0B47FE903C1" box="[583,707,437,457]" gridcol="1" gridrow="6" pageId="8" pageNumber="15">20</td>
|
||
<td id="D558EF5FFF83004FFD11D0B47E5B03C1" box="[758,881,437,457]" gridcol="2" gridrow="6" pageId="8" pageNumber="15">40</td>
|
||
<td id="D558EF5FFF83004FFC42D0B4790A03C1" box="[933,1056,437,457]" gridcol="3" gridrow="6" pageId="8" pageNumber="15">87</td>
|
||
<td id="D558EF5FFF83004FFBB4D0B479E503C1" box="[1107,1231,437,457]" gridcol="4" gridrow="6" pageId="8" pageNumber="15">26</td>
|
||
<td id="D558EF5FFF83004FFAE5D0B4785403C1" box="[1282,1406,437,457]" gridcol="5" gridrow="6" pageId="8" pageNumber="15">36</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D0D5785403E0" box="[177,1406,468,488]" gridrow="7" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D0D57CC103E0" box="[177,491,468,488]" gridcol="0" gridrow="7" pageId="8" pageNumber="15">Granules/mm2 on actinal plates</th>
|
||
<td id="D558EF5FFF83004FFDA0D0D57FE903E0" box="[583,707,468,488]" gridcol="1" gridrow="7" pageId="8" pageNumber="15">20</td>
|
||
<td id="D558EF5FFF83004FFD11D0D57E5B03E0" box="[758,881,468,488]" gridcol="2" gridrow="7" pageId="8" pageNumber="15">46</td>
|
||
<td id="D558EF5FFF83004FFC42D0D5790A03E0" box="[933,1056,468,488]" gridcol="3" gridrow="7" pageId="8" pageNumber="15">101</td>
|
||
<td id="D558EF5FFF83004FFBB4D0D579E503E0" box="[1107,1231,468,488]" gridcol="4" gridrow="7" pageId="8" pageNumber="15">42</td>
|
||
<td id="D558EF5FFF83004FFAE5D0D5785403E0" box="[1282,1406,468,488]" gridcol="5" gridrow="7" pageId="8" pageNumber="15">34</td>
|
||
</tr>
|
||
<tr id="96898623FF83004FFF56D0F57854000F" box="[177,1406,500,519]" gridrow="8" pageId="8" pageNumber="15">
|
||
<th id="D558EF5FFF83004FFF56D0F57CC1000F" box="[177,491,500,519]" gridcol="0" gridrow="8" pageId="8" pageNumber="15">Longitudinal rows of abactinal plates</th>
|
||
<td id="D558EF5FFF83004FFDA0D0F57FE9000F" box="[583,707,500,519]" gridcol="1" gridrow="8" pageId="8" pageNumber="15">5</td>
|
||
<td id="D558EF5FFF83004FFD11D0F57E5B000F" box="[758,881,500,519]" gridcol="2" gridrow="8" pageId="8" pageNumber="15">3</td>
|
||
<td id="D558EF5FFF83004FFC42D0F5790A000F" box="[933,1056,500,519]" gridcol="3" gridrow="8" pageId="8" pageNumber="15">3</td>
|
||
<td id="D558EF5FFF83004FFBB4D0F579E5000F" box="[1107,1231,500,519]" gridcol="4" gridrow="8" pageId="8" pageNumber="15">3</td>
|
||
<td id="D558EF5FFF83004FFAE5D0F57854000F" box="[1282,1406,500,519]" gridcol="5" gridrow="8" pageId="8" pageNumber="15">3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="28068461FF83FFB8FF52D38F7FAE0120" blockId="8.[156,771,589,2076]" pageId="8" pageNumber="15">Each oral plate bears five oral spines on the adoral margin and four to six suboral spines on the rest of the plate. Oral spines are conical or pyramidal, larger than furrow spines. Suboral spines are similar to oral spines in the adradial portion but abradially grading into actinal granules.</paragraph>
|
||
<paragraph id="28068461FF83FFB8FF52D2317FBC0665" blockId="8.[156,771,589,2076]" pageId="8" pageNumber="15">
|
||
The abactinal, marginal, and actinal plates are concealed with a skin which is covered coarsely with granules (
|
||
<figureCitation id="B08298E4FF83FFB8FD30D2507D910181" captionStart="Fig" captionStartId="7.[157,188,1539,1561]" captionTargetBox="[188,1398,206,1499]" captionTargetId="figure-228@7.[187,1399,205,1500]" captionTargetPageId="7" captionText="Fig. 5. Fromia labeosa sp. nov., NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5mm (A, C), 1mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines." figureDoi="http://doi.org/10.5281/zenodo.4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="8" pageNumber="15">Fig. 5B</figureCitation>
|
||
). These granules are domed at the top, polygonal in shape, and slightly enlarged at the center of plates, but never spinous. The diameter is ca.
|
||
<quantity id="EF412984FF83FFB8FE2CD2B37F6C01C3" box="[459,582,946,971]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="3.4" metricValueMin="2.6" pageId="8" pageNumber="15" unit="mm" value="300.0" valueMax="340.0" valueMin="260.0">260–340µm</quantity>
|
||
at center and ca.
|
||
<quantity id="EF412984FF83FFB8FF7BD2D27C3D01E4" box="[156,279,979,1004]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="8" pageNumber="15" unit="mm" value="170.0" valueMax="200.0" valueMin="140.0">140–200µm</quantity>
|
||
on periphery. There are 20 granules per square millimeter on abactinal plates at the base of arms and actinal interradial plates. The papular pores are isolated and confined around abactinal plates except on the interradial areas and tips of arms, where the pores are absent.
|
||
</paragraph>
|
||
<paragraph id="28068461FF83FFB8FF52D5777F1A0718" blockId="8.[156,771,589,2076]" pageId="8" pageNumber="15">
|
||
Many actinal plates possess a large elliptical pedicellaria (
|
||
<figureCitation id="B08298E4FF83FFB8FF44D5977DD806A6" box="[163,242,1174,1198]" captionStart="Fig" captionStartId="7.[157,188,1539,1561]" captionTargetBox="[188,1398,206,1499]" captionTargetId="figure-228@7.[187,1399,205,1500]" captionTargetPageId="7" captionText="Fig. 5. Fromia labeosa sp. nov., NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5mm (A, C), 1mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines." figureDoi="http://doi.org/10.5281/zenodo.4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="8" pageNumber="15">Fig. 5D</figureCitation>
|
||
). It is typically
|
||
<quantity id="EF412984FF83FFB8FE6DD5977CE806A6" box="[394,450,1174,1198]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="15" unit="mm" value="1.0">1 mm</quantity>
|
||
in length and
|
||
<quantity id="EF412984FF83FFB8FDB2D5977FB406A7" box="[597,670,1174,1199]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="15" unit="mm" value="400.0">400 µm</quantity>
|
||
in width. The pedicellariae are loosely arranged in a parallel series to an ambulacrum on arms, and their major axis is often oblique to the ambulacrum (
|
||
<figureCitation id="B08298E4FF83FFB8FE5BD5F97F0E0718" box="[444,548,1272,1296]" captionStart="Fig" captionStartId="7.[157,188,1539,1561]" captionTargetBox="[188,1398,206,1499]" captionTargetId="figure-228@7.[187,1399,205,1500]" captionTargetPageId="7" captionText="Fig. 5. Fromia labeosa sp. nov., NSMT E-9297, holotype. A, actinal surface of the disc; B, abactinal surface of interradial area showing a madreporite (arrow); C, abactinal surface of a denuded arm; D, pedicellaria on the actinal plate; E, proximal portion of a partly denuded arm, actinal side. Arrows indicate pedicellariae; F, adambulacral armature on proximal adambulacral plates. Scale bars: 5mm (A, C), 1mm (B, E), 500µm (D, F). Abbreviations: fs, furrow spines; ss, subambulacral spines." figureDoi="http://doi.org/10.5281/zenodo.4585305" httpUri="https://zenodo.org/record/4585305/files/figure.png" pageId="8" pageNumber="15">Fig. 5A, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="28068461FF83FFB8FF52D4197C4B0759" blockId="8.[156,771,589,2076]" pageId="8" pageNumber="15">
|
||
Color in life is vermilion with white granules and terminal plates (
|
||
<figureCitation id="B08298E4FF83FFB8FEEFD4387C7C0759" box="[264,342,1337,1361]" captionStart="Fig" captionStartId="2.[822,853,1874,1896]" captionTargetBox="[188,1402,207,2036]" captionTargetId="figure-6@2.[187,1400,205,2029]" captionTargetPageId="2" captionText="Fig. 2. Abactinal view of the live specimens (A–D) and ethanol-preserved specimen (E). A, Bathyferdina caelator sp. nov., NSMT E-8265, holotype; B, Fromia eusticha, NSMT E-10595; C, Fromia labeosa sp. nov., NSMT E-9297, holotype; D, Ogmaster capella, NSMT E-9312; E, Tamaria tenella, NSMT E-9277. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.4585299" httpUri="https://zenodo.org/record/4585299/files/figure.png" pageId="8" pageNumber="15">Fig. 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF83FFB8FF52D45B7F9B07FC" pageId="8" pageNumber="15" type="discussion">
|
||
<paragraph id="28068461FF83FFB8FF52D45B7F9B07FC" blockId="8.[156,771,589,2076]" pageId="8" pageNumber="15">
|
||
<emphasis id="1ACD5873FF83FFB8FF52D45B7CA3077A" bold="true" box="[181,393,1370,1394]" pageId="8" pageNumber="15">
|
||
Notes on
|
||
<typeStatus id="F7023AC3FF83FFB8FEFBD45B7CAF077A" box="[284,389,1370,1394]" pageId="8" pageNumber="15" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The difference in the numeric characters among type specimens is shown in
|
||
<tableCitation id="653BB1DAFF83FFB8FDA4D47B7F82079A" box="[579,680,1402,1426]" captionStart="Table 2" captionStartId="8.[156,204,190,212]" captionText="Table 2. Variation in numerical characters of Fromia labeosa sp.nov." pageId="8" pageNumber="15">Table 2. R</tableCitation>
|
||
/r generally increases and the number of granules per square millimeter decreases as R increases. Other characters in the
|
||
<typeStatus id="F7023AC3FF83FFB8FD2ED4BD7DFB07FC" pageId="8" pageNumber="15" type="paratype">paratypes</typeStatus>
|
||
are consistent with the description of
|
||
<typeStatus id="F7023AC3FF83FFB8FDB5D4DD7F8407FC" box="[594,686,1500,1524]" pageId="8" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF83FFB8FF52D4FD7C17045D" pageId="8" pageNumber="15" type="multiple">
|
||
<paragraph id="28068461FF83FFB8FF52D4FD7C17045D" blockId="8.[156,771,589,2076]" pageId="8" pageNumber="15">
|
||
<emphasis id="1ACD5873FF83FFB8FF52D4FD7C70041C" bold="true" box="[181,346,1532,1556]" pageId="8" pageNumber="15">DNA sequence.</emphasis>
|
||
A partial sequence of COI (655 bp) was obtained from NSMT E-9295 and deposited in DDBJ (Acc. No.
|
||
<accessionNumber id="37EA1982FF83FFB8FF21D73C7C18045D" box="[198,306,1597,1621]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC427074" pageId="8" pageNumber="15">LC427074</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF83FFB8FF52D75F781E01E3" pageId="8" pageNumber="15" type="discussion">
|
||
<paragraph id="28068461FF83FFB8FF52D75F781E01E3" blockId="8.[156,771,589,2076]" lastBlockId="8.[820,1435,588,1263]" pageId="8" pageNumber="15">
|
||
<emphasis id="1ACD5873FF83FFB8FF52D75F7C33047E" bold="true" box="[181,281,1630,1654]" pageId="8" pageNumber="15">Remarks.</emphasis>
|
||
The present species is distinguished from its congeners by the following characters: superomarginal plates regularly decreasing in size toward the arm tip in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FD09D79E7DCF04DF" authority="Arai & Fujita, 2021" authorityName="Arai & Fujita" authorityYear="2021" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa" status="sp. nov.">
|
||
<emphasis id="1ACD5873FF83FFB8FD09D79E7DCF04DF" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="01FEE508FF83FFB8FF09D7C17C1404DF" box="[238,318,1728,1751]" pageId="8" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
while they are alternating large and small, or small superomarginal plates are occasionally intercalated between larger ones, in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FE74D6007F9E0510" authority="H. L. Clark, 1921" authorityName="H. L. Clark" authorityYear="1921" box="[403,692,1792,1816]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="pacifica">
|
||
<emphasis id="1ACD5873FF83FFB8FE74D6007CD20510" box="[403,504,1792,1816]" italics="true" pageId="8" pageNumber="15">F. pacifica</emphasis>
|
||
H. L.
|
||
<bibRefCitation id="4C28F990FF83FFB8FDDBD6017F9E0510" author="Clark, H. L." box="[572,692,1792,1816]" pageId="8" pageNumber="15" refId="ref10766" refString="Clark, H. L. 1921. The Echinoderm Fauna of Torres Strait: Its Composition and Its Origin. Carnegie Institution of Washington, viii + 223 pp., 38 pls." type="book" year="1921">Clark, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FD27D6007C9D0531" authority="(Livingstone, 1931)" baseAuthorityName="Livingstone" baseAuthorityYear="1931" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="heffernani">
|
||
<emphasis id="1ACD5873FF83FFB8FD27D6007DCE0531" italics="true" pageId="8" pageNumber="15">F. heffernani</emphasis>
|
||
(Livingstone, 1931)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FE22D6237FE10531" authority="(Perrier, 1869)" baseAuthorityName="Perrier" baseAuthorityYear="1869" box="[453,715,1825,1849]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="monilis">
|
||
<emphasis id="1ACD5873FF83FFB8FE22D6237F030531" box="[453,553,1825,1849]" italics="true" pageId="8" pageNumber="15">F. monilis</emphasis>
|
||
(Perrier, 1869)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FF7BD6437CEE0551" authority="A. M. Clark, 1967" authorityName="A. M. Clark" authorityYear="1967" box="[156,452,1857,1882]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="nodosa">
|
||
<emphasis id="1ACD5873FF83FFB8FF7BD6437DD70551" box="[156,253,1857,1881]" italics="true" pageId="8" pageNumber="15">F. nodosa</emphasis>
|
||
A. M. Clark, 1967
|
||
</taxonomicName>
|
||
; there are five abactinal plates between first superomarginal plates on each side of arms in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FF5FD6827C330592" authorityName="Arai & Fujita" authorityYear="2021" box="[184,281,1922,1946]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FF5FD6827C330592" box="[184,281,1922,1946]" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
while there are more than seven such plates in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FF7BD6A57C8805B3" authority="Fisher, 1913" authorityName="Fisher" authorityYear="1913" box="[156,418,1955,1979]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="hemiopla">
|
||
<emphasis id="1ACD5873FF83FFB8FF7BD6A57C3F05B3" box="[156,277,1955,1979]" italics="true" pageId="8" pageNumber="15">F. hemiopla</emphasis>
|
||
<bibRefCitation id="4C28F990FF83FFB8FEC7D6A27C8805B3" author="Fisher, W. K." box="[288,418,1955,1979]" pageId="8" pageNumber="15" pagination="201 - 224" refId="ref10891" refString="Fisher, W. K. 1913. New starfishes from the Philippine Islands, Celebes, and Moluccas. Proceedings of the United States National Museum 46: 201 - 224" type="journal article" year="1913">Fisher, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FE56D6A57FD105B3" authority="(Lamarck, 1816)" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[433,763,1955,1979]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="milleporella">
|
||
<emphasis id="1ACD5873FF83FFB8FE56D6A57F6B05B3" box="[433,577,1955,1979]" italics="true" pageId="8" pageNumber="15">F. milleporella</emphasis>
|
||
(Lamarck, 1816)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FF7BD6C57CF805D3" authority="H. L. Clark, 1916" authorityName="H. L. Clark" authorityYear="1916" box="[156,466,1987,2011]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="polypora">
|
||
<emphasis id="1ACD5873FF83FFB8FF7BD6C57C2705D3" box="[156,269,1987,2011]" italics="true" pageId="8" pageNumber="15">F. polypora</emphasis>
|
||
H. L. Clark, 1916
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FDE8D6C57DFD05F4" authority="(Doderlein, 1910)" baseAuthorityName="Doderlein" baseAuthorityYear="1910" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="schultzei">
|
||
<emphasis id="1ACD5873FF83FFB8FDE8D6C57F5505D3" box="[527,639,1987,2011]" italics="true" pageId="8" pageNumber="15">F. schultzei</emphasis>
|
||
(Döderlein, 1910)
|
||
</taxonomicName>
|
||
; abactinal plates are uniform in size in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FD85D6E47FE805F4" authorityName="Arai & Fujita" authorityYear="2021" box="[610,706,2020,2044]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FD85D6E47FE805F4" box="[610,706,2020,2044]" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
while several elliptical plates are larger than others in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FD66D9047EED006D" authority="H. L. Clark, 1921" authorityName="H. L. Clark" authorityYear="1921" box="[641,967,588,2076]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="elegans">
|
||
<emphasis id="1ACD5873FF83FFB8FD66D9047FCA0A14" box="[641,736,2052,2076]" italics="true" pageId="8" pageNumber="15">F. elegans</emphasis>
|
||
H. L.
|
||
<bibRefCitation id="4C28F990FF83FFB8FCB7D34D7EED006D" author="Clark, H. L." box="[848,967,588,613]" pageId="8" pageNumber="15" refId="ref10766" refString="Clark, H. L. 1921. The Echinoderm Fauna of Torres Strait: Its Composition and Its Origin. Carnegie Institution of Washington, viii + 223 pp., 38 pls." type="book" year="1921">Clark, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FC1BD34C79DA006C" authority="(Perrier, 1869)" baseAuthorityName="Perrier" baseAuthorityYear="1869" box="[1020,1264,588,612]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="indica">
|
||
<emphasis id="1ACD5873FF83FFB8FC1BD34C7978006C" box="[1020,1106,588,612]" italics="true" pageId="8" pageNumber="15">F. indica</emphasis>
|
||
(Perrier, 1869)
|
||
</taxonomicName>
|
||
; abactinal granules are granular in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FBE2D36F794D008D" authorityName="Arai & Fujita" authorityYear="2021" box="[1029,1127,621,645]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FBE2D36F794D008D" box="[1029,1127,621,645]" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
while they are spinuous in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FA60D36F793200AE" authority="Koehler, 1910" authorityName="Koehler" authorityYear="1910" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="armata">
|
||
<emphasis id="1ACD5873FF83FFB8FA60D36F7E5400AD" italics="true" pageId="8" pageNumber="15">F. armata</emphasis>
|
||
<bibRefCitation id="4C28F990FF83FFB8FC60D38F793200AE" author="Koehler, R." box="[903,1048,654,678]" pageId="8" pageNumber="15" refId="ref11315" refString="Koehler, R. 1910. An Account of the Shallow-water Asteroidea. Trustees of the Indian Museum, Calcutta, 191 pp., 20 pls." type="book" year="1910">Koehler, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
; they are also coarse (20/mm
|
||
<superScript id="DFCC2929FF83FFB8FAA8D38F787D0094" attach="left" box="[1359,1367,654,668]" fontSize="6" pageId="8" pageNumber="15">2</superScript>
|
||
) in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FA60D38E7E5400CE" authorityName="Arai & Fujita" authorityYear="2021" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FA60D38E7E5400CE" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
while they are fine (80–90/mm
|
||
<superScript id="DFCC2929FF83FFB8FB2ED3AF79FB00B4" attach="left" box="[1225,1233,686,700]" fontSize="6" pageId="8" pageNumber="15">2</superScript>
|
||
) in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FAE7D3AE7ECC00EF" authority="H. L. Clark, 1921" authorityName="H. L. Clark" authorityYear="1921" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="hadracantha">
|
||
<emphasis id="1ACD5873FF83FFB8FAE7D3AE78B200CE" box="[1280,1432,686,710]" italics="true" pageId="8" pageNumber="15">F. hadracantha</emphasis>
|
||
H. L.
|
||
<bibRefCitation id="4C28F990FF83FFB8FC88D3CE7ECC00EF" author="Clark, H. L." box="[879,998,719,743]" pageId="8" pageNumber="15" refId="ref10766" refString="Clark, H. L. 1921. The Echinoderm Fauna of Torres Strait: Its Composition and Its Origin. Carnegie Institution of Washington, viii + 223 pp., 38 pls." type="book" year="1921">Clark, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
; papulae are lacking on the actinal surface in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FCA8D3F17E85010F" authorityName="Arai & Fujita" authorityYear="2021" box="[847,943,751,775]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FCA8D3F17E85010F" box="[847,943,751,775]" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
while there are two rows of actinal papulae in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FA60D3F1793D0120" authority="Perrier, 1875" authorityName="Perrier" authorityYear="1875" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="balansae">
|
||
<emphasis id="1ACD5873FF83FFB8FA60D3F17EA10120" italics="true" pageId="8" pageNumber="15">F. balansae</emphasis>
|
||
Perrier, 1875
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FBAAD21078BE0120" authority="Mortensen, 1938" authorityName="Mortensen" authorityYear="1938" box="[1101,1428,784,808]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="ghardaqana">
|
||
<emphasis id="1ACD5873FF83FFB8FBAAD21079F70120" box="[1101,1245,784,808]" italics="true" pageId="8" pageNumber="15">F. ghardaqana</emphasis>
|
||
Mortensen, 1938
|
||
</taxonomicName>
|
||
. Five to nine subambulacral tubercles continuous in size and not arranged in definite rows in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FB9BD25379F60161" authorityName="Arai & Fujita" authorityYear="2021" box="[1148,1244,849,873]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FB9BD25379F60161" box="[1148,1244,849,873]" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
while subambulacral spines are conspicuously larger than outer granules on the adambulacral plate, and arranged in a straight row in
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FA60D2927EAF01C3" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="eusticha">
|
||
<emphasis id="1ACD5873FF83FFB8FA60D2927EAF01C3" italics="true" pageId="8" pageNumber="15">F. eusticha</emphasis>
|
||
</taxonomicName>
|
||
. In addition to these differences, large elliptical pedicellariae on actinal plates are specific to
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FB29D2D5780401E3" authorityName="Arai & Fujita" authorityYear="2021" box="[1230,1326,979,1003]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FB29D2D5780401E3" box="[1230,1326,979,1003]" italics="true" pageId="8" pageNumber="15">F. labeosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF83FFB8FCA9D2F27E510624" pageId="8" pageNumber="15" type="distribution">
|
||
<paragraph id="28068461FF83FFB8FCA9D2F27E510624" blockId="8.[820,1435,588,1263]" pageId="8" pageNumber="15">
|
||
<emphasis id="1ACD5873FF83FFB8FCA9D2F27EF30603" bold="true" box="[846,985,1011,1035]" pageId="8" pageNumber="15">Distribution.</emphasis>
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FC15D2F579EB0604" authority="Islands" authorityName="Islands" box="[1010,1217,1012,1036]" class="Insecta" family="Cerambycidae" genus="Ogasawara" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="genus">Ogasawara Islands</taxonomicName>
|
||
,
|
||
<quantity id="EF412984FF83FFB8FB3AD2F2787B0603" box="[1245,1361,1011,1035]" metricMagnitude="1" metricUnit="m" metricValue="9.68" metricValueMax="13.7" metricValueMin="5.66" pageId="8" pageNumber="15" unit="m" value="96.8" valueMax="137.0" valueMin="56.6">56.6–137 m</quantity>
|
||
(this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60A3D7EAFF83FFB8FCA9D5347EA806E7" pageId="8" pageNumber="15" type="etymology">
|
||
<paragraph id="28068461FF83FFB8FCA9D53479FA0686" blockId="8.[820,1435,588,1263]" pageId="8" pageNumber="15">
|
||
<emphasis id="1ACD5873FF83FFB8FCA9D5347EEF0645" bold="true" box="[846,965,1077,1101]" pageId="8" pageNumber="15">Etymology.</emphasis>
|
||
The species epithet,
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FB47D53479C30645" authorityName="Arai & Fujita" authorityYear="2021" box="[1184,1257,1077,1101]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="species" species="labeosa">
|
||
<emphasis id="1ACD5873FF83FFB8FB47D53479C30645" box="[1184,1257,1077,1101]" italics="true" pageId="8" pageNumber="15">labeosa</emphasis>
|
||
</taxonomicName>
|
||
, is a Latin feminine adjective meaning having large lips and referring to the large pedicellariae on the actinal surface.
|
||
</paragraph>
|
||
<paragraph id="28068461FF83FFB8FCA9D5977EA806E7" blockId="8.[820,1435,588,1263]" pageId="8" pageNumber="15">
|
||
<emphasis id="1ACD5873FF83FFB8FCA9D5977EDF06A6" bold="true" box="[846,1013,1174,1198]" pageId="8" pageNumber="15">Japanese name.</emphasis>
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FBE6D596795B06A6" box="[1025,1137,1175,1198]" class="Insecta" family="Cerambycidae" genus="Ogasawara" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1ACD5873FF83FFB8FBE6D596795B06A6" box="[1025,1137,1175,1198]" italics="true" pageId="8" pageNumber="15">Ogasawara</emphasis>
|
||
</taxonomicName>
|
||
is taken from the
|
||
<typeStatus id="F7023AC3FF83FFB8FAA3D596785A06A7" box="[1348,1392,1175,1199]" pageId="8" pageNumber="15">type</typeStatus>
|
||
locality and
|
||
<emphasis id="1ACD5873FF83FFB8FC45D5B6791006C7" box="[930,1082,1207,1231]" italics="true" pageId="8" pageNumber="15">juzuberi-hitode</emphasis>
|
||
is a Japanese name for the genus
|
||
<taxonomicName id="EFB9FFE2FF83FFB8FCD3D5D97E5606E7" box="[820,892,1240,1263]" class="Asteroidea" family="Goniasteridae" genus="Fromia" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="15" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="1ACD5873FF83FFB8FCD3D5D97E5606E7" box="[820,892,1240,1263]" italics="true" pageId="8" pageNumber="15">Fromia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |