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<mods:title id="69EAA7E2DB7F11CB69F16B4F034098B8">A monographic catalogue on the systematics and phylogeny of the South American iguanian lizard family Liolaemidae (Squamata, Iguania)</mods:title>
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Boulenger
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<taxonomicName id="7BB630F1FFA961016CFBEEAFFD7B8C9E" authority="Boulenger 1885 a: 152" authorityName="Boulenger" authorityYear="1885" box="[151,692,1456,1479]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="variety" species="pictus" variety="major">
<emphasis id="8EC29760FFA961016CFBEEAFFE838C9E" box="[151,332,1456,1479]" italics="true" pageId="25" pageNumber="26">Liolaemus pictus</emphasis>
var.
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(
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locality: “
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”).
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<treatmentCitation id="3D176D63FFA961016CFBEECFFDE38CBE" author="Hulse, A. C." box="[151,556,1488,1511]" page="204" pageId="25" pageNumber="26" year="1979">
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<emphasis id="8EC29760FFA961016CFBEECFFEBE8CBE" box="[151,369,1488,1511]" italics="true" pageId="25" pageNumber="26">Liolaemus capillitas</emphasis>
<bibRefCitation id="D8273683FFA961016D15EECFFDE38CBE" author="Hulse" box="[377,556,1488,1511]" firstAuthor="Hulse" page="204" pageId="25" pageNumber="26" pagination="203 - 209" refId="ref61768" refString="Hulse, A. C. (1979) A new Liolaemus (Sauria, Iguanidae) from the high Andes of Argentina, with ecological comments. Annals of the Carnegie Museum, 48, 203 - 209." type="journal article" year="1979">Hulse 1979: 204</bibRefCitation>
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(synonymy
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<bibRefCitation id="D8273683FFA961016E88EECFFCA48CBE" author="Nunez, H." box="[740,875,1488,1511]" pageId="25" pageNumber="26" pagination="28 - 34" refId="ref67402" refString="Nunez, H. (2004) Cambios taxonomicos para la herpetofauna de Argentina, Bolivia y Chile. Noticiario Mensual Museo Nacional de Historia Natural de Chile, 353, 28 - 34." type="journal article" year="2004">Núñez 2004</bibRefCitation>
);
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<bibRefCitation id="D8273683FFA961016F13EECFFB748CBE" author="McCoy &amp; Censky" box="[895,1211,1488,1511]" firstAuthor="McCoy" page="326" pageId="25" pageNumber="26" pagination="317 - 330" refId="ref65705" refString="McCoy, C. J. &amp; Censky, E. J. (1982) Herpetological type specimens in Carnegie Museum of Natural History. Annals of the Carnegie Museum, 51, 317 - 330." type="journal article" year="1982">McCoy &amp; Censky 1982: 326</bibRefCitation>
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;
<treatmentCitation id="3D176D63FFA9610168ABEECFFA588CBE" author="Vanzolini, P. E." box="[1223,1431,1488,1511]" page="13" pageId="25" pageNumber="26" year="1986">
<bibRefCitation id="D8273683FFA9610168ABEECFFA588CBE" author="Vanzolini" box="[1223,1431,1488,1511]" firstAuthor="Vanzolini" page="13" pageId="25" pageNumber="26" pagination="1 - 25" refId="ref73558" refString="Vanzolini, P. E. (1986) Addenda and corrigenda to the Catalogue of Neotropical Squamata. Smithsonian Herpetological Information Service, 70, 1 - 25." type="journal article" year="1986">Vanzolini 1986: 13</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="D8273683FFA961016CABEEEFFE4D8F5E" author="Laurent" box="[199,386,1520,1543]" firstAuthor="Laurent" page="31" pageId="25" pageNumber="26" pagination="1 - 33" refId="ref64066" refString="Laurent, R. F. (1992) On some overlooked species of the genus Liolaemus Wiegmann (Reptilia-Tropiduridae) from Peru. Breviora, 494, 1 - 33." type="journal article" year="1992">Laurent 1992: 31</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016DE3EEEFFDE48F5E" author="Cei" box="[399,555,1520,1543]" firstAuthor="Cei" page="280" pageId="25" pageNumber="26" refId="ref54875" refString="Cei, J. M. (1993) Reptiles del noroeste, nordeste y este de la Argentina. Herpetofauna de las selvas subtropicales, Puna y Pampas. Museo Regionale di Scienze Naturali di Torino, Monografie 14, 949 pp." type="book" year="1993">Cei 1993: 280</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016E5BEEEFFCC78F5E" author="Etheridge" box="[567,776,1520,1543]" firstAuthor="Etheridge" page="33" pageId="25" pageNumber="26" pagination="1 - 34" refId="ref59230" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995: 33</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016F78EEEFFB928F5E" author="Etheridge &amp; Espinoza" box="[788,1117,1520,1543]" firstAuthor="Etheridge" page="5" pageId="25" pageNumber="26" pagination="1 - 64" refId="ref59521" refString="Etheridge, R &amp; Espinoza, R. E. (2000) Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semiannotated bibliography. Smithsonian Herpetological Information Service, 126, 1 - 64." type="journal article" year="2000">Etheridge &amp; Espinoza 2000: 5</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016806EEEFFAD68F5E" author="Lobo" box="[1130,1305,1520,1543]" firstAuthor="Lobo" page="139" pageId="25" pageNumber="26" pagination="137 - 150" refId="ref64638" refString="Lobo, F. (2001) A phylogenetic analysis of lizards of the Liolaemus chiliensis group (Iguania: Tropiduridae). Herpetological Journal, 11, 137 - 150." type="journal article" year="2001">Lobo 2001: 139</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016949EEEFFEFA8F7E" author="Avila, L. J. &amp; Perez, C. H. F. &amp; Morando, M." etAl="et al." firstAuthor="Avila" page="534" pageId="25" pageNumber="26" pagination="532 - 543" refId="ref51745" refString="Avila, L. J., Perez, C. H. F. &amp; Morando, M. (2003) A new species of Liolaemus (Squamata: Iguania: Liolaemidae) from northwestern Patagonia (Neuquen, Argentina). Herpetologica, 59, 532 - 543." type="journal article" year="2003">
Avila
<emphasis id="8EC29760FFA961016909EEEFFA538F5E" box="[1381,1436,1520,1543]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2003: 534
</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016D2CED0FFDFB8F7E" author="Chebez" box="[320,564,1552,1575]" etAl="et al." firstAuthor="Chebez" page="29" pageId="25" pageNumber="26" refId="ref56330" refString="Chebez, J. C., Rey, N. R. &amp; Williams, J. D. (2005) Reptiles de los parques nacionales de la Argentina. Literatura of Latin America, Buenos Aires, 37 pp." type="book" year="2005">
Chebez
<emphasis id="8EC29760FFA961016DFBED0FFE028F7E" box="[407,461,1552,1575]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2005: 29
</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016E53ED0FFD2A8F7E" author="Sura" box="[575,741,1552,1575]" firstAuthor="Sura" page="428" pageId="25" pageNumber="26" refId="ref73002" refString="Sura, P. 2005. Encyclopedia of extant amphibians and reptiles, Wydawnictwo Fundacja, Nowy Sacz, 544 pp." type="book" year="2005">Sura 2005: 428</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016E9CED0FFC3E8F7E" author="Diaz &amp; Lobo" box="[752,1009,1552,1575]" firstAuthor="Diaz" page="264" pageId="25" pageNumber="26" pagination="261 - 274" refId="ref57471" refString="Diaz, J. M. &amp; Lobo, F. (2006) Historical biogeography of a clade of Liolaemus (Iguania: Liolaemidae) based on ancestral areas and dispersal-vicariance analysis (DIVA). Papeis Avulsos de Zoologia, 46, 261 - 274." type="journal article" year="2006">Díaz &amp; Lobo 2006: 264</bibRefCitation>
;
<bibRefCitation id="D8273683FFA961016F90ED0FFB6C8F7E" author="Diaz" box="[1020,1187,1552,1575]" firstAuthor="Diaz" page="114" pageId="25" pageNumber="26" pagination="105 - 118" refId="ref57434" refString="Diaz, J. M. (2007) Reptilia, Iguania, Liolaeminae, Liolaemus, Puna, Prepuna, and mountain ranges, northwestern Argentina. Check List, 3, 105 - 118." type="journal article" year="2007">Díaz 2007: 114</bibRefCitation>
.
</paragraph>
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<taxonomicName id="7BB630F1FFA961016CFBED2FFDCE8F1E" authority="Nunez 2004: 32" authorityName="Nunez" authorityPageNumber="32" authorityYear="2004" box="[151,513,1584,1607]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="major">
<emphasis id="8EC29760FFA961016CFBED2FFE828F1E" box="[151,333,1584,1607]" italics="true" pageId="25" pageNumber="26">Liolaemus major</emphasis>
<bibRefCitation id="D8273683FFA961016D3FED2FFDCE8F1E" author="Nunez, H." box="[339,513,1584,1607]" pageId="25" pageNumber="26" pagination="28 - 34" refId="ref67402" refString="Nunez, H. (2004) Cambios taxonomicos para la herpetofauna de Argentina, Bolivia y Chile. Noticiario Mensual Museo Nacional de Historia Natural de Chile, 353, 28 - 34." type="journal article" year="2004">Núñez 2004: 32</bibRefCitation>
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.
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<paragraph id="BC094B72FFA961016CFBED68FA9B8EDB" blockId="25.[151,1436,1655,1922]" pageId="25" pageNumber="26">
<emphasis id="8EC29760FFA961016CFBED68FE898FC8" bold="true" box="[151,326,1655,1681]" pageId="25" pageNumber="26">Observations:</emphasis>
After studying the collection of
<taxonomicName id="7BB630F1FFA961016EA3ED67FCAE8FCB" authorityName="Frost &amp; Etheridge" authorityYear="1989" box="[719,865,1656,1682]" class="Reptilia" family="Liolaemidae" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="family">Liolaemidae</taxonomicName>
<typeStatus id="630DF5D0FFA961016F06ED67FC678FCB" box="[874,936,1656,1682]" pageId="25" pageNumber="26">types</typeStatus>
housed in the British Museum of Natural History of London,
<bibRefCitation id="D8273683FFA961016DEFEDBFFDE98FE3" author="Nunez, H." box="[387,550,1696,1722]" pageId="25" pageNumber="26" pagination="28 - 34" refId="ref67402" refString="Nunez, H. (2004) Cambios taxonomicos para la herpetofauna de Argentina, Bolivia y Chile. Noticiario Mensual Museo Nacional de Historia Natural de Chile, 353, 28 - 34." type="journal article" year="2004">Núñez (2004)</bibRefCitation>
concluded that the
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specimens originally named
<taxonomicName id="7BB630F1FFA9610168F3EDBFFF128FB8" authorityName="Boulenger" authorityYear="1885" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="variety" species="pictus" variety="major">
<emphasis id="8EC29760FFA9610168F3EDBFFAA78FE0" box="[1183,1384,1696,1721]" italics="true" pageId="25" pageNumber="26">Liolaemus pictus</emphasis>
var.
<emphasis id="8EC29760FFA961016CFBEDD7FF128FB8" box="[151,221,1736,1761]" italics="true" pageId="25" pageNumber="26">major</emphasis>
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by
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are the species currently known as
<taxonomicName id="7BB630F1FFA961016FE9EDD7FBDE8FB8" authorityName="Hulse" authorityYear="1979" box="[901,1041,1736,1761]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="capillitas">
<emphasis id="8EC29760FFA961016FE9EDD7FBDE8FB8" box="[901,1041,1736,1761]" italics="true" pageId="25" pageNumber="26">L. capillitas</emphasis>
</taxonomicName>
, from
<collectingCountry id="C4A10BE2FFA961016833EDD7FB1B8FBB" box="[1119,1236,1736,1762]" name="Argentina" pageId="25" pageNumber="26">Argentina</collectingCountry>
(
<bibRefCitation id="D8273683FFA96101688FEDD7FA9B8FBB" author="Cei" box="[1251,1364,1736,1762]" firstAuthor="Cei" pageId="25" pageNumber="26" refId="ref54875" refString="Cei, J. M. (1993) Reptiles del noroeste, nordeste y este de la Argentina. Herpetofauna de las selvas subtropicales, Puna y Pampas. Museo Regionale di Scienze Naturali di Torino, Monografie 14, 949 pp." type="book" year="1993">Cei 1993</bibRefCitation>
). This
<typeStatus id="630DF5D0FFA961016CFBEDEFFF068E53" box="[151,201,1776,1802]" pageId="25" pageNumber="26">type</typeStatus>
material is labeled as collected from “
<collectingCountry id="C4A10BE2FFA961016EF0EDEFFD148E53" box="[668,731,1776,1802]" name="Chile" pageId="25" pageNumber="26">Chili</collectingCountry>
” (
<collectingCountry id="C4A10BE2FFA961016E94EDEFFCF28E53" box="[760,829,1776,1802]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
). Nevertheless, when
<bibRefCitation id="D8273683FFA96101682AEDEFFB258E53" author="Nunez, H." box="[1094,1258,1776,1802]" pageId="25" pageNumber="26" pagination="28 - 34" refId="ref67402" refString="Nunez, H. (2004) Cambios taxonomicos para la herpetofauna de Argentina, Bolivia y Chile. Noticiario Mensual Museo Nacional de Historia Natural de Chile, 353, 28 - 34." type="journal article" year="2004">Núñez (2004)</bibRefCitation>
reassessed the taxonomic status of these individuals, he did not propose a new
<typeStatus id="630DF5D0FFA961016FE0EC07FC718E6B" box="[908,958,1816,1842]" pageId="25" pageNumber="26">type</typeStatus>
locality in accordance with the distribution of its synonymous
<taxonomicName id="7BB630F1FFA961016DC2EC5FFDF38E00" authorityName="Hulse" authorityYear="1979" box="[430,572,1856,1881]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="capillitas">
<emphasis id="8EC29760FFA961016DC2EC5FFDF38E00" box="[430,572,1856,1881]" italics="true" pageId="25" pageNumber="26">L. capillitas</emphasis>
</taxonomicName>
(now
<taxonomicName id="7BB630F1FFA961016EE5EC5FFC0E8E03" authority="Nunez 2004" authorityName="Nunez" authorityYear="2004" box="[649,961,1856,1882]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="subSpecies" species="major" subSpecies="fide">
<emphasis id="8EC29760FFA961016EE5EC5FFCEC8E00" box="[649,803,1856,1881]" italics="true" pageId="25" pageNumber="26">L. major fide</emphasis>
<bibRefCitation id="D8273683FFA961016F40EC5FFC0E8E03" author="Nunez, H." box="[812,961,1856,1882]" pageId="25" pageNumber="26" pagination="28 - 34" refId="ref67402" refString="Nunez, H. (2004) Cambios taxonomicos para la herpetofauna de Argentina, Bolivia y Chile. Noticiario Mensual Museo Nacional de Historia Natural de Chile, 353, 28 - 34." type="journal article" year="2004">Núñez 2004</bibRefCitation>
</taxonomicName>
). Consequently, although
<taxonomicName id="7BB630F1FFA96101689FEC5FFA968E00" authorityName="Nunez" authorityYear="2004" box="[1267,1369,1856,1881]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="major">
<emphasis id="8EC29760FFA96101689FEC5FFA968E00" box="[1267,1369,1856,1881]" italics="true" pageId="25" pageNumber="26">L. major</emphasis>
</taxonomicName>
(=
<taxonomicName id="7BB630F1FFA9610169E8EC5FFECB8ED8" authorityName="Hulse" authorityYear="1979" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="capillitas">
<emphasis id="8EC29760FFA9610169E8EC5FFECB8ED8" italics="true" pageId="25" pageNumber="26">L. capillitas</emphasis>
</taxonomicName>
) would be endemic to
<collectingCountry id="C4A10BE2FFA961016E7DEC77FD4A8EDB" box="[529,645,1896,1922]" name="Argentina" pageId="25" pageNumber="26">Argentina</collectingCountry>
(see
<bibRefCitation id="D8273683FFA961016EAFEC77FCFE8EDB" author="Cei" box="[707,817,1896,1922]" firstAuthor="Cei" pageId="25" pageNumber="26" refId="ref54875" refString="Cei, J. M. (1993) Reptiles del noroeste, nordeste y este de la Argentina. Herpetofauna de las selvas subtropicales, Puna y Pampas. Museo Regionale di Scienze Naturali di Torino, Monografie 14, 949 pp." type="book" year="1993">Cei 1993</bibRefCitation>
), its
<typeStatus id="630DF5D0FFA961016F06EC77FC538EDB" box="[874,924,1896,1922]" pageId="25" pageNumber="26">type</typeStatus>
locality is still recognized as “
<collectingCountry id="C4A10BE2FFA96101696FEC77FA8A8EDB" box="[1283,1349,1896,1922]" name="Chile" pageId="25" pageNumber="26">Chili</collectingCountry>
”.
</paragraph>
</subSubSection>
</treatment>
</document>