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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.104699" ID-Pensoft-Pub="1313-2970-1168-329" ID-Pensoft-UUID="EBBBDFFED8C15BF593E1E2D493E1C06B" ID-ZooBank="251CDBED1028441D8DC4AAEA3BF4A4E7" ModsDocID="1313-2970-1168-329" checkinTime="1688525912427" checkinUser="pensoft" docAuthor="Zhou, Jiale, Evenhuis, Neal L. &amp; Yang, Ding" docDate="2023" docId="71E0657FC3155D5095866048468ED52A" docLanguage="en" docName="ZooKeys 1168: 329-353" docOrigin="ZooKeys 1168" docPubDate="2023-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1168.104699" docTitle="Strongylophthalmyia narwhal Evenhuis 2020" docType="treatment" docVersion="1" id="EBBBDFFED8C15BF593E1E2D493E1C06B" lastPageNumber="329" masterDocId="EBBBDFFED8C15BF593E1E2D493E1C06B" masterDocTitle="A review of the Strongylophthalmyia coarctata subgroup (Diptera, Brachycera, Strongylophthalmyiidae) from China, with the descriptions of three new species" masterLastPageNumber="353" masterPageNumber="329" pageNumber="329" updateTime="1688525912427" updateUser="pensoft">
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<mods:title id="916AE2892D1589B379F4B3249426F58F">A review of the Strongylophthalmyia coarctata subgroup (Diptera, Brachycera, Strongylophthalmyiidae) from China, with the descriptions of three new species</mods:title>
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<mods:namePart id="8EB1619C9F7BD7B8CD8CFF229A5345F7">Zhou, Jiale</mods:namePart>
<mods:nameIdentifier id="432B7C02A2409F182197FDE080E0E26E" type="ORCID">https://orcid.org/0000-0002-8901-3187</mods:nameIdentifier>
<mods:affiliation id="8D1F8716543E18FCD40CB81568689CFD">Department of Entomology, College of Plant Protection, China Agricultural University, 2 Yuanmingyuan West Road, 100193 Beijing, China</mods:affiliation>
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<mods:namePart id="1C63BA696738BB6DBA1A0587F4738F21">Evenhuis, Neal L.</mods:namePart>
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<mods:affiliation id="D6B0F58307A7A522D50954B2D6B40D4E">Department of Natural Sciences, Bernice Pauahi Bishop Museum, 1525 Bernice Street, Honolulu, 96817 - 2704 Hawaii, USA</mods:affiliation>
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<mods:namePart id="CE91BE10A201F8138F097CBF8B08274B">Yang, Ding</mods:namePart>
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<mods:affiliation id="67CDD263302C69FEA9611D2774BAEE12">Department of Entomology, College of Plant Protection, China Agricultural University, 2 Yuanmingyuan West Road, 100193 Beijing, China</mods:affiliation>
<mods:nameIdentifier id="F3D17F92D629F784DF7796FA0754927D" type="email">dyangcau@126.com</mods:nameIdentifier>
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<mods:date id="0F685E7877497B1E689159004212F64B">2023</mods:date>
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<treatment id="71E0657FC3155D5095866048468ED52A" LSID="urn:lsid:plazi:treatment:71E0657FC3155D5095866048468ED52A" httpUri="http://treatment.plazi.org/id/71E0657FC3155D5095866048468ED52A" lastPageNumber="329" pageId="0" pageNumber="329">
<subSubSection id="B866A4D03F3AE7E38BA6801BCD226644" pageId="0" pageNumber="329" type="nomenclature">
<paragraph id="57795299600F55185FD9B40958F07FDF" pageId="0" pageNumber="329">
<taxonomicName id="D77C772B960E176B58AA38AF13D246E7" LSID="71E0657F-C315-5D50-9586-6048468ED52A" authority="Evenhuis, 2020" authorityName="Evenhuis" authorityYear="2020" class="Insecta" family="Strongylophthalmyiidae" genus="Strongylophthalmyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylophthalmyia narwhal" order="Diptera" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="narwhal">Strongylophthalmyia narwhal Evenhuis, 2020</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="640A99B1AE43B098A494DE1A0FBE1E21" pageId="0" pageNumber="329" type="description">
<paragraph id="5136D6909FDF49D384EF3BBA97164C36" pageId="0" pageNumber="329">
<figureCitation id="CA98FBF2B41DDE830B940201DB8005E5" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male, habitus, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1168.104699.figure24" httpUri="https://binary.pensoft.net/fig/872799" pageId="0" pageNumber="329">Figs 24</figureCitation>
<figureCitation id="18C9261B734B051351B69B9D8EB438E2" captionStart="Figures 2529" captionStartId="F9" captionText="Figures 25 - 29. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male: 25 head, dorsal view 26 same, lateral view 27 left antenna, lateral view 28 thorax, dorsal view 29 wing. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1168.104699.figures25-29" httpUri="https://binary.pensoft.net/fig/872800" pageId="0" pageNumber="329">, 25-29</figureCitation>
<figureCitation id="AE262143AFAEB36D8E6A9384EFB18346" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">, 30-33</figureCitation>
</paragraph>
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<paragraph id="64F3301B38609B21BBD801DB4FDE4F59" pageId="0" pageNumber="329">
<taxonomicName id="B043867BEF42DEA93A395A91C6868706" authorityName="Evenhuis" authorityYear="2020" class="Insecta" family="Strongylophthalmyiidae" genus="Strongylophthalmyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylophthalmyia narwhal" order="Diptera" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="narwhal">Strongylophthalmyia narwhal</taxonomicName>
Evenhuis, 2020: 5 (protologue). Holotype (♂): Thailand, Chiang Mai, Chiang Mai, QSBG.
</paragraph>
</subSubSection>
<subSubSection id="074B3B70E4C42082AC11469644620FED" pageId="0" pageNumber="329" type="type material">
<paragraph id="087E8DC0BF50F70F21CB50E69EA545C9" pageId="0" pageNumber="329">Type material examined.</paragraph>
<paragraph id="C06DD355E234A083C0C167F13C6838CE" pageId="0" pageNumber="329">
<emphasis id="44ED21B9F8236A4C4898A0AFCC82D2E1" bold="true" italics="true" pageId="0" pageNumber="329">Holotype</emphasis>
(♂): Thailand, Chiang Mai, Queen Sirikit Botanical Garden, 2013.vii.11-25, leg. M. Hauser, by Malaise trap (QSBG).
</paragraph>
</subSubSection>
<subSubSection id="20B9ADE6EEDD0F6B9C5BFA559E3F9DB6" pageId="0" pageNumber="329" type="materials_examined">
<paragraph id="9B09C177FA62AD3867FB79645CF58723" pageId="0" pageNumber="329">Other materials examined.</paragraph>
<paragraph id="3E7119EAC57D358322ED75E2D02432A7" pageId="0" pageNumber="329">
<materialsCitation id="F55387BA44521955EB5ADB0BC0B6E6F7" collectingDate="2018-04-15" collectingDateMax="2020-11-25" collectingDateMin="2018-04-15" collectorName="Liang Wang" country="China" county="Honghe" elevation="380" location="Huanglianshan" municipality="Lvchun" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan">
<collectingCountry id="213976BD3FC8DE95D83B7007A8AD8D28" name="China">China</collectingCountry>
.
<collectingRegion id="B078D4BA2A3B643C8EED551D86601F54" country="China" name="Yunnan">Yunnan</collectingRegion>
:
<collectingCounty id="B15A757610C1EDCFA12B5382F91E2260">Honghe</collectingCounty>
,
<collectingMunicipality id="60DFB3D4CC4851A75ED335B405723CDB">Lvchun</collectingMunicipality>
,
<location id="49CDA6BCAA18BD9D10201239B9585434" LSID="urn:lsid:plazi:treatment:71E0657FC3155D5095866048468ED52A:49CDA6BCAA18BD9D10201239B9585434" country="China" county="Honghe" municipality="Lvchun" name="Huanglianshan" stateProvince="Yunnan">Huanglianshan</location>
,
<quantity id="675AC252D0CA5AF807AE08934722FE71" metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">
<elevation id="C74D1DE2347CCF6FCE03A4B62309F39B" metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">380 m</elevation>
</quantity>
,
<collectingDate id="5484ED3A2C6CDC1F35FC8261DAAFB722" value="2018-04-15">2018.iv.15</collectingDate>
, leg.
<collectorName id="BB7539E75F6DB63F26BD2BBBBA5B09D2">Liang Wang</collectorName>
(
<specimenCount id="786AC565759DFDB2BAF508F22F151B91" type="male">1♂</specimenCount>
, CAU); same locality and collector as preceding,
<quantity id="1D2DE915E70B67E57B3B2025262D1D97" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation id="0356C69F2275BB6CDE8844F8104DD948" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<collectingDate id="2B2F8959DE250FBF54DCB86A77696F0A" value="2018-04-16">2018.iv.16</collectingDate>
(
<specimenCount id="0248CEBD49CAFF6D5477661C6DB33305" type="male">1♂</specimenCount>
, CAU); Xishuangbanna, Jinghong,
<location id="C7B5FB5B95388494C004FE80C095AB98" LSID="urn:lsid:plazi:treatment:71E0657FC3155D5095866048468ED52A:C7B5FB5B95388494C004FE80C095AB98" country="China" county="Honghe" municipality="Lvchun" name="Naban River Watershed National Nature Reserve" stateProvince="Yunnan">Naban River Watershed National Nature Reserve</location>
, Mandian Waterfall,
<quantity id="E41E8F2ADDBD192688B193C59CD8A4BE" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">
<elevation id="8AED9200877C11F2A40F29E344F0F273" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<collectingDate id="605B25F33756D49B4C5F445A2FDBE0A4" value="2020-11-25">2020.xi.25</collectingDate>
, leg.
<collectorName id="DBCE7766F2F93A98810D02360DA76F47">Liang Wang</collectorName>
(
<specimenCount id="64931CAB0FA58A2409779660C40473E4" type="male">1♂</specimenCount>
, CAU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="383A72D91CB87CFDA54BAA64B58F8750" pageId="0" pageNumber="329" type="diagnosis">
<paragraph id="38339FAC41AC406BE807A96C752A93FE" pageId="0" pageNumber="329">Diagnosis.</paragraph>
<paragraph id="1023AB886555B1A5A28E1F14E47A4C16" pageId="0" pageNumber="329">
Generally shiny blackish brown (Fig.
<figureCitation id="CA150B5A7B482A7136FB0134C4756E46" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male, habitus, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1168.104699.figure24" httpUri="https://binary.pensoft.net/fig/872799" pageId="0" pageNumber="329">24</figureCitation>
); antennal first flagellomere of male yellowish brown with apical half darkened, round, with a long, thick, sword-like process (Figs
<figureCitation id="0EF3E377098C0AC6AB68FE15D7B81A6B" captionStart="Figures 2529" captionStartId="F9" captionText="Figures 25 - 29. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male: 25 head, dorsal view 26 same, lateral view 27 left antenna, lateral view 28 thorax, dorsal view 29 wing. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1168.104699.figures25-29" httpUri="https://binary.pensoft.net/fig/872800" pageId="0" pageNumber="329">26</figureCitation>
,
<figureCitation id="B31D49A78F1D2165EF2E953389414934" captionStart="Figures 2529" captionStartId="F9" captionText="Figures 25 - 29. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male: 25 head, dorsal view 26 same, lateral view 27 left antenna, lateral view 28 thorax, dorsal view 29 wing. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1168.104699.figures25-29" httpUri="https://binary.pensoft.net/fig/872800" pageId="0" pageNumber="329">27</figureCitation>
); wing infumate, with large dark suffusion at apex and narrow median transverse band at level of dm-m (Fig.
<figureCitation id="82FCFBA099533ECCF32A88A43E0B6D29" captionStart="Figures 2529" captionStartId="F9" captionText="Figures 25 - 29. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male: 25 head, dorsal view 26 same, lateral view 27 left antenna, lateral view 28 thorax, dorsal view 29 wing. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1168.104699.figures25-29" httpUri="https://binary.pensoft.net/fig/872800" pageId="0" pageNumber="329">29</figureCitation>
); fore femur yellow with narrow brown ring subapically (Fig.
<figureCitation id="F5AE3939A6BABBF62C57C11E9628021C" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male, habitus, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1168.104699.figure24" httpUri="https://binary.pensoft.net/fig/872799" pageId="0" pageNumber="329">24</figureCitation>
); mid and hind femora yellowish brown, with apical half of mid femur and broad subbasal and subapical rings of hind femur dark brown (Fig.
<figureCitation id="405050E8840FCF65F04F51B7144AB532" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Strongylophthalmyia narwhal Evenhuis, 2020, non-type male, habitus, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1168.104699.figure24" httpUri="https://binary.pensoft.net/fig/872799" pageId="0" pageNumber="329">24</figureCitation>
); hind femur of male with one wart-like inner basal process; distiphallus approx. 1.3
<normalizedToken id="CA6011384AF6211735E1886BA9861389" originalValue="×">x</normalizedToken>
as long as phallapodeme, with small apical
<normalizedToken id="1DB92B5AF5C02F2EF22FE3C2A5D635A8" originalValue="“glans”">&quot;glans&quot;</normalizedToken>
(Figs
<figureCitation id="608AF448E66F846BBB8203DBDCCB1D08" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">32</figureCitation>
,
<figureCitation id="195B99BDA3FD09BAD940B70E7FA4AEAC" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">33</figureCitation>
).
</paragraph>
<caption id="F581A368C14BCF811B600139F5C3B84D" doi="10.3897/zookeys.1168.104699.figure24" httpUri="https://binary.pensoft.net/fig/872799" pageId="0" pageNumber="329" start="Figure 24" startId="F8">
<paragraph id="894B0778126CD5A015D6EB720AEBB95A" pageId="0" pageNumber="329">
<emphasis id="A1644AE3FC03476BDA1D1267CDD73702" bold="true" pageId="0" pageNumber="329">Figure 24.</emphasis>
<taxonomicName id="CEAA525408152742FE18738D1D4AD94A" authorityName="Evenhuis" authorityYear="2020" class="Insecta" family="Strongylophthalmyiidae" genus="Strongylophthalmyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylophthalmyia narwhal" order="Diptera" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="narwhal">
<emphasis id="0EC3440E3DCA05E3E553CF91F3CFD9BF" italics="true" pageId="0" pageNumber="329">Strongylophthalmyia narwhal</emphasis>
</taxonomicName>
Evenhuis, 2020, non-type male, habitus, lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="F9AB52DE9B54458F25226D77F18D7180" doi="10.3897/zookeys.1168.104699.figures25-29" httpUri="https://binary.pensoft.net/fig/872800" pageId="0" pageNumber="329" start="Figures 2529" startId="F9">
<paragraph id="88A1D43CC68EB47F5E492B991C416748" pageId="0" pageNumber="329">
<emphasis id="BB4864626D0603E8CF5641037A363FF5" bold="true" pageId="0" pageNumber="329">Figures 25-29.</emphasis>
<taxonomicName id="6B82F1B5AAFA28E0F7C0A5ABAD802FEE" authorityName="Evenhuis" authorityYear="2020" class="Insecta" family="Strongylophthalmyiidae" genus="Strongylophthalmyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylophthalmyia narwhal" order="Diptera" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="narwhal">
<emphasis id="4AD73918FB9E19E5C2EFAE3F36FDBA0B" italics="true" pageId="0" pageNumber="329">Strongylophthalmyia narwhal</emphasis>
</taxonomicName>
Evenhuis, 2020, non-type male:
<emphasis id="FAEF55365D5BA060EAF442A1CEA0ADE1" bold="true" pageId="0" pageNumber="329">25</emphasis>
head, dorsal view
<emphasis id="B141B0F4B32041CD195AA34C0384A112" bold="true" pageId="0" pageNumber="329">26</emphasis>
same, lateral view
<emphasis id="3C575FA5DA0349F182E088522DE720ED" bold="true" pageId="0" pageNumber="329">27</emphasis>
left antenna, lateral view
<emphasis id="5F3C37B1F5CBA69B922627527F254D87" bold="true" pageId="0" pageNumber="329">28</emphasis>
thorax, dorsal view
<emphasis id="4254C369464AAA0A80005F61B09FD1DD" bold="true" pageId="0" pageNumber="329">29</emphasis>
wing. Scale bars: 0.25 mm.
</paragraph>
</caption>
<caption id="C8646D09BCF39D381C9F0742A93822DC" doi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329" start="Figures 3033" startId="F10">
<paragraph id="C03A9FCFB9F097DF37195BEF8F57F6CB" pageId="0" pageNumber="329">
<emphasis id="9815AD04A7D68CA0D387A7A0A8FCCF21" bold="true" pageId="0" pageNumber="329">Figures 30-33.</emphasis>
<taxonomicName id="4380BEBC4DC0A0D5BAA5DCD9BDBAFADE" authorityName="Evenhuis" authorityYear="2020" class="Insecta" family="Strongylophthalmyiidae" genus="Strongylophthalmyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylophthalmyia narwhal" order="Diptera" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="narwhal">
<emphasis id="66483419064C6F9570357D8DE7C0069C" italics="true" pageId="0" pageNumber="329">Strongylophthalmyia narwhal</emphasis>
</taxonomicName>
Evenhuis, 2020, male genitalia:
<emphasis id="9C8C91D41EF2D5244A877C0123DF71EC" bold="true" pageId="0" pageNumber="329">30</emphasis>
external genitalia, posterior view
<emphasis id="0408E71FB9CC87F5FDA74A12D6C6C8BA" bold="true" pageId="0" pageNumber="329">31</emphasis>
same, lateral view
<emphasis id="DB7E5048C7898AA06E88C080DC6EBFA7" bold="true" pageId="0" pageNumber="329">32</emphasis>
internal genitalia, ventral view
<emphasis id="9A3C6D6F4568C29F8E9954943911AA65" bold="true" pageId="0" pageNumber="329">33</emphasis>
same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="59579D4FAFB9CE6D39CCB20F6853CA88" pageId="0" pageNumber="329" type="description">
<paragraph id="42E11E92F8B53B61D519DC4C568C166F" pageId="0" pageNumber="329">Description of male genitalia.</paragraph>
<paragraph id="C4B4D7D4797C707085658B5C7AD99A29" pageId="0" pageNumber="329">
Epandrium (Figs
<figureCitation id="2A72C9CE5B1325742750375569DDE9AF" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">30</figureCitation>
,
<figureCitation id="7E4456792B50D7CDF0D19F2A934E08D1" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">31</figureCitation>
) short and narrow, with long, dense setae. Surstylus (Figs
<figureCitation id="1463572E10062E722D951BC93EBA82AA" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">30</figureCitation>
,
<figureCitation id="F191731A50E3A8DBD78FBB5F461A277F" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">31</figureCitation>
) with short stout setae on inner distal surface. Cerci (Figs
<figureCitation id="51004679BED2454BE53CF3EB7F1A0D4D" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">30</figureCitation>
,
<figureCitation id="EA46B27ED5D90464076BB7913E3EFC89" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">31</figureCitation>
) narrow, elongate, finger-like, fused along length, with 1 long subapical seta and several short setae. Hypandrium (Figs
<figureCitation id="3C621CDA54B7511438AAD84DE13C67B2" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">32</figureCitation>
,
<figureCitation id="0EFE7D3B6BC35D3DF096F3FDD83369D0" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">33</figureCitation>
) narrowly rounded anteriorly, strongly arched medially, with one pair of bifid anterior lobes. Phallapodeme (Figs
<figureCitation id="8F59D64692D474A905B294DF37A00CFF" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">32</figureCitation>
,
<figureCitation id="EA971ED58C2C21304886CDF87731478A" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">33</figureCitation>
) short, slightly curved, distinctly shorter than distiphallus. Pregonite (Fig.
<figureCitation id="0B61AF8D18ACBC1EAB2C5E00A3DCB35A" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">32</figureCitation>
) very long, narrow, band-like, basally fused to inner surface of hypandrium. Phallic plate (Fig.
<figureCitation id="2D06000867D6B23C588B4ED3F5D6009B" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">33</figureCitation>
) divided into two articulating sclerites. Distiphallus (Figs
<figureCitation id="4A99DE255C53510D76DA1A766D5689EE" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">32</figureCitation>
,
<figureCitation id="0CAF514B54146D26713C9A32D3363D86" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">33</figureCitation>
) extremely long, nearly 1.3
<normalizedToken id="8B226650A0B5381F03A8D5253604BAEB" originalValue="×">x</normalizedToken>
as long as phallapodeme, unsegmented, with sclerotized bands and apical
<normalizedToken id="6783814B2340E4F8E0A76FC2D03E55FB" originalValue="“glans”">&quot;glans&quot;</normalizedToken>
, membrane microtrichose. Ejaculatory apodeme (Fig.
<figureCitation id="BCC00AA1413992AD8C89675F3CE64403" captionStart="Figures 3033" captionStartId="F10" captionText="Figures 30 - 33. Strongylophthalmyia narwhal Evenhuis, 2020, male genitalia: 30 external genitalia, posterior view 31 same, lateral view 32 internal genitalia, ventral view 33 same, lateral view. Abbreviations: cerc = cercus; dph = distiphallus; ea = ejaculatory apodeme; ep = epandrium; hyp = hypandrium; pgt = pregonite; phap = phallapodeme; pp = phallic plate; sur = surstylus. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1168.104699.figures30-33" httpUri="https://binary.pensoft.net/fig/872801" pageId="0" pageNumber="329">33</figureCitation>
) relatively large, slightly curved.
</paragraph>
</subSubSection>
<subSubSection id="F1489A88874DBA613B94E0D5FD486AF1" pageId="0" pageNumber="329" type="distribution">
<paragraph id="1A33A2E83E9CAEC71F7BBB1CBE565E2F" pageId="0" pageNumber="329">Distribution.</paragraph>
<paragraph id="D132E507359A664BE0E3DCDB59BF28BB" pageId="0" pageNumber="329">
China - Yunnan: Jinghong*, Lvchun* (Fig.
<figureCitation id="D5F2591446A2987B17164C21E8CC4A93" captionStart="Figure 56" captionStartId="F17" captionText="Figure 56. Known distribution of species of the Strongylophthalmyia coarctata subgroup in China and South Korea." figureDoi="10.3897/zookeys.1168.104699.figure56" httpUri="https://binary.pensoft.net/fig/872808" pageId="0" pageNumber="329">56</figureCitation>
). Thailand - Chiang Mai: Chiang Mai (
<bibRefCitation id="EA6974895E95BDB6427B1C736E41EEAC" author="Evenhuis, NL" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="329" pagination="1 - 7" refId="B2" refString="Evenhuis, NL, 2020. Mine's longer than yours: two new Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae) with remarkably long antennal processes. Bishop Museum Occasional Papers 132: 1 - 7" title="Mine's longer than yours: two new Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae) with remarkably long antennal processes." volume="132" year="2020">Evenhuis 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57A7640FDF0E79D677D1A0CD6F107137" pageId="0" pageNumber="329" type="remarks">
<paragraph id="BFE4449CDD3F2BD0BA21A003C90373B0" pageId="0" pageNumber="329">Remarks.</paragraph>
<paragraph id="06B056DA1CA1FD402968ED2218C2316D" pageId="0" pageNumber="329">
This species was described based on a male holotype from Chiang Mai, Thailand (
<bibRefCitation id="8488BF228B1456591C4690C60ACEA4AD" author="Evenhuis, NL" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="329" pagination="1 - 7" refId="B2" refString="Evenhuis, NL, 2020. Mine's longer than yours: two new Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae) with remarkably long antennal processes. Bishop Museum Occasional Papers 132: 1 - 7" title="Mine's longer than yours: two new Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae) with remarkably long antennal processes." volume="132" year="2020">Evenhuis 2020</bibRefCitation>
). Newly available specimens from Yunnan, China are identical in most of the features described by
<bibRefCitation id="6F31A392721B0BBEF359A38205675157" author="Evenhuis, NL" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="329" pagination="1 - 7" refId="B2" refString="Evenhuis, NL, 2020. Mine's longer than yours: two new Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae) with remarkably long antennal processes. Bishop Museum Occasional Papers 132: 1 - 7" title="Mine's longer than yours: two new Strongylophthalmyia Heller (Diptera: Strongylophthalmyiidae) with remarkably long antennal processes." volume="132" year="2020">Evenhuis (2020)</bibRefCitation>
, but differs in the following aspects: (i) gena blackish brown as general body color; (ii) both wings with narrow median transverse band at level of dm-m; (iii) legs yellow to yellowish brown, with basal half of fore coxa, narrow subapical ring of fore femur, apical half of mid femur (except extreme apex), broad subbasal and subapical rings of hind femur, mid and hind tibiae (except bases and apices) and tarsomeres 4 and 5 brown to dark brown.
</paragraph>
<paragraph id="5C99610C8740F52BF4FF14D7B36137ED" pageId="0" pageNumber="329">The chaetotaxy, which was not mentioned in the original description, can now be added as: head with 1 inner vertical seta, 1 outer vertical seta, 3 fronto-orbital setae, 1 ocellar seta and 1 postocellar seta; thorax with 1 anepisternal seta, 1 postpronotal seta, 2 notopleural setae, 2 dorsocentral setae, 2 posterior supra-alar setae and 1 scutellar seta; mid femur with row of long erect black setae. The male genitalia are described as above.</paragraph>
</subSubSection>
</treatment>
</document>