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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.661.10867" ID-GBIF-Dataset="5e74ad60-e9d3-4707-af5b-7b67f9648c02" ID-PMC="PMC5539367" ID-Pensoft-Pub="1313-2970-661-61" ID-PubMed="28769603" ID-ZBK="68925924A20A4D30BA1660CF33F7CF21" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-661-61" ModsDocOrigin="ZooKeys 661" ModsDocTitle="Two psammophilic noctuids newly associated with beach plum, Prunusmaritima (Rosaceae): The Dune Noctuid (Sympistisriparia) and Coastal Heathland Cutworm (Abagrotisbenjamini) in Northeastern North America (Lepidoptera, Noctuidae)" checkinTime="1489572798561" checkinUser="pensoft" docAuthor="Goldstein, Paul Z. &amp; Nelson, Michael W." docDate="2017" docId="17F1C7E4FF7F2E6A6A5E7DD6ACA4DDAB" docLanguage="en" docName="ZooKeys 661: 61-89" docOrigin="ZooKeys 661" docSource="http://dx.doi.org/10.3897/zookeys.661.10867" docTitle="Abagrotis benjamini Franclemont, stat. rev." docType="treatment" docVersion="4" lastPageNumber="70" masterDocId="FF8EFFE4FF84FFB1FF89FF9FFFFC4858" masterDocTitle="Two psammophilic noctuids newly associated with beach plum, Prunusmaritima (Rosaceae): The Dune Noctuid (Sympistisriparia) and Coastal Heathland Cutworm (Abagrotisbenjamini) in Northeastern North America (Lepidoptera, Noctuidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="69" updateTime="1668164299231" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two psammophilic noctuids newly associated with beach plum, Prunusmaritima (Rosaceae): The Dune Noctuid (Sympistisriparia) and Coastal Heathland Cutworm (Abagrotisbenjamini) in Northeastern North America (Lepidoptera, Noctuidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goldstein, Paul Z.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nelson, Michael W.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>661</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>89</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.661.10867</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.661.10867</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-661-61</mods:identifier>
<mods:identifier type="ZBK">68925924A20A4D30BA1660CF33F7CF21</mods:identifier>
<mods:identifier type="ZooBank">68925924A20A4D30BA1660CF33F7CF21</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127886011" LSID="urn:lsid:plazi:treatment:17F1C7E4FF7F2E6A6A5E7DD6ACA4DDAB" httpUri="http://treatment.plazi.org/id/17F1C7E4FF7F2E6A6A5E7DD6ACA4DDAB" lastPageId="9" lastPageNumber="70" pageId="8" pageNumber="69">
<subSubSection pageId="8" pageNumber="69" type="nomenclature">
<paragraph pageId="8" pageNumber="69">
<taxonomicName authority="Franclemont" class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis benjamini" order="Lepidoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="benjamini">Abagrotis benjamini Franclemont</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="69">stat. rev.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="70" pageId="8" pageNumber="69" type="materials_examined">
<paragraph pageId="8" pageNumber="69">Material examined.</paragraph>
<paragraph pageId="8" pageNumber="69">
Type material (
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis crumbi" order="Lepidoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="subspecies" species="crumbi" subspecies="benjamini">Abagrotis crumbi benjamini</taxonomicName>
Franclemont, 1955)
</paragraph>
<paragraph pageId="9" pageNumber="70">
<pageBreakToken pageId="9" pageNumber="70" start="start">Holotype</pageBreakToken>
. (Fig. 16a, b). New York: E.N.Y. 7.17.00; Collection BrklynMus; Acc. 17,151; ♂ Gen. #1203 F.H.B. 17 July 1935; HOLOTYPE
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis crumbi" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="subspecies" species="crumbi" subspecies="benjamini">Abagrotis crumbi benjamini</taxonomicName>
J.G. Franclemont.
</paragraph>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 16. Holotype of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" infraspecific-rank="race" kingdom="Animalia" lsidName="Abagrotis crumbi" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" race="benjamini" rank="race" species="crumbi">Abagrotis crumbi race benjamini</taxonomicName>
Franclemont. a Dorsal b Ventral. Scale bars: 1 cm (Photos B. Proshek).
</paragraph>
</caption>
<paragraph pageId="9" pageNumber="70">
Slide labels [Paratypes]: (Fig. 22a, b) USNM 36838 ♂; ♂ Gen. #1202
<taxonomicName class="Insecta" family="Buprestidae" genus="Lampra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lampra crumbi" order="Coleoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="crumbi">Lampra crumbi</taxonomicName>
n. sp. var. E.N.Y. A.C. Weeks 7.17.00 Coll. Brklyn Mus. F.H.B. 17 July 1935. Paratypes (2 of 5♂, 3 of 3♀ designated by Franclemont): New York: East New York (2♂, 1♀). Connecticut: East River (1♀). Location uncertain: &quot;Collection J.B. Smith&quot; (1♀).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="70" type="materials_examined">
<paragraph pageId="9" pageNumber="70">Other material examined.</paragraph>
<paragraph pageId="9" pageNumber="70">(not including ~13 with DNA barcodes pending)</paragraph>
<paragraph pageId="9" pageNumber="70">
MASSACHUSETTS (28♂, 18♀): Edgartown (1♂, 2♀): Katama Plain
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.35787">41°21.472'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.520935">70°31.256'W</geoCoordinate>
, 19 June 2010 (1♂); Wintucket Cove frost bottom 24 July 1991 (1♀); Felix Neck Wildlife Sanctuary 7 VII 1989 P.Z. Goldstein, leg. (1♀). Aquinnah (22♂, 15♀): East Pasture Road,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.341568">41°20.494'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.78747">70°47.248'W</geoCoordinate>
, @ UV Trap: 30 June 2013 (1♂, 1♀); 1 July 2011 (1♂); 15 July 2011 (1♂); off Moshup Trail/E. of Moshup Trail, 41.324894, -70.810698 / 41°19'29.6'N, 70°48'38.5'W (7♂, 7♀): 17 vii 2003 (3♂, 1♀) [USNM Dissection #s 148015♂; 148029♂ (Fig. 23); 148031♂]; 17 July 2015 (1♀) Barcode of Life DNA voucher specimen SampleID CCDB-28975-E01 BOLD Proc ID LNAUU 1474-15; USNM Dissection #148022♀; 31 July 2014 (1♀); 17 July 2003 (3♂, 4♀);
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.323032">41°19.382'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-70.811">70°48.66'W</geoCoordinate>
, 14 August 2002 (1♂); off Moshup Trail, above
<normalizedToken originalValue="Zacks">Zack's</normalizedToken>
Cliffs,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.323067">41°19.384'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.81091">70°48.655'W</geoCoordinate>
, (5♂, 1♀): 10 Sept 2002 (3♂); 8 sept 2011 [USNM Dissection 148028♂] (1♂); 12 Sept 2010 (1♂, 1♀); Lobsterville, 41°20'49.7'N, 70°46'54.5'W, (7♂, 6♀): 22 Sept 2012 148026♀ (1♀); 23 August 2012 (2♀); 5 July 2015 148021♂ USNMENT01203938 (1♂); 30 Aug 2014 148019♀ USNMENT01203918 (1♀) (Fig. 17a, b); 3 October 2014 148020♀USNMENT01203909 (1♀); 19 July 2012 (3♂, 1♀); 22 Sept 2012 (3♂). West Tisbury (5♂, 1♀): Long Point Wildlife Refuge, maritime heathland E. of Long Cove Pond
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.353867">41°21.232'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.631065">70°37.864'W</geoCoordinate>
, (2♂, 1♀) 29 June 2010 @ UV trap (USNM Dissection #s 148030♂; 148056♀; 148016♂);
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.359516">41°21.571'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.60052">70°36.031'W</geoCoordinate>
, 14 June 2010 @ UV trap (1♂);
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.359535">41°21.572'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.60053">70°36.032'W</geoCoordinate>
, 29 August 2010 @ UV trap (1♂); Maritime heath W. of Long Cove Pond
<geoCoordinate direction="north" orientation="latitude" precision="1" value="41.35275">41°21.165'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-70.6388">70°38.328'W</geoCoordinate>
, 10 September 2010 @ UV trap (1♂).
</paragraph>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 17.
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis benjamini" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="benjamini">Abagrotis benjamini</taxonomicName>
Franclemont, Aquinnah, MA. P.Z. Goldstein. a Dorsal b Ventral. Scale bars: 1 cm (Photos B. Proshek).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="9" pageNumber="70" type="diagnosis">
<paragraph pageId="9" pageNumber="70">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="70">
Forewing ground color ranging from pale gray brown to, more commonly, a rusty brown, occasionally reddish brown (paprika colored), never dark; both FW and soma more heavily peppered with black scales than in
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis nefascia" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="nefascia">Abagrotis nefascia</taxonomicName>
; both fasciae and subterminal line (as dashes on underside, scalloping on upperside) less conspicuous than in
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis nefascia" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="nefascia">Abagrotis nefascia</taxonomicName>
. Scobinate patch on vesicae of male genitalia with 12 or fewer denticles, at least two of them markedly larger than others. Female bursa with a single signum.
</paragraph>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 18.
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis benjamini" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="benjamini">Abagrotis benjamini</taxonomicName>
Franclemont, Dartmouth, MA. M.W. Nelson. a Dorsal b Ventral. Scale bars: 1 cm (Photos B. Proshek).
</paragraph>
</caption>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 19. Everted vesica of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis nefascia" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="nefascia">Abagrotis nefascia</taxonomicName>
, Holotype. Scale bar: 1 mm.
</paragraph>
</caption>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 20. Everted vesica of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis nefascia" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="nefascia">Abagrotis nefascia</taxonomicName>
, Yakima, WA. USNM dissection #s 148023, 148024. Scale bar: 1 mm.
</paragraph>
</caption>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 21. Genitalia of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis crumbi" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="crumbi">Abagrotis crumbi</taxonomicName>
(=
<taxonomicName lsidName="nefascia" pageId="9" pageNumber="70" rank="species" species="nefascia">nefascia</taxonomicName>
). a Holotype, ♂. USNM dissection #36819 b Paratype, ♂. USNM dissection #36821. Scale bars: 1 mm.
</paragraph>
</caption>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 22. Genitalia of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis crumbi" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="subspecies" species="crumbi" subspecies="benjamini">Abagrotis crumbi benjamini</taxonomicName>
(Paratype). USNM dissection #36838. a Clasper b Phallus.
</paragraph>
</caption>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 23. Everted vesica of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis benjamini" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="benjamini">Abagrotis benjamini</taxonomicName>
, Aquinnah, MA. a USNM dissection #148029 b Everted vesica of
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis benjamini" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="benjamini">Abagrotis benjamini</taxonomicName>
, Dartmouth, MA. USNM dissection #148067. Cf. Figs 9d (larva), 11(adult), and 18a, b (adult).
</paragraph>
</caption>
<caption pageId="9" pageNumber="70">
<paragraph pageId="9" pageNumber="70">
Figure 24. Female genitalia,
<taxonomicName class="Insecta" family="Noctuidae" genus="Abagrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abagrotis nefascia" order="Lepidoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="nefascia">Abagrotis nefascia</taxonomicName>
USNM dissection #148066. Note presence of two signa, one reduced.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>