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<document id="7A588051029679EED20DA1376E19C79A" ID-DOI="10.5852/ejt.2020.695" ID-GBIF-Dataset="c2e17975-440e-40f9-8b37-f078f9aadf1b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3973377" ID-ZooBank="DC24EAB0-DCF5-44A8-B1A0-82BF25D280C2" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1596646124793" checkinUser="valdenar" docAuthor="Dietrich, Christopher H., Magalhães, Raysa Brito de &amp; Takiya, Daniela M." docDate="2020" docId="03A587FCFF82B52BFDFD80E6FBFE5646" docLanguage="en" docName="ejt-695_dietrich_magalhaes_takiya.pdf" docOrigin="European Journal of Taxonomy 695" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Betsileonas Kirkaldy 1903" docType="treatment" docVersion="11" lastPageNumber="10" masterDocId="FF9CFF84FF8BB521FF848643FFFA5425" masterDocTitle="Revision of the endemic Malagasy leafhopper tribe Platyjassini (Hemiptera: Cicadellidae: Iassinae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="10" updateTime="1733753971925" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A693C28AA6CF644485C84DA52632D752">Revision of the endemic Malagasy leafhopper tribe Platyjassini (Hemiptera: Cicadellidae: Iassinae)</mods:title>
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<mods:namePart id="0749B8CCA6365A6D77B885C426DBB5A0">Dietrich, Christopher H.</mods:namePart>
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<mods:affiliation id="0FE279A13CCABC4CAF5BABD0435D07E1">Illinois Natural History Survey, Prairie Research Institute, University of Illinois, 1816 S. Oak St., Champaign, IL 61820, USA.</mods:affiliation>
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<mods:namePart id="445D08C6E5EC039B131E4D94CD2ADD6D">Magalhães, Raysa Brito de</mods:namePart>
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<mods:affiliation id="8B1D86D0CAC7C5DA119825855E6FAC3B">Laboratório de Entomologia, Departamento de Zoologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ Brazil.</mods:affiliation>
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<mods:affiliation id="2101D4F7988C930FD63C454EA5D6AAD5">Laboratório de Entomologia, Departamento de Zoologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ Brazil.</mods:affiliation>
<mods:nameIdentifier id="8F56A33F895277910D790FA00076ADD1" type="email">takiya@acd.ufrj.br</mods:nameIdentifier>
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<mods:date id="7A983E3CCA584D1FCFB7C9145252BCD6">2020</mods:date>
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<treatment id="03A587FCFF82B52BFDFD80E6FBFE5646" ID-DOI="http://doi.org/10.5281/zenodo.4342214" ID-GBIF-Taxon="166095895" ID-Zenodo-Dep="4342214" LSID="urn:lsid:plazi:treatment:03A587FCFF82B52BFDFD80E6FBFE5646" httpUri="http://treatment.plazi.org/id/03A587FCFF82B52BFDFD80E6FBFE5646" lastPageId="10" lastPageNumber="10" pageId="9" pageNumber="10" scope_family="Cicadellidae" scope_infraOrder="Cicadomorpha" scope_order="Hemiptera" scope_subFamily="Iassinae" scope_subOrder="Auchenorrhyncha" scope_superFamily="Membracoidea" scope_tribe="Platyjassini">
<subSubSection id="C3166561FF82B528FDFD80E6FC41529A" box="[633,955,1701,1728]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BB336EAFF82B528FDFD80E6FC41529A" blockId="9.[633,955,1701,1728]" box="[633,955,1701,1728]" pageId="9" pageNumber="10">
<heading id="D0FB8186FF82B528FDFD80E6FC41529A" box="[633,955,1701,1728]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<taxonomicName id="4C0C4D69FF82B528FDFD80E6FC41529A" ID-CoL="8HDVN" authority="Kirkaldy, 1903" authorityName="Kirkaldy" authorityYear="1903" box="[633,955,1701,1728]" class="Insecta" family="Cicadellidae" genus="Betsileonas" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF8FF82B528FDFD80E6FCFE52E5" bold="true" box="[633,772,1701,1728]" italics="true" pageId="9" pageNumber="10">Betsileonas</emphasis>
<bibRefCitation id="EF9D4B1BFF82B528FC8F80E6FC41529A" author="Kirkaldy G. W." box="[779,955,1701,1727]" firstAuthor="Kirkaldy" journalOrPublisher="Wiener Entomologische Zeitung" pageId="9" pageNumber="10" pagination="13 - 17" part="22" refId="ref49131" refString="Kirkaldy G. W. 1903. Einige neue und wenig bekannte Rhynchoten. Wiener Entomologische Zeitung 22: 13 - 17." title="Einige neue und wenig bekannte Rhynchoten" type="journal article" year="1903">Kirkaldy, 1903</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3166561FF82B528FF3980A5FAFC5362" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8BB336EAFF82B528FF3980A5FECC5300" blockId="9.[189,1398,1766,1863]" pageId="9" pageNumber="10">
<taxonomicName id="4C0C4D69FF82B528FF3980A5FE055325" authority="Stal, 1864 b: 67" authorityName="Stal" authorityPageNumber="67" authorityYear="1864" box="[189,511,1766,1793]" class="Insecta" family="Cicadellidae" genus="Thaumastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF8FF82B528FF3980A5FEB15325" box="[189,331,1766,1792]" italics="true" pageId="9" pageNumber="10">Thaumastus</emphasis>
<bibRefCitation id="EF9D4B1BFF82B528FED480A5FE335325" author="Stal C." box="[336,457,1766,1793]" firstAuthor="Stal" journalOrPublisher="Annales de la Societe entomologique de France" pageId="9" pageNumber="10" pagination="47 - 68" part="4" refId="ref49378" refString="Stal C. 1864. Hemiptera nonnulla nova vel minus cognita. Annales de la Societe entomologique de France (Ser. 4) 4: 47 - 68." title="Hemiptera nonnulla nova vel minus cognita" type="journal article" year="1864">Stål, 1864</bibRefCitation>
b: 67
</taxonomicName>
(
<typeStatus id="54B78848FF82B528FD8980A4FDBB5324" box="[525,577,1767,1793]" pageId="9" pageNumber="10">type</typeStatus>
species:
<taxonomicName id="4C0C4D69FF82B528FD2E80A5FBB95324" ID-CoL="8JVKJ" authority="Blanchard, 1840" authorityName="Blanchard" authorityYear="1840" box="[682,1091,1766,1793]" class="Insecta" family="Cicadellidae" genus="Ledra" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis id="B978EAF8FF82B528FD2E80A5FC835325" box="[682,889,1766,1792]" italics="true" pageId="9" pageNumber="10">Ledra marmorata</emphasis>
<bibRefCitation id="EF9D4B1BFF82B528FCFB80A5FBB95324" accessDate="2020-07-09" author="Blanchard E." box="[895,1091,1766,1793]" firstAuthor="Blanchard" journalOrPublisher="Dumenil, Paris" pageId="9" pageNumber="10" publicationUrl="https://www.biodiversitylibrary.org/page/39159721" refId="ref48834" refString="Blanchard E. 1840. Histoire naturelle des Insectes: Orthopteres, Nevropteres, Hemipteres, Hymenopteres, Lepidopteres et Dipteres. P. Dumenil, Paris. Available from https: // www. biodiversitylibrary. org / page / 39159721 [accessed 9 Jul. 2020]." title="Histoire naturelle des Insectes: Orthopteres, Nevropteres, Hemipteres, Hymenopteres, Lepidopteres et Dipteres. P" type="url" year="1840">Blanchard, 1840</bibRefCitation>
</taxonomicName>
; nec
<taxonomicName id="4C0C4D69FF82B528FBFF80A5FED35300" authority="Martens, 1860" authorityName="Martens" authorityYear="1860" class="Insecta" family="Cicadellidae" genus="Thaumastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF8FF82B528FBFF80A5FAF35325" box="[1147,1289,1766,1792]" italics="true" pageId="9" pageNumber="10">Thaumastus</emphasis>
Martens, 1860
</taxonomicName>
).
</paragraph>
<paragraph id="8BB336EAFF82B528FF39816EFAFC5362" blockId="9.[189,1398,1766,1863]" box="[189,1286,1837,1863]" pageId="9" pageNumber="10">
<treatmentCitation id="0AAD10FBFF82B528FF39816EFDD05362" author="Kirkaldy G. W." box="[189,554,1837,1863]" page="13" pageId="9" pageNumber="10" year="1903">
<taxonomicName id="4C0C4D69FF82B528FF39816EFDD05362" ID-CoL="8HDVN" authority="Kirkaldy, 1903: 13" authorityName="Kirkaldy" authorityPageNumber="13" authorityYear="1903" box="[189,554,1837,1863]" class="Insecta" family="Cicadellidae" genus="Betsileonas" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF8FF82B528FF39816EFEBE5362" box="[189,324,1837,1863]" italics="true" pageId="9" pageNumber="10">Betsileonas</emphasis>
<bibRefCitation id="EF9D4B1BFF82B528FEC8816EFDD05362" author="Kirkaldy G. W." box="[332,554,1837,1863]" firstAuthor="Kirkaldy" journalOrPublisher="Wiener Entomologische Zeitung" pageId="9" pageNumber="10" pagination="13 - 17" part="22" refId="ref49131" refString="Kirkaldy G. W. 1903. Einige neue und wenig bekannte Rhynchoten. Wiener Entomologische Zeitung 22: 13 - 17." title="Einige neue und wenig bekannte Rhynchoten" type="journal article" year="1903">Kirkaldy, 1903: 13</bibRefCitation>
</taxonomicName>
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, nom. nov. (
<typeStatus id="54B78848FF82B528FD3E816EFD145362" box="[698,750,1837,1863]" pageId="9" pageNumber="10">type</typeStatus>
species:
<taxonomicName id="4C0C4D69FF82B528FCDE816EFB035362" ID-CoL="8JVKJ" authority="Blanchard, 1840" authorityName="Blanchard" authorityYear="1840" box="[858,1273,1837,1863]" class="Insecta" family="Cicadellidae" genus="Ledra" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis id="B978EAF8FF82B528FCDE816EFBD65362" box="[858,1068,1837,1863]" italics="true" pageId="9" pageNumber="10">Ledra marmorata</emphasis>
<bibRefCitation id="EF9D4B1BFF82B528FBB7816EFB035362" accessDate="2020-07-09" author="Blanchard E." box="[1075,1273,1837,1863]" firstAuthor="Blanchard" journalOrPublisher="Dumenil, Paris" pageId="9" pageNumber="10" publicationUrl="https://www.biodiversitylibrary.org/page/39159721" refId="ref48834" refString="Blanchard E. 1840. Histoire naturelle des Insectes: Orthopteres, Nevropteres, Hemipteres, Hymenopteres, Lepidopteres et Dipteres. P. Dumenil, Paris. Available from https: // www. biodiversitylibrary. org / page / 39159721 [accessed 9 Jul. 2020]." title="Histoire naturelle des Insectes: Orthopteres, Nevropteres, Hemipteres, Hymenopteres, Lepidopteres et Dipteres. P" type="url" year="1840">Blanchard, 1840</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3166561FF82B52BFF39812CFBFE5646" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BB336EAFF82B528FF39812CFED553AC" blockId="9.[189,303,1903,1929]" box="[189,303,1903,1929]" pageId="9" pageNumber="10">
<heading id="D0FB8186FF82B528FF39812CFED553AC" bold="true" box="[189,303,1903,1929]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B978EAF8FF82B528FF39812CFED553AC" bold="true" box="[189,303,1903,1929]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336EAFF82B52BFF3981DEFBFE5646" blockId="9.[189,1398,1949,2011]" lastBlockId="10.[189,1399,267,611]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
This monotypic genus was placed in Penthimiini by
<bibRefCitation id="EF9D4B1BFF82B528FCCC81DEFC115392" author="Evans J. W." box="[840,1003,1949,1975]" firstAuthor="Evans" journalOrPublisher="Transactions of the Entomological Society of London" pageId="9" pageNumber="10" pagination="105 - 271" part="98" refId="ref48943" refString="Evans J. W. 1947. A natural classification of leaf-hoppers (Jassoidea, Homoptera). Transactions of the Entomological Society of London 98: 105 - 271. https: // doi. org / 10.1111 / j. 1365 - 2311.1947. tb 01054. x" title="A natural classification of leaf-hoppers (Jassoidea, Homoptera)" type="journal article" year="1947">Evans (1947)</bibRefCitation>
but later returned to Ledrini by
<bibRefCitation id="EF9D4B1BFF82B528FF398182FEA753FE" author="Evans J. W." box="[189,349,1984,2011]" firstAuthor="Evans" journalOrPublisher="Memoires de l'Institut scientifique de Madagascar Tananarive (E" pageId="9" pageNumber="10" pagination="87 - 137" part="4" refId="ref48993" refString="Evans J. W. 1953. Les Cicadellidae de Madagascar. Memoires de l'Institut scientifique de Madagascar Tananarive (E) 4: 87 - 137." title="Les Cicadellidae de Madagascar" type="journal article" year="1953">Evans (1953)</bibRefCitation>
.
<bibRefCitation id="EF9D4B1BFF82B528FEE38182FD9B53FF" author="Jones J. R. &amp; Deitz L. L." box="[359,609,1984,2011]" firstAuthor="Jones" journalOrPublisher="Zootaxa" pageId="9" pageNumber="10" pagination="1 - 120" part="2186" refId="ref49082" refString="Jones J. R. &amp; Deitz L. L. 2009. Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae). Zootaxa 2186 (1): 1 - 120. https: // doi. org / 10.11646 / zootaxa. 2186.1.1" title="Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae)" type="journal article" year="2009">Jones &amp; Deitz (2009)</bibRefCitation>
treated it as a genus
<emphasis id="B978EAF8FF82B528FCD78182FC0E53FF" box="[851,1012,1984,2010]" italics="true" pageId="9" pageNumber="10">incertae sedis</emphasis>
because their morphology-based phylogenetic analysis failed to support its inclusion in
<taxonomicName id="4C0C4D69FF81B52BFCC38748FC495500" authorityName="Fairmaire" authorityYear="1855" box="[839,947,267,293]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subFamily" subFamily="Ledrinae">Ledrinae</taxonomicName>
. The genus is known only from a few very old museum specimens and was not encountered in the recent extensive terrrestrial arthropod inventory of
<collectingCountry id="F31B767AFF81B52BFEDF8711FE115549" box="[347,491,338,364]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
, suggesting that
<taxonomicName id="4C0C4D69FF81B52BFD3C8711FCC55549" authorityName="Kirkaldy" authorityYear="1903" box="[696,831,338,364]" class="Insecta" family="Cicadellidae" genus="Betsileonas" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAF8FF81B52BFD3C8711FCC55549" box="[696,831,338,364]" italics="true" pageId="10" pageNumber="11">Betsileonas</emphasis>
</taxonomicName>
is now either extremely rare or extinct. Males remain unknown, but several characters mentioned or illustrated for the genus by
<bibRefCitation id="EF9D4B1BFF81B52BFBFF8736FA8C55AA" author="Jones J. R. &amp; Deitz L. L." box="[1147,1398,373,399]" firstAuthor="Jones" journalOrPublisher="Zootaxa" pageId="10" pageNumber="11" pagination="1 - 120" part="2186" refId="ref49082" refString="Jones J. R. &amp; Deitz L. L. 2009. Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae). Zootaxa 2186 (1): 1 - 120. https: // doi. org / 10.11646 / zootaxa. 2186.1.1" title="Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae)" type="journal article" year="2009">Jones &amp; Deitz (2009)</bibRefCitation>
support its placement in
<taxonomicName id="4C0C4D69FF81B52BFE5F87DBFD9F5597" authority="Dietrich &amp; Magalhães &amp; Takiya, 2020" authorityName="Dietrich &amp; Magalhães &amp; Takiya" authorityYear="2020" box="[475,613,408,434]" class="Insecta" family="Cicadellidae" genus="Platyjassini" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">Platyjassini</taxonomicName>
, including the spatulate head with dorsal ocelli well separated from the crown apex, the rugose texture on the crown, the anterolateral extensions of the vertex overlapping the eyes, and the distally fused RP and MA in the hindwing. The ovipositor structure also resembles that of some other
<taxonomicName id="4C0C4D69FF81B52BFEEF8441FE0C5639" authority="Dietrich &amp; Magalhães &amp; Takiya, 2020" authorityName="Dietrich &amp; Magalhães &amp; Takiya" authorityYear="2020" box="[363,502,514,540]" class="Insecta" family="Cicadellidae" genus="Platyjassini" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">Platyjassini</taxonomicName>
. Known from a single species, the genus is easily distinguished from other
<taxonomicName id="4C0C4D69FF81B52BFF398466FEBE561A" authority="Dietrich &amp; Magalhães &amp; Takiya, 2020" authorityName="Dietrich &amp; Magalhães &amp; Takiya" authorityYear="2020" box="[189,324,549,575]" class="Insecta" family="Cicadellidae" genus="Platyjassini" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">Platyjassini</taxonomicName>
by its very large size (length
<quantity id="4CF49B0FFF81B52BFD188466FCE4561A" box="[668,798,549,576]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="10" pageNumber="11" unit="mm" value="17.0" valueMax="19.0" valueMin="15.0">1519 mm</quantity>
), the head being distinctly wider than the pronotum and its marmorate dorsal color pattern (see
<bibRefCitation id="EF9D4B1BFF81B52BFD3E840AFC5D5647" author="Jones J. R. &amp; Deitz L. L." box="[698,935,584,611]" firstAuthor="Jones" journalOrPublisher="Zootaxa" pageId="10" pageNumber="11" pagination="1 - 120" part="2186" refId="ref49082" refString="Jones J. R. &amp; Deitz L. L. 2009. Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae). Zootaxa 2186 (1): 1 - 120. https: // doi. org / 10.11646 / zootaxa. 2186.1.1" title="Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae)" type="journal article" year="2009">Jones &amp; Deitz 2009</bibRefCitation>
: pl. 5c).
</paragraph>
</subSubSection>
</treatment>
</document>