treatments-xml/data/59/06/0E/59060E37938150B0A55547C6757449BA.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.907.36563" ID-GBIF-Dataset="69422977-aa4d-4fe3-a450-8ef54d6fa905" ID-PMC="PMC7002455" ID-Pensoft-Pub="1313-2970-907-1" ID-Pensoft-UUID="9920267E73CF5E00B644DED1F101D965" ID-PubMed="32063727" ID-ZooBank="DBF339E5EBBC461994388359C769473F" ModsDocID="1313-2970-907-1" checkinTime="1580398733216" checkinUser="pensoft" docAuthor="Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. &amp; Nuss, Matthias" docDate="2020" docId="59060E37938150B0A55547C6757449BA" docLanguage="en" docName="ZooKeys 907: 1-99" docOrigin="ZooKeys 907" docSource="http://dx.doi.org/10.3897/zookeys.907.36563" docTitle="Hoploscopa sumatrensis Leger &amp; Nuss 2020, sp. nov." docType="treatment" docUuid="22F33777-64CB-4E37-9AB2-41A52020ABEB" docUuidSource="ZooBank" docVersion="4" id="9920267E73CF5E00B644DED1F101D965" lastPageNumber="1" masterDocId="9920267E73CF5E00B644DED1F101D965" masterDocTitle="Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding" masterLastPageNumber="99" masterPageNumber="1" pageNumber="1" updateTime="1668168367651" updateUser="ExternalLinkService">
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<mods:title>Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Leger, Theo</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kehlmaier, Christian</mods:namePart>
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<mods:namePart>Vairappan, Charles S.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nuss, Matthias</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>907</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.907.36563</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.907.36563</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-907-1</mods:identifier>
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<treatment ID-GBIF-Taxon="161926059" LSID="urn:lsid:zoobank.org:act:22F33777-64CB-4E37-9AB2-41A52020ABEB" httpUri="http://treatment.plazi.org/id/59060E37938150B0A55547C6757449BA" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="59060E37-9381-50B0-A555-47C6757449BA" authority="Léger &amp; Nuss" authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa sumatrensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sumatrensis" status="sp. nov.">
Hoploscopa sumatrensis 
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; Nuss
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">Figs 36</figureCitation>
<figureCitation captionStart="Figures 70–75" captionStartId="F9" captionText="Figures 70 - 75. Male genitalia features of Hoploscopa. 70 Hoploscopa nauticorum, TL 725 ♂ 71 Hoploscopa anacantha sp. nov., holotype, TL 759 ♂ 72 Hoploscopa kelama sp. nov., paratype, TL 511 ♂ 73 Hoploscopa ignitamaculae sp. nov., paratype, TL 545 ♂ 74 Hoploscopa albomaculata sp. nov., paratype, TL 363 ♂ 75 Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂." figureDoi="10.3897/zookeys.907.36563.figures70-75" httpUri="https://binary.pensoft.net/fig/375764" pageId="0" pageNumber="1">, 75</figureCitation>
<figureCitation captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">, 83</figureCitation>
<figureCitation captionStart="Figures 115–118" captionStartId="F19" captionText="Figures 115 - 118. Female genitalia features of Hoploscopa. 115 Hoploscopa sumatrensis sp. nov., paratype, TL 530 ♀ 116 Hoploscopa mallyi sp. nov., paratype, TL 513 ♀ 117 Hoploscopa gracilis sp. nov., paratype, TL 527 ♀ 118 Hoploscopa agtuuganonensis sp. nov., holotype, TL 616 ♀." figureDoi="10.3897/zookeys.907.36563.figures115-118" httpUri="https://binary.pensoft.net/fig/375774" pageId="0" pageNumber="1">, 115</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂, with labels: &quot;Sumatra-Holzweg | 25km SSW-Pematangsiantar | 
<normalizedToken originalValue="Straße">Strasse</normalizedToken>
nach Prapat [road to Prapat] | 6.-26.i.1995, leg. A. Kallies&quot;; &quot;coll[ection]. M. Nuss | Geschenk 2000 | Museum 
<normalizedToken originalValue="für">fuer</normalizedToken>
Tier- | kunde Dresden&quot;; &quot;DNA voucher | Lepidoptera | date: xi.2018 | MTD8260 | [vertically written:] DNA | -voucher&quot;; &quot;TL755 | ♂&quot;. Deposited in MTD.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 3 ♂, 4 ♀. Indonesia. 3 ♂ (DNA vouchers MTD9119, MTD9129 &amp; MTD9121 and genitalia on slides TL761 ♂, TL763 ♂, TL538 ♂), 3 ♀ (DNA vouchers MTD LEP3205, MTD8257, MTD8262 &amp; genitalia on slides TL524 ♀, TL752 ♀, TL757 ♀), same data as holotype; 1 ♀ (DNA voucher MTD7880 &amp; genitalia on slide TL670 ♀), Sumatra, Asahan, Huta Padang, 1990, leg. E. W. Diehl (MTD).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">4 ♀, 2 ♀ (DNA vouchers MTD LEP3202, MTD 8258 &amp; genitalia on slides TL530 ♀, TL753 ♀), same data as holotype; 1 ♀ (DNA voucher MTD LEP3210 &amp; genitalia on slide TL522 ♀), same data as holotype except 23.viii.1989, leg. E. W. Diehl; 1 ♀ (DNA voucher MTD8261 &amp; genitalia on slide TL756 ♀), same data as holotype except 13.ii.1996 (MTD).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The forewings of 
<taxonomicName lsidName="H. sumatrensis" pageId="0" pageNumber="1" rank="species" species="sumatrensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. sumatrensis</emphasis>
</taxonomicName>
sp. nov. display a pale yellow crescent-shaped median discoidal stigma filled with reddish brown, as well as a postmedian area broadly suffused with pale yellow. In male genitalia, the gnathos projection is slender, ca. 4/5 of uncus length, with a tongue-shaped apex, and the juxta has a rounded base and a notched apex. In female genitalia, the short and slender ductus bursae bent before corpus and the pear-shaped corpus bursae with small straight thorn resemble those of other 
<taxonomicName authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
</taxonomicName>
species, e.g., 
<taxonomicName lsidName="H. danaoensis" pageId="0" pageNumber="1" rank="species" species="danaoensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. danaoensis</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="H. parvimacula" pageId="0" pageNumber="1" rank="species" species="parvimacula">
<emphasis italics="true" pageId="0" pageNumber="1">H. parvimacula</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa pangrangoensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pangrangoensis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa pangrangoensis</emphasis>
</taxonomicName>
sp. nov. (q.v.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Antennae dorsally with brown scales. Proboscis pale yellow to brown. Maxillary palpi brown, base and inner side pale yellow. Labial palpi brown, ventral base and inner side pale yellow.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
(Fig. 
<figureCitation captionStart="Figures 34–45" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">36</figureCitation>
). Collar pale yellow. Forewing length: 9-10 mm (♂ &amp; ♀); forewing ground colour brown; small basal dark brown dash distally pale yellow; base of dorsum with patch of pale yellow scales; cubital reddish brown fascia running up to median area; median discoidal stigma reddish brown, edged with crescent-shaped pale yellow patch; postmedian patch reddish brown, with pale yellow blotch at costa; subterminal line broad, pale yellow; subterminal field tawny to brown; fringes brown, with pale yellow dots. Hindwing pale yellow. Forelegs bronze. Midlegs with femur brown; tibia pale yellow to brown; tarsi bronze. Hindlegs brown; tibia pale yellow speckled with brown; tarsi bronze.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Fig. 
<figureCitation captionStart="Figures 82–86" captionStartId="F11" captionText="Figures 82 - 86. Male genitalia and abdomen features of Hoploscopa. 82 Hoploscopa jubata sp. nov., paratype, TL 542 ♂ 83 Tympanal organs of Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂ 84 Male tympanal organs of Hoploscopa metacrossa, TL 472 ♂ 85 Sternite VIII of Hoploscopa sp. near isarogensis, TL 626 ♂ 86 Sternite VIII of Hoploscopa metacrossa, TL 443 ♂. Abbreviations: tb (tympanic bridge), td (tympanic drum), tdp (tympanic depression), tm (tympanum), tr (transverse ridge)." figureDoi="10.3897/zookeys.907.36563.figures82-86" httpUri="https://binary.pensoft.net/fig/375766" pageId="0" pageNumber="1">83</figureCitation>
). Male sternum A8 posterior margin bilobed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 4) (Fig. 
<figureCitation captionStart="Figures 70–75" captionStartId="F9" captionText="Figures 70 - 75. Male genitalia features of Hoploscopa. 70 Hoploscopa nauticorum, TL 725 ♂ 71 Hoploscopa anacantha sp. nov., holotype, TL 759 ♂ 72 Hoploscopa kelama sp. nov., paratype, TL 511 ♂ 73 Hoploscopa ignitamaculae sp. nov., paratype, TL 545 ♂ 74 Hoploscopa albomaculata sp. nov., paratype, TL 363 ♂ 75 Hoploscopa sumatrensis sp. nov., paratype, TL 538 ♂." figureDoi="10.3897/zookeys.907.36563.figures70-75" httpUri="https://binary.pensoft.net/fig/375764" pageId="0" pageNumber="1">75</figureCitation>
). Uncus medially broad, narrowed at apical 1/4, apex tongue-shaped. Gnathos projection slender, ca. 4/5 of uncus length, apex tongue-shaped. Valva ventral margin straight, bent dorsad on distal 1/4, dorsal margin convex, apex pointed. Juxta base rounded, distal half narrowed, apex notched. Saccus triangular, pointed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 4) (Fig. 
<figureCitation captionStart="Figures 115–118" captionStartId="F19" captionText="Figures 115 - 118. Female genitalia features of Hoploscopa. 115 Hoploscopa sumatrensis sp. nov., paratype, TL 530 ♀ 116 Hoploscopa mallyi sp. nov., paratype, TL 513 ♀ 117 Hoploscopa gracilis sp. nov., paratype, TL 527 ♀ 118 Hoploscopa agtuuganonensis sp. nov., holotype, TL 616 ♀." figureDoi="10.3897/zookeys.907.36563.figures115-118" httpUri="https://binary.pensoft.net/fig/375774" pageId="0" pageNumber="1">115</figureCitation>
). Anterior apophyses with dorsal tip at posterior 1/3. Antrum sclerotisation short. Ductus bursae short, bent before corpus opening. Corpus bursae large, pear-shaped, with sclerotisation between thorn and corpus opening and a median sclerotised band. Thorn small, straight, with small dents.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Known from North Sumatra (Indonesia).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="dna barcoding">
<paragraph pageId="0" pageNumber="1">DNA barcoding.</paragraph>
<paragraph pageId="0" pageNumber="1">Two MOTUs are found in morphologically identical specimens from the same locality. The K2P-distance between the two MOTUs is 4.1-6%. The first MOTU, which is the one of the type material, shows an intraspecific variation of 0.7%. The second MOTU is found in three females (samples BC MTD LEP01422, MTD8258 and MTD8261) and shows an intraspecific divergence of 0.6%.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Named after the island of Sumatra where the species is encountered.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">Future examination of male specimens from the second lineage will help to determine whether or not it represents a separate species.</paragraph>
<caption doi="10.3897/zookeys.907.36563.figures52-57" httpUri="https://binary.pensoft.net/fig/375761" pageId="0" pageNumber="1" start="Figures 52–57" startId="F6">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 52-57.</emphasis>
Male genitalia features of 
<taxonomicName authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="1">52</emphasis>
<taxonomicName authorityName="Nuss" authorityYear="1998" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa luteomacula" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luteomacula">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa luteomacula</emphasis>
</taxonomicName>
Nuss, holotype, GU prep. Nuss 744 ♂ 
<emphasis bold="true" pageId="0" pageNumber="1">53</emphasis>
<taxonomicName baseAuthorityName="Rothschild" baseAuthorityYear="1915" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa obliqua" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa obliqua</emphasis>
</taxonomicName>
, holotype, 
<taxonomicName family="Pyralidae" lsidName="" pageId="0" pageNumber="1" rank="family">Pyralidae</taxonomicName>
Brit. Mus. Slide N° BMNH20252 ♂ (phallus with juxta attached) 
<emphasis bold="true" pageId="0" pageNumber="1">54</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa gombongi" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gombongi">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa gombongi</emphasis>
</taxonomicName>
sp. nov., holotype, TL653 ♂ 
<emphasis bold="true" pageId="0" pageNumber="1">55</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa tonsepi" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tonsepi">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa tonsepi</emphasis>
</taxonomicName>
sp. nov., paratype, TL661 ♂ 
<emphasis bold="true" pageId="0" pageNumber="1">56</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa niveofascia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveofascia">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa niveofascia</emphasis>
</taxonomicName>
sp. nov., holotype, TL442 ♂ 
<emphasis bold="true" pageId="0" pageNumber="1">57</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa marijoweissae" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marijoweissae">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa marijoweissae</emphasis>
</taxonomicName>
sp. nov., holotype, TL710 ♂.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>