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<document id="E9953B13A2832F951A09E828738B0358" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A857FFD00CBA12DDFBCB97FB" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus soekhnandanae Perkins 2019, new status" docType="treatment" docVersion="8" lastPageNumber="53" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="53" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:title id="7C78E35914AC173F997DA3001C85218E">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="001FF3A305A455C7DDB80012E9B963FF">Perkins, Philip D.</mods:namePart>
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<treatment id="03A587E7A857FFD00CBA12DDFBCB97FB" ID-DOI="http://doi.org/10.5281/zenodo.5941813" ID-GBIF-Taxon="160798411" ID-Zenodo-Dep="5941813" LSID="urn:lsid:plazi:treatment:03A587E7A857FFD00CBA12DDFBCB97FB" httpUri="http://treatment.plazi.org/id/03A587E7A857FFD00CBA12DDFBCB97FB" lastPageNumber="53" pageId="52" pageNumber="53">
<subSubSection id="C316657AA857FFD00CBA12DDFD0792E1" box="[151,742,153,180]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BB336F1A857FFD00CBA12DDFD0792E1" blockId="52.[151,742,153,215]" box="[151,742,153,180]" pageId="52" pageNumber="53">
<heading id="D0FB819DA857FFD00CBA12DDFD0792E1" bold="true" box="[151,742,153,180]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<emphasis id="B978EAE3A857FFD00CBA12DDFD0792E1" bold="true" box="[151,742,153,180]" pageId="52" pageNumber="53">
<taxonomicName id="4C0C4D72A857FFD00CBA12DDFDB792E1" ID-CoL="3N6C8" authority="Makhan" authorityName="Perkins" authorityYear="2019" baseAuthorityName="Makhan" baseAuthorityYear="1992" box="[151,598,153,180]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="soekhnandanae" status="stat. nov.">
<emphasis id="B978EAE3A857FFD00CBA12DDFE0292E1" bold="true" box="[151,483,153,180]" italics="true" pageId="52" pageNumber="53">Hydrochus soekhnandanae</emphasis>
Makhan
</taxonomicName>
,
<taxonomicNameLabel id="A24B5798A857FFD00E4E12DEFD0792E1" box="[611,742,154,180]" pageId="52" pageNumber="53">new status</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA857FFD00CBA12F9FEEC9282" box="[151,269,189,215]" pageId="52" pageNumber="53" type="description">
<paragraph id="8BB336F1A857FFD00CBA12F9FEEC9282" blockId="52.[151,742,153,215]" box="[151,269,189,215]" pageId="52" pageNumber="53">
<figureCitation id="13372A74A857FFD00CBA12F9FF049282" box="[151,229,189,215]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="52" pageNumber="53">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A857FFD00CDC12F9FEEC9282" box="[241,269,189,215]" captionStart="FIGURE 29" captionStartId="51.[151,250,1957,1982]" captionTargetBox="[238,1380,380,1906]" captionTargetId="figure@51.[196,1391,345,1932]" captionTargetPageId="51" captionText="FIGURE 29. Hydrochus soekhnandanae Makhan, habitus and male genitalia" figureDoi="http://doi.org/10.5281/zenodo.3587241" httpUri="https://zenodo.org/record/3587241/files/figure.png" pageId="52" pageNumber="53">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA857FFD00CBA12B8FCBA9366" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="8BB336F1A857FFD00CBA12B8FD889341" blockId="52.[151,859,252,307]" box="[151,617,252,277]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="AB1C11DFA857FFD00CBA12B8FD889341" box="[151,617,252,277]" pageId="52" pageNumber="53">
<taxonomicName id="4C0C4D72A857FFD00CBA12B8FD849341" ID-CoL="3N6C8" authority="Makhan 1992: 98" authorityName="Makhan" authorityPageNumber="98" authorityYear="1992" box="[151,613,252,277]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B978EAE3A857FFD00CBA12B8FE4B9341" box="[151,426,252,276]" italics="true" pageId="52" pageNumber="53">Hydrochus soekhnandanae</emphasis>
<treatmentCitation id="0AAD10E0A857FFD00D9D12B8FD849341" author="Makhan, D." box="[432,613,252,277]" page="98" pageId="52" pageNumber="53" year="1992">
<bibRefCitation id="EF9D4B00A857FFD00D9D12B8FD849341" author="Makhan, D." box="[432,613,252,277]" pageId="52" pageNumber="53" pagination="95 - 103" refId="ref30207" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992: 98</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB336F1A857FFD00CBA135FFCBA9366" blockId="52.[151,859,252,307]" box="[151,859,282,307]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="AB1C11DFA857FFD00CBA135FFCBA9366" box="[151,859,282,307]" pageId="52" pageNumber="53">
<taxonomicName id="4C0C4D72A857FFD00CBA135FFE3A9366" ID-CoL="6MK94" authority="Orchymont" authorityName="Orchymont. I" authorityYear="1939" box="[151,475,282,307]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pupillus">
<emphasis id="B978EAE3A857FFD00CBA135FFEBF9366" box="[151,350,283,307]" italics="true" pageId="52" pageNumber="53">Hydrochus pupillus</emphasis>
Orchymont
</taxonomicName>
:
<treatmentCitation id="0AAD10E0A857FFD00DC9135EFD7C9366" author="Oliva, A." box="[484,669,282,307]" page="331" pageId="52" pageNumber="53" year="1996">
<bibRefCitation id="EF9D4B00A857FFD00DC9135EFD7C9366" author="Oliva, A." box="[484,669,282,307]" pageId="52" pageNumber="53" pagination="301 - 341" refId="ref30673" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996: 331)</bibRefCitation>
</treatmentCitation>
; misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C316657AA857FFD00CBA131FFB8C907F" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8BB336F1A857FFD00CBA131FFA669323" blockId="52.[151,1437,347,1454]" box="[151,1415,347,374]" pageId="52" pageNumber="53">
<emphasis id="B978EAE3A857FFD00CBA131FFE4B9323" bold="true" box="[151,426,347,374]" pageId="52" pageNumber="53">Material examined (2):</emphasis>
<materialsCitation id="3B643CACA857FFD00D9C1318FC269323" ID-GBIF-Occurrence="2521263700" box="[433,967,347,374]" collectionCode="SEMC" country="Venezuela" location="Sucre" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Sucre">
<emphasis id="B978EAE3A857FFD00D9C1318FD669323" bold="true" box="[433,647,347,374]" pageId="52" pageNumber="53">
<collectingCountry id="F31B7661A857FFD00D9C1318FDCF9323" box="[433,558,348,374]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
:
<collectingRegion id="49C8F813A857FFD00E17131FFD639323" box="[570,642,347,374]" country="Venezuela" name="Sucre" pageId="52" pageNumber="53">Sucre</collectingRegion>
:
</emphasis>
VZ10-0129-04A (1
<collectionCode id="ED1DAE34A857FFD00F431318FC5E9323" box="[878,959,348,374]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="52" pageNumber="53" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA857FFD00FFF1318FA629323" ID-GBIF-Occurrence="2521263696" box="[978,1411,348,374]" collectionCode="MIZA" country="Venezuela" location="Monagas" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Monagas">
<emphasis id="B978EAE3A857FFD00FFF1318FBA99323" bold="true" box="[978,1096,348,374]" pageId="52" pageNumber="53">
<collectingRegion id="49C8F813A857FFD00FFF1318FBA29323" box="[978,1091,348,374]" country="Venezuela" name="Monagas" pageId="52" pageNumber="53">Monagas</collectingRegion>
:
</emphasis>
VZ10-0202-03A (1
<collectionCode id="ED1DAE34A857FFD009021318FA9D9323" box="[1327,1404,348,374]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="52" pageNumber="53" type="Museum">MIZA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A857FFD00CEA133BFB8C907F" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">
<emphasis id="B978EAE3A857FFD00CEA133BFE3793CF" bold="true" box="[199,470,383,410]" pageId="52" pageNumber="53">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of small size (ca.
<quantity id="4CF49B14A857FFD0094613C4FF2693EA" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="52" pageNumber="53" unit="mm" value="1.93">1.93 mm</quantity>
), the bright silver dorsum, the elytra with odd numbered interstriae raised, the large and very closely spaced strial punctures, the pronotum with moderately shallow depressions much more densely punctate than reliefs, the pronotal shape, the lack of black spots on the elytra, and the male genitalia (
<figureCitation id="13372A74A857FFD00FC913A8FBD69053" box="[996,1079,492,518]" captionStart="FIGURE 29" captionStartId="51.[151,250,1957,1982]" captionTargetBox="[238,1380,380,1906]" captionTargetId="figure@51.[196,1391,345,1932]" captionTargetPageId="51" captionText="FIGURE 29. Hydrochus soekhnandanae Makhan, habitus and male genitalia" figureDoi="http://doi.org/10.5281/zenodo.3587241" httpUri="https://zenodo.org/record/3587241/files/figure.png" pageId="52" pageNumber="53">Fig. 29</figureCitation>
; described below). Reliable differentiation from other similarly small sized species should include dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA857FFD00CEA1077FEBC96AF" pageId="52" pageNumber="53" type="description">
<paragraph id="8BB336F1A857FFD00CEA1077FE0C9157" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">
<emphasis id="B978EAE3A857FFD00CEA1077FEB0901B" bold="true" box="[199,337,563,590]" pageId="52" pageNumber="53">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A857FFD00E4B1070FD5C901B" box="[614,701,564,590]" captionStart="FIGURE 29" captionStartId="51.[151,250,1957,1982]" captionTargetBox="[238,1380,380,1906]" captionTargetId="figure@51.[196,1391,345,1932]" captionTargetPageId="51" captionText="FIGURE 29. Hydrochus soekhnandanae Makhan, habitus and male genitalia" figureDoi="http://doi.org/10.5281/zenodo.3587241" httpUri="https://zenodo.org/record/3587241/files/figure.png" pageId="52" pageNumber="53">Fig. 29</figureCitation>
). Size of imaged specimen (length/width, mm): body (length to elytral apices) 1.93/0.75; head width 0.50; pronotum l/w 0.45/0.50; PA 0.48; PB 0.39; elytra 1.18/0.75. Body size small (ca.
<quantity id="4CF49B14A857FFD00D261038FE9590C2" box="[267,372,636,663]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="52" pageNumber="53" unit="mm" value="1.93">1.93 mm</quantity>
). Dorsum bright silver metallic, with slight bluish iridescence, elytra without black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2
<date id="FFB21031A857FFD00F9310E4FC3B90EF" box="[958,986,672,698]" pageId="52" pageNumber="53">4x</date>
that of pronotum. Elytra interstriae ca. 0.5
<date id="FFB21031A857FFD00CE11087FF05908B" box="[204,228,707,734]" pageId="52" pageNumber="53">1x</date>
strial puncture diameter. Odd numbered interstriae raised. Interstria 9
<superScript id="7C799BB9A857FFD008241087FBF79084" attach="left" box="[1033,1046,707,721]" fontSize="6" pageId="52" pageNumber="53">th</superScript>
overhanging 10
<superScript id="7C799BB9A857FFD008FC1087FB3F9084" attach="left" box="[1233,1246,707,721]" fontSize="6" pageId="52" pageNumber="53">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C799BB9A857FFD00D4910A3FE9090A0" attach="left" box="[356,369,743,757]" fontSize="6" pageId="52" pageNumber="53">th</superScript>
not raised.
</paragraph>
<paragraph id="8BB336F1A857FFD00CEA1149FDB3913B" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">Pronotum slightly wider than long (as ca. 45/50), widest slightly behind anterior margin, markedly narrowed at base, sides arcuate or very slightly sinuate, smooth; anterior margin bisinuate; depressions moderately shallow, punctation much denser than on reliefs.</paragraph>
<paragraph id="8BB336F1A857FFD00CEA113CFCAD91E3" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">Elytra with spaces between strial punctures narrow ridges; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with very slight angulation.</paragraph>
<paragraph id="8BB336F1A857FFD00CEA1184FF3391AA" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">Ventral characters: Mentum densely, coarsely punctate, with shallow median fovea. Submentum without foveae.</paragraph>
<paragraph id="8BB336F1A857FFD00CEA164CFAE496DB" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">
Male genitalia (
<figureCitation id="13372A74A857FFD00D57164CFE2C9677" box="[378,461,1032,1058]" captionStart="FIGURE 29" captionStartId="51.[151,250,1957,1982]" captionTargetBox="[238,1380,380,1906]" captionTargetId="figure@51.[196,1391,345,1932]" captionTargetPageId="51" captionText="FIGURE 29. Hydrochus soekhnandanae Makhan, habitus and male genitalia" figureDoi="http://doi.org/10.5281/zenodo.3587241" httpUri="https://zenodo.org/record/3587241/files/figure.png" pageId="52" pageNumber="53">Fig. 29</figureCitation>
) general characters: genitalia very small; parameres extending well beyond distal end of aedeagus, tips in dorsal/ventral views wide in proximal part, narrowing apically, with medial margins diverging distally, in lateral view tips arcuate toward ventrad; basal piece long, longer than parameres, slightly constricted just before base, in lateral view arcuate, orifice with rim weakly sclerotized and characteristically shaped.
</paragraph>
<paragraph id="8BB336F1A857FFD00CEA16DCFB1F96E7" blockId="52.[151,1437,347,1454]" box="[199,1278,1176,1202]" pageId="52" pageNumber="53">Dorsal surface: adtl lacking; pdmm slightly overlapping lateral margins of aedeagus; adbl small.</paragraph>
<paragraph id="8BB336F1A857FFD00CBA16F8FEBC96AF" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">Ventral surface: avtl indistinct; pvmm overlapping lateral margins of aedeagus, medial margins more widely separated than pdmm.</paragraph>
</subSubSection>
<subSubSection id="C316657AA857FFD00CEA1740FBCB97FB" pageId="52" pageNumber="53" type="discussion">
<paragraph id="8BB336F1A857FFD00CEA1740FD9997DF" blockId="52.[151,1437,347,1454]" pageId="52" pageNumber="53">
<emphasis id="B978EAE3A857FFD00CEA1740FEDA974B" bold="true" box="[199,315,1284,1310]" pageId="52" pageNumber="53">Remarks.</emphasis>
<bibRefCitation id="EF9D4B00A857FFD00D6B1740FE38974A" author="Oliva, A." box="[326,473,1284,1311]" pageId="52" pageNumber="53" pagination="301 - 341" refId="ref30673" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
synonymized
<taxonomicName id="4C0C4D72A857FFD00EA91740FBFC974B" authority="Makhan" authorityName="Makhan" authorityYear="1992" box="[644,1053,1284,1310]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B978EAE3A857FFD00EA91740FC57974B" box="[644,950,1284,1310]" italics="true" pageId="52" pageNumber="53">Hydrochus soekhnandanae</emphasis>
Makhan
</taxonomicName>
with
<taxonomicName id="4C0C4D72A857FFD008491740FF069716" authority="Orchymont. I" authorityName="Orchymont. I" authorityYear="1939" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pupillus">
<emphasis id="B978EAE3A857FFD008491740FAA3974B" box="[1124,1346,1284,1310]" italics="true" pageId="52" pageNumber="53">Hydrochus pupillus</emphasis>
Orchymont. I
</taxonomicName>
have examined a dissected male
<typeStatus id="54B78853A857FFD00E7B176DFD579716" box="[598,694,1321,1347]" pageId="52" pageNumber="53" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C0C4D72A857FFD00EF4176DFC499717" authorityName="Makhan" authorityYear="1992" box="[729,936,1320,1346]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B978EAE3A857FFD00EF4176DFC499717" box="[729,936,1320,1346]" italics="true" pageId="52" pageNumber="53">H. soekhnandanae</emphasis>
</taxonomicName>
, and a dissected male
<typeStatus id="54B78853A857FFD008B2176DFB1E9716" box="[1183,1279,1321,1347]" pageId="52" pageNumber="53" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C0C4D72A857FFD0090F176CFE9B9733" authority="Orchymont" authorityName="Orchymont. I" authorityYear="1939" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pupillus">
<emphasis id="B978EAE3A857FFD0090F176CFF109733" italics="true" pageId="52" pageNumber="53">Hydrochus pupillus</emphasis>
Orchymont
</taxonomicName>
, and conclude that they are different species.
<bibRefCitation id="EF9D4B00A857FFD00F581708FBC39733" author="Makhan, D." box="[885,1058,1356,1382]" pageId="52" pageNumber="53" pagination="95 - 103" refId="ref30207" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan (1992)</bibRefCitation>
does not state where the
<typeStatus id="54B78853A857FFD009171708FA7D9733" box="[1338,1436,1356,1382]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0C4D72A857FFD00C9B1734FE0597DF" authorityName="Makhan" authorityYear="1992" box="[182,484,1392,1418]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B978EAE3A857FFD00C9B1734FE0597DF" box="[182,484,1392,1418]" italics="true" pageId="52" pageNumber="53">Hydrochus soekhnandanae</emphasis>
</taxonomicName>
is deposited.
</paragraph>
<paragraph id="8BB336F1A857FFD00CEA17D0FBCB97FB" blockId="52.[151,1437,347,1454]" box="[199,1066,1428,1454]" pageId="52" pageNumber="53">
The specimen used for the images (
<figureCitation id="13372A74A857FFD00E7917D0FD4797FB" box="[596,678,1428,1454]" captionStart="FIGURE 29" captionStartId="51.[151,250,1957,1982]" captionTargetBox="[238,1380,380,1906]" captionTargetId="figure@51.[196,1391,345,1932]" captionTargetPageId="51" captionText="FIGURE 29. Hydrochus soekhnandanae Makhan, habitus and male genitalia" figureDoi="http://doi.org/10.5281/zenodo.3587241" httpUri="https://zenodo.org/record/3587241/files/figure.png" pageId="52" pageNumber="53">Fig. 29</figureCitation>
) is from locality VZ10-0129-04A.
</paragraph>
</subSubSection>
</treatment>
</document>