219 lines
19 KiB
XML
219 lines
19 KiB
XML
<document ID-DOI="10.3897/zookeys.16.229" ID-GBIF-Dataset="8d4d91e2-7ba2-41dd-a5d3-47a50b82837c" ID-ISSN="1313–2970" ID-Zenodo-Dep="576478" ID-ZooBank="D2CB2FAD-B1A2-4AA0-88A6-AB5BB6D735B8" checkinTime="1588007293194" checkinUser="plazi" docAuthor="Marusik, Yuri & Fet, Victor" docDate="2009" docId="591AA14AFFBD6734FF7B09305B4450DB" docLanguage="en" docName="ZK_article_2011.pdf" docOrigin="ZooKeys 16 (16)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@1b4312d" docStyleName="zookeys.2008.journal_article" docTitle="Hersiliola lindbergi Marusik & Fet 2009, sp. n." docType="treatment" docUuid="3609714D-77BE-4D7F-AA5F-34125B7693B9" docUuidSource="ZooBank" docVersion="9" lastPageNumber="85" masterDocId="A523D932FFB7673EFFF70B365850565C" masterDocTitle="A survey of East Palaearctic Hersiliola Thorell, 1870 (Araneae, Hersiliidae), with a description of three new genera" masterLastPageNumber="114" masterPageNumber="75" pageNumber="85" updateTime="1677608779287" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A survey of East Palaearctic Hersiliola Thorell, 1870 (Araneae, Hersiliidae), with a description of three new genera</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marusik, Yuri</mods:namePart>
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<mods:affiliation>IBPN RAS, Magadan,</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fet, Victor</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Department of Biological Sciences, Marshall University, Huntington, West Virginia, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-07-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>16</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>16</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>114</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.16.229</mods:identifier>
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<mods:identifier type="GBIF-Dataset">8d4d91e2-7ba2-41dd-a5d3-47a50b82837c</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576478</mods:identifier>
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||
<mods:identifier type="ZooBank">D2CB2FAD-B1A2-4AA0-88A6-AB5BB6D735B8</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792008" ID-GBIF-Taxon="163755998" ID-Zenodo-Dep="3792008" LSID="urn:lsid:zoobank.org:act:3609714D-77BE-4D7F-AA5F-34125B7693B9" httpUri="http://treatment.plazi.org/id/591AA14AFFBD6734FF7B09305B4450DB" lastPageNumber="85" pageId="10" pageNumber="85">
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<subSubSection box="[140,445,518,545]" pageId="10" pageNumber="85" type="nomenclature">
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||
<paragraph blockId="10.[140,965,518,616]" box="[140,445,518,545]" pageId="10" pageNumber="85">
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<heading box="[140,445,518,545]" fontSize="11" level="2" pageId="10" pageNumber="85" reason="3">
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<emphasis bold="true" box="[140,445,518,545]" pageId="10" pageNumber="85">
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<taxonomicName ID-CoL="3L3TY" authority="Marusik & Fet, 2009" authorityName="Marusik & Fet" authorityYear="2009" box="[140,374,518,545]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="lindbergi" status="sp. n.">
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<emphasis bold="true" box="[140,374,518,545]" italics="true" pageId="10" pageNumber="85">
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<emphasis box="[140,258,518,545]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Hersiliola</emphasis>
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lindbergi
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[380,445,519,545]" pageId="10" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="85" type="description">
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<paragraph blockId="10.[140,965,518,616]" box="[140,965,554,581]" pageId="10" pageNumber="85">
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<uri box="[140,965,554,581]" pageId="10" pageNumber="85">
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urn:lsid:zoobank.org:act:
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<uuid box="[416,965,554,581]" pageId="10" pageNumber="85">3609714D-77BE-4D7F-AA5F-34125B7693B9</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="10.[140,965,518,616]" lastBlockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">
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Figs
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<figureCitation box="[192,206,589,615]" captionStart-0="Figure 1" captionStart-1="Map 1" captionStartId-0="4.[140,212,1622,1644]" captionStartId-1="33.[139,187,789,811]" captionTargetBox-0="[147,1099,165,1602]" captionTargetBox-1="[138,1108,164,771]" captionTargetId-0="figure@4.[147,1100,165,1602]" captionTargetId-1="figure@33.[139,1108,165,769]" captionTargetPageId-0="4" captionTargetPageId-1="33" captionText-0="Figure 1. Habitus of females of Hersiliola sternbergsi sp. n. (1), H. esyunini sp. n. (2), H. lindbergi sp. n. (3) and H. afghanica (4)." captionText-1="Map 1. Distribution of Hersiliidae in Central Asia. ' Hersiliola afghanica; × Hersiliola esyunini; • Hersiliola foordi; · Hersiliola lindbergi; • Hersiliola sternbergsi; + Hersiliola xinjiangensis; ′ Duninia baehrae; ‵ Duninia rheimsae;. Deltshevia danovi; * Deltshevia gromovi; - Ovtsharenkoia pallida; -? identity uncertain" figureDoi-0="http://doi.org/10.5281/zenodo.3817414" figureDoi-1="http://doi.org/10.5281/zenodo.3817426" httpUri-0="https://zenodo.org/record/3817414/files/figure.png" httpUri-1="https://zenodo.org/record/3817426/files/figure.png" pageId="10" pageNumber="85">1</figureCitation>
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.3,
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<figureCitation box="[243,261,589,616]" captionStart="Figure 7" captionStartId="17.[140,212,1309,1331]" captionTargetBox="[140,1108,165,1275]" captionTargetId="figure@17.[140,1108,165,1276]" captionTargetPageId="17" captionText="Figure 7. Epigynes of Hersiliola versicolor (1), Hersiliola sp. (2), H. lindbergi sp. n. (3, 6), H. afghanica (4-5), H. sternbergsi sp. n. (7-9), H. esyunini sp. n. (10-12), and H. foordi sp. n. (13-14). 1-8, 10-11, 13-14 ventral 9, 12 dorsal. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3770581" httpUri="https://zenodo.org/record/3770581/files/figure.png" pageId="10" pageNumber="85">7</figureCitation>
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.5-6,
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<figureCitation box="[319,335,589,615]" captionStart="Figure 8" captionStartId="18.[140,212,1622,1644]" captionTargetBox="[140,1107,837,1599]" captionTargetId="figure@18.[140,1108,828,1602]" captionTargetPageId="18" captionText="Figure 8. Epigynes, ventral view, of Hersiliola afghanica (1), H. sternbergsi sp. n. (2), H. lindbergi sp. n. (3), H. esyunini sp. n. (4), H. foordi sp. n (5), Duninia baehrae sp. n. (6), and D. rheimsae sp. n. (7)." figureDoi="http://doi.org/10.5281/zenodo.3770583" httpUri="https://zenodo.org/record/3770583/files/figure.png" pageId="10" pageNumber="85">8</figureCitation>
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.3
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<emphasis bold="true" box="[140,314,659,685]" pageId="10" pageNumber="85">
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<emphasis box="[140,198,659,685]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Type</emphasis>
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material.
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608997611" country="Afghanistan" location="Paratyp" pageId="10" pageNumber="85" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
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<typeStatus box="[323,430,659,685]" pageId="10" pageNumber="85">Holotype</typeStatus>
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<specimenCount box="[440,455,660,686]" pageId="10" pageNumber="85" type="female">♀</specimenCount>
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with label “
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<specimenCount box="[608,635,660,686]" pageId="10" pageNumber="85" type="female">1♀</specimenCount>
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<location LSID="urn:lsid:plazi:treatment:591AA14AFFBD6734FF7B09305B4450DB:D46C4687FFBD6734FD7B09A25AB454F2" box="[652,740,660,686]" country="Afghanistan" name="Paratyp" pageId="10" pageNumber="85">Paratyp</location>
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,
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<accessionNumber box="[753,895,659,686]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF12996" pageId="10" pageNumber="85">SMF 12996</accessionNumber>
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, No. 15,
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<collectingCountry name="Afghanistan" pageId="10" pageNumber="85">Afghanistan</collectingCountry>
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”, in SMF.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="85" type="etymology">
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<paragraph blockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">
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<emphasis bold="true" box="[185,320,730,756]" pageId="10" pageNumber="85">
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<emphasis box="[185,315,730,756]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Etymology</emphasis>
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.
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</emphasis>
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The species name is a patronym honoring its collector, Dr. Knut Lindberg (
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<date box="[206,335,765,792]" pageId="10" pageNumber="85" value="1892" valueMax="1962">1892-1962</date>
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) of Lund,
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<collectingCountry box="[462,551,765,791]" name="Sweden" pageId="10" pageNumber="85">Sweden</collectingCountry>
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, who conducted an important expedition to
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<collectingCountry name="Afghanistan" pageId="10" pageNumber="85">Afghanistan</collectingCountry>
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between
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<date box="[356,534,800,827]" pageId="10" pageNumber="85" value="1957" valueMax="1960">1957 and 1960</date>
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.
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</paragraph>
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<paragraph blockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">
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<emphasis bold="true" box="[185,243,835,861]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Note</emphasis>
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.
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<bibRefCitation author="Roewer CF" box="[259,430,835,862]" pageId="10" pageNumber="85" pagination="3 - 53" refId="ref19918" refString="Roewer CF (1960) Araneae Orthognathae, Haplogynae und Entelegynae. Solifugen und Opilioniden. (Contribution a l'etude de la faune d'Afghanistan 23). Goteborgs Kunglige Vetenskaps och Vitterhets Samhalles Handlingar, B 8 (7): 3 - 53." type="journal article" year="1960">Roewer (1960)</bibRefCitation>
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in his description of
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<taxonomicName authorityName="Marusik & Fet" authorityYear="2009" box="[677,820,835,861]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="afghanica">
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<emphasis box="[677,820,835,861]" italics="true" pageId="10" pageNumber="85">H. afghanica</emphasis>
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</taxonomicName>
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indicated only one adult specimen, the
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<typeStatus box="[303,402,871,897]" pageId="10" pageNumber="85">holotype</typeStatus>
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female. All
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<typeStatus box="[539,645,871,897]" pageId="10" pageNumber="85">paratypes</typeStatus>
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were juveniles. Labels of the
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<typeStatus box="[978,1077,871,897]" pageId="10" pageNumber="85">holotype</typeStatus>
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of
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<taxonomicName authorityName="Marusik & Fet" authorityYear="2009" box="[140,282,906,932]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="afghanica">
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<emphasis box="[140,282,906,932]" italics="true" pageId="10" pageNumber="85">H. afghanica</emphasis>
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</taxonomicName>
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(GNM) and the
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<accessionNumber box="[481,618,906,932]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF12996" pageId="10" pageNumber="85">SMF 12996</accessionNumber>
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“
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<typeStatus box="[636,735,906,932]" pageId="10" pageNumber="85">paratype</typeStatus>
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” were written by the same hand, probably by Roewer.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="85" type="description">
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<paragraph blockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">
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<emphasis bold="true" box="[185,330,976,1002]" pageId="10" pageNumber="85">
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<emphasis box="[185,325,976,1002]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Description</emphasis>
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.
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</emphasis>
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Male unknown. Female: Total length 3.6. Carapace 1.5 long, 1.7 wide, femur I 2.5 long, femur I/carapace length ratio 1.66. General colouration light yellowbrown. Carapace without pattern but with dark margins. Abdomen with a wide heart spot and one transverse stripe; sides with brown bands. Legs without distinct annulations. Chelicerae and maxillae without strong setae. Coxae IV separated by less than one diameter.
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</paragraph>
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<paragraph blockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">Epigyne as in Figs 7.5-6, 8.3, with septum as wide as epigyne openings; median plate not separated from the epigynal plate; translucent spermathecae small and round, close to each other, their diameter approximately equals width of epigyne openings, windows absent; spermathecae separated from the opening by three diameters; insemination ducts long, making six coils around fertilization duct; the upper loop welldeveloped; accessory glands small, digitiform.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="85" type="diagnosis">
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<paragraph blockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">
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<emphasis bold="true" box="[185,312,1363,1389]" pageId="10" pageNumber="85">
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<emphasis box="[185,307,1363,1389]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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<taxonomicName authority="Marusik & Fet, 2009" authorityName="Marusik & Fet" authorityYear="2009" box="[321,457,1363,1389]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="lindbergi" status="sp. n.">
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<emphasis box="[321,457,1363,1389]" italics="true" pageId="10" pageNumber="85">H. lindbergi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[467,530,1364,1390]" pageId="10" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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clearly differs from other species by the very long insemination ducts making six coils and an apical loop. It can be also distinguished by small round spermathecae, the shape of the septum and the epigyne openings.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="85" type="discussion">
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<paragraph blockId="10.[140,1108,659,1672]" pageId="10" pageNumber="85">
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<emphasis bold="true" box="[185,324,1469,1495]" pageId="10" pageNumber="85">
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<emphasis box="[185,319,1469,1495]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Comments</emphasis>
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.
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</emphasis>
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There is a certain possibility that
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<taxonomicName authority="Marusik & Fet, 2009" authorityName="Marusik & Fet" authorityYear="2009" box="[722,859,1469,1495]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="lindbergi" status="sp. n.">
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<emphasis box="[722,859,1469,1495]" italics="true" pageId="10" pageNumber="85">H. lindbergi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[868,932,1470,1496]" pageId="10" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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belongs to another genus. Unlike in
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<taxonomicName box="[400,566,1504,1530]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="maculullata">
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<emphasis box="[400,566,1504,1530]" italics="true" pageId="10" pageNumber="85">H. maculullata</emphasis>
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</taxonomicName>
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,
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<taxonomicName box="[581,709,1504,1530]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="afganica">
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<emphasis box="[581,709,1504,1530]" italics="true" pageId="10" pageNumber="85">H. afganica</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName=": Benoit" authorityYear="1974" box="[723,832,1505,1530]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="simoni">
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<emphasis box="[723,832,1505,1530]" italics="true" pageId="10" pageNumber="85">H. simoni</emphasis>
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</taxonomicName>
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,
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<taxonomicName authority="Marusik & Fet, 2009" authorityName="Marusik & Fet" authorityYear="2009" box="[846,995,1504,1530]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="sternbergsi" status="sp. n.">
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<emphasis box="[846,995,1504,1530]" italics="true" pageId="10" pageNumber="85">H. sternbergsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1002,1065,1505,1531]" pageId="10" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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,
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<taxonomicName baseAuthorityName="Blackwall" baseAuthorityYear="1865" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="versicolor">
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<emphasis italics="true" pageId="10" pageNumber="85">H. versicolor</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authority="Marusik & Fet, 2009" authorityName="Marusik & Fet" authorityYear="2009" box="[303,489,1539,1565]" class="Arachnida" family="Hersiliidae" genus="Duninia" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="baehrae" status="sp. n.">
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<emphasis box="[303,489,1539,1565]" italics="true" pageId="10" pageNumber="85">Duninia baehrae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[496,560,1540,1566]" pageId="10" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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, the median plate in this species is not separated from the epigynal plates and has no accessory sclerites or windows. The position of
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<taxonomicName authority="Marusik & Fet, 2009" authorityName="Marusik & Fet" authorityYear="2009" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="lindbergi" status="sp. n.">
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<emphasis italics="true" pageId="10" pageNumber="85">H. lindbergi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[244,305,1610,1636]" pageId="10" pageNumber="85" rank="species">sp. n.</taxonomicNameLabel>
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cannot be confirmed until a male is found.
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</paragraph>
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</subSubSection>
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<subSubSection box="[185,788,1645,1672]" pageId="10" pageNumber="85" type="distribution">
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<paragraph blockId="10.[140,1108,659,1672]" box="[185,788,1645,1672]" pageId="10" pageNumber="85">
|
||
<emphasis bold="true" box="[185,344,1645,1671]" pageId="10" pageNumber="85">
|
||
<emphasis box="[185,339,1645,1671]" inLineHeading="true" pageId="10" pageNumber="85" reason="1">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry box="[351,490,1645,1671]" name="Afghanistan" pageId="10" pageNumber="85">Afghanistan</collectingCountry>
|
||
, without a precise locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |