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<document id="08A90D526ADBD45E25E6C8D9B5195F51" ID-CLB-Dataset="53693" ID-DOI="10.11646/phytotaxa.599.3.7" ID-GBIF-Dataset="c63d7d3e-8150-4e7e-bae2-bf85671d8fc4" ID-ISSN="1179-3163" ID-Zenodo-Dep="8012611" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1686118845818" checkinUser="plazi" docAuthor="Cornejo, Xavier, Homeier, Jürgen &amp; Ulloa, Carmen Ulloa" docDate="2023" docId="C36D87E1FF97245818A8FBE30E7E991A" docLanguage="en" docName="phytotaxa.599.3.7.pdf" docOrigin="Phytotaxa 599 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.599.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Heisteria austroecuadorica Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="204" masterDocId="3F54FF99FF96245B1820FFD70B379F42" masterDocTitle="Heisteria austroecuadorica (Erythropalaceae): A new species from southeastern Ecuador" masterLastPageNumber="206" masterPageNumber="201" pageNumber="202" updateTime="1699521435169" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="30C4B50359291FCDE5A3226BECE41CCB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="928B65FE21BFEB38267FB41ED1CB7C7A">
<mods:title id="761D362E6D9E7C0F2900BB6A789FDD58">Heisteria austroecuadorica (Erythropalaceae): A new species from southeastern Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="35A91A32F28B58CB241EEB88643E371C" type="personal">
<mods:role id="795264F679DAC0B1FDBDFB1089CD818C">
<mods:roleTerm id="4EF8922E6F08F8D7C0B01055C9725E96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB02748D222ABD3E95FA51C7F64176F0">Cornejo, Xavier</mods:namePart>
<mods:nameIdentifier id="F140E78729E76A82AE7C9DCC8436BA1C" type="ORCID">0000-0002-4081-4047</mods:nameIdentifier>
<mods:affiliation id="C31985CE3928D7B84C5FA2C21F820A3B">Herbario GUAY, Departamento de Botánica, Facultad de Ciencias Naturales, Universidad de Guayaquil, Av. Raúl Gómez Lince s. n. y Av. Juan Tanca Marengo (campus Mapasingue), Guayaquil, Ecuador &amp; xcornejoguay @ gmail. com; https: // orcid. org / 0000 - 0002 - 4081 - 4047</mods:affiliation>
<mods:nameIdentifier id="F68C2EC76CC0636DE6012D40E499DB2C" type="email">xcornejoguay@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="07C656AF8DA569F82E1687A9ACA6F3B9" type="personal">
<mods:role id="0D50362D0485366EA74B33ACE66FCEF3">
<mods:roleTerm id="F510EF8743D7830A4BB41B5BBFBC12EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="96AFA8B93DA9C8C6BAAB94D7F7D73167">Homeier, Jürgen</mods:namePart>
<mods:nameIdentifier id="560E6B2212C8F17A520938C68FB48E58" type="ORCID">0000-0001-5676-3267</mods:nameIdentifier>
<mods:affiliation id="E6E7528D89147AC791D5991AB0CA7711">Plant Ecology and Ecosystems Research, University of Goettingen, Goettingen, Germany &amp; Faculty of Resource Management, HAWK University of Applied Sciences and Arts, Goettingen, Germany. &amp; jhomeie @ gwdg. de; https: // orcid. org / 0000 - 0001 - 5676 - 3267</mods:affiliation>
<mods:nameIdentifier id="20C636B5FF0ABB952BCA5AD18FCBCC98" type="email">jhomeie@gwdg.de</mods:nameIdentifier>
</mods:name>
<mods:name id="B8E483A982DDC8F22A5E12B4C4A4828E" type="personal">
<mods:role id="25B066996BB06A9DAD238A548D81FB86">
<mods:roleTerm id="E5F1CDF353C0AA84D2A6BFACB6BFDAC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3DBFCE29A8ECDF78A3507346E091AB7">Ulloa, Carmen Ulloa</mods:namePart>
<mods:nameIdentifier id="B28F9B93A662F277321F35541B0416C6" type="ORCID">0000-0002-4081-4047</mods:nameIdentifier>
<mods:affiliation id="F2BA3EC8F4B31B401969997D7447BE17">Missouri Botanical Garden, 4344 Shaw Blvd., Saint Louis, MO 63110, U. S. A. &amp; carmen. ulloa @ mobot. org; https: // orcid. org / 0000 - 0002 - 4081 - 4047</mods:affiliation>
<mods:nameIdentifier id="41755AABA94C380035F8E2526B8C547B" type="email">carmen.ulloa@mobot.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B28B75E4B4339067A2B5B7A7384CFDC0">text</mods:typeOfResource>
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<mods:title id="0CDFBD1B7B1E7CAB4F0C2E13D56AE91D">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="8B26A2370C1BD5A278D53A5933590AB1">
<mods:date id="1AA9886D337437D1BEEFFB807E844329">2023</mods:date>
<mods:detail id="F557F53D2B5ABD0B5EF936F0A105687A" type="pubDate">
<mods:number id="0E3D51DCDC54BDAF30944FC501BDAD46">2023-06-07</mods:number>
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<mods:detail id="929DBE5BCA0A9CCF7A8E5F8213566FA3" type="volume">
<mods:number id="7B05C354AAC37C648BB51B6931BA0F20">599</mods:number>
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<mods:number id="DD7FA8D167D6CD99FE0D66CC7A489799">3</mods:number>
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<mods:url id="8CE9F338B432D645EB9E410F1940ED80">http://dx.doi.org/10.11646/phytotaxa.599.3.7</mods:url>
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<mods:classification id="413DE30FB3706437770C424C2DC62E37">journal article</mods:classification>
<mods:identifier id="30B6F58CB34D6798D8714561D77C514C" type="CLB-Dataset">53693</mods:identifier>
<mods:identifier id="198013110BBC7C55962AA26BAB0D0C16" type="DOI">10.11646/phytotaxa.599.3.7</mods:identifier>
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<treatment id="C36D87E1FF97245818A8FBE30E7E991A" ID-DOI="http://doi.org/10.5281/zenodo.8012613" ID-GBIF-Taxon="209670456" ID-Zenodo-Dep="8012613" LSID="urn:lsid:plazi:treatment:C36D87E1FF97245818A8FBE30E7E991A" httpUri="http://treatment.plazi.org/id/C36D87E1FF97245818A8FBE30E7E991A" lastPageId="3" lastPageNumber="204" pageId="1" pageNumber="202">
<subSubSection id="03DE657CFF97245A18A8FBE308E39B0C" box="[136,980,1076,1103]" pageId="1" pageNumber="202" type="nomenclature">
<paragraph id="4B7B36F7FF97245A18A8FBE308E39B0C" blockId="1.[136,980,1076,1103]" box="[136,980,1076,1103]" pageId="1" pageNumber="202">
<heading id="1033819BFF97245A18A8FBE308E39B0C" box="[136,980,1076,1103]" fontSize="11" level="2" pageId="1" pageNumber="202" reason="3">
<taxonomicName id="8CC44D74FF97245A18A8FBE308229B0C" ID-CoL="BRHPB" authority="Cornejo, Homeier &amp; C. Ulloa" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[136,789,1076,1103]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="species" species="austroecuadorica" status="sp. nov.">
<emphasis id="79B0EAE5FF97245A18A8FBE30AF59B0D" bold="true" box="[136,450,1076,1103]" italics="true" pageId="1" pageNumber="202">Heisteria austroecuadorica</emphasis>
Cornejo, Homeier &amp; C. Ulloa
</taxonomicName>
<taxonomicNameLabel id="6283579EFF97245A1B3CFBE208449B0D" box="[796,883,1077,1103]" pageId="1" pageNumber="202" rank="species">
<emphasis id="79B0EAE5FF97245A1B3CFBE2085C9B0C" box="[796,875,1077,1102]" italics="true" pageId="1" pageNumber="202">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="D3FF2A72FF97245A1BA2FBE308F49B0C" box="[898,963,1076,1102]" captionStart="FIGURE 1" captionStartId="2.[136,245,1952,1978]" captionTargetBox="[183,1404,187,1929]" captionTargetId="figure-17@2.[183,1404,187,1929]" captionTargetPageId="2" captionText="FIGURE 1. Heisteria austroecuadorica Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence with flower buds. B, Terminal branchlet and immature fruits. C, Fruits surrounded at base by an accrescent persistent reddish-orange calyx. D, Pedicel and abaxial view of fruiting calyx. E, Abaxial view of a leafy branch and immature fruits. Photos by Jürgen Homeier, from the type population." figureDoi="http://doi.org/10.5281/zenodo.8012616" httpUri="https://zenodo.org/record/8012616/files/figure.png" pageId="1" pageNumber="202">Fig. 1</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF97245A18A8FB8C0BC89BD7" pageId="1" pageNumber="202" type="diagnosis">
<paragraph id="4B7B36F7FF97245A18A8FB8C0BC89BD7" blockId="1.[136,1452,1115,1246]" pageId="1" pageNumber="202">
<emphasis id="79B0EAE5FF97245A18A8FB8C0BC49B33" bold="true" box="[136,243,1115,1137]" pageId="1" pageNumber="202">Diagnosis:</emphasis>
—
<emphasis id="79B0EAE5FF97245A192BFB8B0BC89BD7" italics="true" pageId="1" pageNumber="202">
Species nova affinis
<taxonomicName id="8CC44D74FF97245A19F7FB8B09A29B33" authorityName="Sleumer" authorityYear="1984" box="[471,661,1116,1137]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="species" species="asplundii">Heisteria asplundii</taxonomicName>
, a qua folia angusta, stamina decandra, calycem fructuum basin non occultans differt.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF97245A18A8FB730F129B9C" pageId="1" pageNumber="202" type="materials_examined">
<paragraph id="4B7B36F7FF97245A18A8FB730F129B9C" blockId="1.[136,1452,1115,1246]" pageId="1" pageNumber="202">
<emphasis id="79B0EAE5FF97245A18A8FB730BF59BFB" bold="true" box="[136,194,1188,1209]" pageId="1" pageNumber="202">Type:</emphasis>
—
<materialsCitation id="FBAC3CAAFF97245A18FAFB73081E9B9F" ID-GBIF-Occurrence="4127574302" collectingDate="2005-09-06" collectorName="J. Homeier &amp; S. Hanel" country="Ecuador" elevation="1980" latitude="-3.9666667" location="Reserva San Francisco" longLatPrecision="1307" longitude="-79.066666" pageId="1" pageNumber="202" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="holotype">
<collectingCountry id="33D37667FF97245A18FAFB730A799BFB" box="[218,334,1188,1209]" name="Ecuador" pageId="1" pageNumber="202">ECUADOR</collectingCountry>
.
<collectingRegion id="8900F815FF97245A197DFB73092F9BF8" box="[349,536,1188,1210]" country="Ecuador" name="Zamora-Chinchipe" pageId="1" pageNumber="202">Zamora Chinchipe</collectingRegion>
: area of the
<location id="4E1B602CFF97245A1AB7FB73084D9BF8" LSID="urn:lsid:plazi:treatment:C36D87E1FF97245818A8FBE30E7E991A:4E1B602CFF97245A1AB7FB73084D9BF8" box="[663,890,1188,1210]" country="Ecuador" latitude="-3.9666667" longLatPrecision="1307" longitude="-79.066666" name="Reserva San Francisco" pageId="1" pageNumber="202" stateProvince="Zamora-Chinchipe">Reserva San Francisco</location>
,
<geoCoordinate id="2EF05030FF97245A1BA6FB7308FE9BF8" box="[902,969,1188,1210]" degrees="3" direction="south" minutes="58" orientation="latitude" pageId="1" pageNumber="202" precision="925" value="-3.9666667">3°58’S</geoCoordinate>
<geoCoordinate id="2EF05030FF97245A1BF2FB730F1D9BFB" box="[978,1066,1188,1209]" degrees="79" direction="west" minutes="04" orientation="longitude" pageId="1" pageNumber="202" precision="925" value="-79.066666">79°04’W</geoCoordinate>
,
<quantity id="8C3C9B12FF97245A1C17FB730FB29BFB" box="[1079,1157,1188,1209]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="1" pageNumber="202" unit="m" value="1980.0">
<elevation id="C0E9D1C4FF97245A1C17FB730FB29BFB" box="[1079,1157,1188,1209]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="1" pageNumber="202" unit="m" value="1980.0">1980 m</elevation>
</quantity>
,
<date id="3F7A1037FF97245A1CB1FB730E729BF8" box="[1169,1349,1188,1210]" pageId="1" pageNumber="202" value="2005-09-06">
<collectingDate id="2F3EE9DFFF97245A1CB1FB730E729BF8" box="[1169,1349,1188,1210]" pageId="1" pageNumber="202" value="2005-09-06">6 September 2005</collectingDate>
</date>
(fl, fr),
<emphasis id="79B0EAE5FF97245A1DBBFB730A8E9B9F" italics="true" pageId="1" pageNumber="202">
<collectorName id="E6315321FF97245A1DBBFB730A269B9F" pageId="1" pageNumber="202">J. Homeier</collectorName>
&amp;
<collectorName id="E6315321FF97245A1910FB1F0AB49B9F" box="[304,387,1224,1245]" pageId="1" pageNumber="202">S. Hänel</collectorName>
1609
</emphasis>
(
<typeStatus id="947F8855FF97245A19E5FB1F09299B9C" box="[453,542,1224,1246]" pageId="1" pageNumber="202" type="holotype">holotype</typeStatus>
: GUAY!; isotypes: GOET!,
</materialsCitation>
<collectingRegion id="8900F815FF97245A1B0EFB1F085E9B9F" box="[814,873,1224,1245]" country="Ecuador" name="Loja" pageId="1" pageNumber="202">LOJA</collectingRegion>
!, QCNE-195572!).
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF97245A18A8FADB0AD3985D" pageId="1" pageNumber="202" type="description">
<paragraph id="4B7B36F7FF97245A18A8FADB0AD3985D" blockId="1.[136,1452,1292,2075]" pageId="1" pageNumber="202">
<emphasis id="79B0EAE5FF97245A18A8FADB0A2B9A64" bold="true" box="[136,284,1292,1318]" pageId="1" pageNumber="202">Description:</emphasis>
—
<emphasis id="79B0EAE5FF97245A1918FADA0A5E9A64" box="[312,361,1293,1318]" italics="true" pageId="1" pageNumber="202">Tree</emphasis>
, to
<quantity id="8C3C9B12FF97245A19B2FADB0AFD9A65" box="[402,458,1292,1319]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="202" unit="m" value="20.0">20 m</quantity>
tall and
<quantity id="8C3C9B12FF97245A1A08FADB095B9A65" box="[552,620,1292,1319]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="202" unit="cm" value="30.0">30 cm</quantity>
DBH; milky white latex present in stems and branches; branches ca.
<quantity id="8C3C9B12FF97245A1D4CFADB0E9B9A65" box="[1388,1452,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="202" unit="mm" value="2.0">2 mm</quantity>
thick, terete, striate, glabrous.
<emphasis id="79B0EAE5FF97245A19FAFAE6091F9A08" box="[474,552,1329,1354]" italics="true" pageId="1" pageNumber="202">Leaves</emphasis>
alternate;
<emphasis id="79B0EAE5FF97245A1ABDFAE709DE9A08" box="[669,745,1328,1354]" italics="true" pageId="1" pageNumber="202">petiole</emphasis>
<quantity id="8C3C9B12FF97245A1AD0FAE7086C9A09" box="[752,859,1328,1355]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="1" pageNumber="202" unit="mm" value="11.5" valueMax="16.0" valueMin="7.0">7–16 mm</quantity>
long, rugose, light- to dark-brown, channeled above, glabrous; lamina chartaceous, (4.5–)7–19 ×
<quantity id="8C3C9B12FF97245A1AA4FA8309EA9A2D" box="[644,733,1364,1391]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="1" pageNumber="202" unit="cm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 cm</quantity>
, elliptic-lanceolate to narrowly elliptic or narrowly lanceolate, apex acuminate to long-caudate, drip-tip
<quantity id="8C3C9B12FF97245A1A70FAAF098D9AD1" box="[592,698,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="1" pageNumber="202" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">7–12 mm</quantity>
long, base cuneate, margins revolute, dull, smooth, and glabrous on both sides, venation brochidodromous, midrib slightly sulcate above, prominent below, secondary nerves 6 to 9 pairs, prominulous, tertiary venation inconspicuous.
<emphasis id="79B0EAE5FF97245A1AB3FA17081E9A98" box="[659,809,1472,1498]" italics="true" pageId="1" pageNumber="202">Inflorescence</emphasis>
ramiflorous, a short axillary fascicle, 3 to 15 flowers per fascicle; bracts cucullate-deltoid, 0.5–1 ×
<quantity id="8C3C9B12FF97245A1A41FA3309E29ABD" box="[609,725,1508,1535]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="202" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
, the apex obtuse to broadly obtuse; pedicel
<quantity id="8C3C9B12FF97245A1CE5FA330E159ABD" box="[1221,1314,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="202" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long.
<emphasis id="79B0EAE5FF97245A1D4AFA330E9C9ABC" box="[1386,1451,1508,1534]" italics="true" pageId="1" pageNumber="202">Calyx</emphasis>
patelliform or cup-shaped, 5-lobed, lobes ca.
<quantity id="8C3C9B12FF97245A1A5FF9DF09E39961" box="[639,724,1544,1571]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="202" unit="mm" value="0.5">0.5 mm</quantity>
long, triangular to broadly triangular, acute at apex, green.
<emphasis id="79B0EAE5FF97245A1D45F9DF0E9C9960" box="[1381,1451,1544,1570]" italics="true" pageId="1" pageNumber="202">Petals</emphasis>
5, ca. 3 ×
<quantity id="8C3C9B12FF97245A18DDF9FB0A769905" box="[253,321,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="202" unit="mm" value="1.0">1 mm</quantity>
, free, densely papillose or shortly pilose on the adaxial side, greenish-white.
<emphasis id="79B0EAE5FF97245A1C92F9FB0E389904" box="[1202,1295,1580,1606]" italics="true" pageId="1" pageNumber="202">Stamens</emphasis>
10, filaments ligulate, ca. 2 ×
<quantity id="8C3C9B12FF97245A191CF9870AA39929" box="[316,404,1616,1643]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="202" unit="mm" value="0.3">0.3 mm</quantity>
, the 5 inner ones ca.
<quantity id="8C3C9B12FF97245A1A5FF98709E09929" box="[639,727,1616,1643]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="202" unit="mm" value="1.8">1.8 mm</quantity>
, all free, anthers ca. 0.5 ×
<quantity id="8C3C9B12FF97245A1BDEF9870F609929" box="[1022,1111,1616,1643]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="202" unit="mm" value="0.8">0.8 mm</quantity>
.
<emphasis id="79B0EAE5FF97245A1C41F9860F8D9928" box="[1121,1210,1617,1642]" italics="true" pageId="1" pageNumber="202">Nectary</emphasis>
absent; ovary oblatesubglobose (
<emphasis id="79B0EAE5FF97245A1936F9A20A5599CC" box="[278,354,1653,1678]" italics="true" pageId="1" pageNumber="202">in vivo</emphasis>
), pyriform (
<emphasis id="79B0EAE5FF97245A19CDF9A2097399CC" box="[493,580,1653,1678]" italics="true" pageId="1" pageNumber="202">in sicco</emphasis>
); style ca.
<quantity id="8C3C9B12FF97245A1A9EF9A3082199CD" box="[702,790,1652,1679]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="202" unit="mm" value="0.5">0.5 mm</quantity>
, stigma slightly 3-lobed.
<emphasis id="79B0EAE5FF97245A1C12F9A20FD099CC" box="[1074,1255,1652,1678]" italics="true" pageId="1" pageNumber="202">Fruiting pedicel</emphasis>
<quantity id="8C3C9B12FF97245A1CCEF9A30E5C99CD" box="[1262,1387,1652,1679]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="1" pageNumber="202" unit="cm" value="1.9" valueMax="2.3" valueMin="1.5">1.5–2.3 cm</quantity>
long, slightly swollen,
<quantity id="8C3C9B12FF97245A196AF94F0A9F99F1" box="[330,424,1688,1715]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="202" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
distally.
<emphasis id="79B0EAE5FF97245A1A30F94E098799F0" box="[528,688,1688,1714]" italics="true" pageId="1" pageNumber="202">Fruiting calyx</emphasis>
slightly 5-lobed,
<quantity id="8C3C9B12FF97245A1B59F94F08C099F1" box="[889,1015,1688,1715]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="1" pageNumber="202" unit="cm" value="2.5" valueMax="2.8" valueMin="2.2">2.2–2.8 cm</quantity>
in diam., reflexed, stiffly chartaceous, orange to red, bright and shiny above, dull below.
<emphasis id="79B0EAE5FF97245A1AF5F96A082A9994" box="[725,797,1725,1750]" italics="true" pageId="1" pageNumber="202">Drupe</emphasis>
barrel-shape, olive green with maroon at apex (
<emphasis id="79B0EAE5FF97245A1D6DF96A0EAC9994" box="[1357,1435,1725,1750]" italics="true" pageId="1" pageNumber="202">in vivo</emphasis>
), 1.2–2 ×
<quantity id="8C3C9B12FF97245A18C4F9370A7999B9" box="[228,334,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="202" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
, rugulose, faintly ribbed longitudinally and tuberculate (from endocarp,
<emphasis id="79B0EAE5FF97245A1C55F9360FFC99B8" box="[1141,1227,1761,1786]" italics="true" pageId="1" pageNumber="202">in sicco</emphasis>
), with a clear, deep, circular depression at the apex.
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF97245A189DF8FF0A069825" pageId="1" pageNumber="202" type="etymology">
<paragraph id="4B7B36F7FF97245A189DF8FF0A069825" blockId="1.[136,1452,1292,2075]" pageId="1" pageNumber="202">
<emphasis id="79B0EAE5FF97245A189DF8FF0A7F9800" bold="true" box="[189,328,1832,1858]" pageId="1" pageNumber="202">Etymology:</emphasis>
—The epithet refers to the geographic pattern of distribution at southeastern
<collectingCountry id="33D37667FF97245A1CF1F8FF0E199800" box="[1233,1326,1832,1858]" name="Ecuador" pageId="1" pageNumber="202">Ecuador</collectingCountry>
where the species occurs.
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF972458189DF8A70E099DF2" lastPageId="3" lastPageNumber="204" pageId="1" pageNumber="202" type="biology_ecology">
<paragraph id="4B7B36F7FF972458189DF8A70FA79F9E" blockId="1.[136,1452,1292,2075]" lastBlockId="3.[136,1452,158,1625]" lastPageId="3" lastPageNumber="204" pageId="1" pageNumber="202">
<emphasis id="79B0EAE5FF97245A189DF8A70AC198C8" bold="true" box="[189,502,1904,1930]" pageId="1" pageNumber="202">Habitat and distribution:</emphasis>
—
<taxonomicName id="8CC44D74FF97245A1A32F8A6087C98C8" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[530,843,1904,1930]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF97245A1A32F8A6087C98C8" box="[530,843,1904,1930]" italics="true" pageId="1" pageNumber="202">Heisteria austroecuadorica</emphasis>
</taxonomicName>
is known from the San Francisco reserve, a forest remnant close to the settlement of El
<collectingCountry id="33D37667FF97245A1A64F84309A498EC" box="[580,659,1940,1966]" name="Lebanon" pageId="1" pageNumber="202">Líbano</collectingCountry>
north of the Podocarpus National Park, and the Jamboe river basin in southeastern
<collectingCountry id="33D37667FF97245A191DF86F0AAB9890" box="[317,412,1976,2002]" name="Ecuador" pageId="1" pageNumber="202">Ecuador</collectingCountry>
, between
<quantity id="8C3C9B12FF97245A1A31F86F09DC9890" box="[529,747,1976,2003]" metricMagnitude="3" metricUnit="m" metricValue="1.525" metricValueMax="2.05" metricValueMin="1.0" pageId="1" pageNumber="202" unit="m" value="1525.0" valueMax="2050.0" valueMin="1000.0">1000 to 2050 masl</quantity>
; all three localities are on the Loja-Zamora road. The area is characterized by a rugged terrain and small-scale structural and floristic vegetation changes, which result mainly from the decreasing soil fertility along ravine-slope-ridge gradients (Werner &amp; Homeier 2015). The newly described species is restricted to the ravines and lower slope positions with more fertile soils within the species-rich evergreen lower montane forests below
<quantity id="8C3C9B12FF95245819ADFF1509339F9E" box="[397,516,194,221]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="3" pageNumber="204" unit="m" value="2050.0">2050 masl</quantity>
, that reach up to
<quantity id="8C3C9B12FF9524581AE3FF1509CB9F9F" box="[707,764,194,221]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="204" unit="m" value="30.0">30 m</quantity>
height (
<bibRefCitation id="2F554B06FF9524581B77FF150F0B9F9E" author="Homeier, J. &amp; Werner, F. A. &amp; Gradstein, S. R. &amp; Breckle, S. - W. &amp; Richter, M." box="[855,1084,194,220]" pageId="3" pageNumber="204" pagination="87 - 100" refId="ref3029" refString="Homeier, J., Werner, F. A., Gradstein, S. R., Breckle, S. - W. &amp; Richter, M. (2008) Potential vegetation and floristic composition of Andean forests in South Ecuador, with a focus on the RBSF. In: Beck, E., Bendix, J., Kottke, I., Makeschin, F. &amp; Mosandl, R. (eds.) Gradients in a Tropical Mountain Ecosystem of Ecuador. Ecological Studies 198: 87 - 100. https: // doi. org / 10.1007 / 978 - 3 - 540 - 73526 - 7 _ 10" type="journal article" year="2008">
Homeier
<emphasis id="79B0EAE5FF9524581BE2FF1408CD9F9E" box="[962,1018,194,220]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2008
</bibRefCitation>
, 2010).
</paragraph>
<caption id="1FBB667FFF94245918A8F8770A929764" ID-DOI="http://doi.org/10.5281/zenodo.8012616" ID-Zenodo-Dep="8012616" httpUri="https://zenodo.org/record/8012616/files/figure.png" pageId="2" pageNumber="203" startId="2.[136,245,1952,1978]" targetBox="[183,1404,187,1929]" targetPageId="2" targetType="figure">
<paragraph id="4B7B36F7FF94245918A8F8770A929764" blockId="2.[136,1452,1952,2087]" pageId="2" pageNumber="203">
<emphasis id="79B0EAE5FF94245918A8F8770A3E98F8" bold="true" box="[136,265,1952,1978]" pageId="2" pageNumber="203">FIGURE 1</emphasis>
.
<taxonomicName id="8CC44D74FF9424591936F8760F6D98F8" authority="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[278,1114,1952,1979]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="2" pageNumber="203" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF9424591936F876097E98F8" box="[278,585,1952,1978]" italics="true" pageId="2" pageNumber="203">Heisteria austroecuadorica</emphasis>
Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence
</taxonomicName>
with flower buds. B, Terminal branchlet and immature fruits. C, Fruits surrounded at base by an accrescent persistent reddish-orange calyx. D, Pedicel and abaxial view of fruiting calyx. E, Abaxial view of a leafy branch and immature fruits. Photos by Jürgen Homeier, from the type population.
</paragraph>
</caption>
<paragraph id="4B7B36F7FF952458189DFF300BE69D62" blockId="3.[136,1452,158,1625]" pageId="3" pageNumber="204">
In the ravines and on the lower slopes of this specific forest
<typeStatus id="947F8855FF9524581B5BFF3008999E43" box="[891,942,231,257]" pageId="3" pageNumber="204">type</typeStatus>
,
<taxonomicName id="8CC44D74FF9524581B9CFF300FC49E42" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[956,1267,230,256]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF9524581B9CFF300FC49E42" box="[956,1267,230,256]" italics="true" pageId="3" pageNumber="204">Heisteria austroecuadorica</emphasis>
</taxonomicName>
is accompanied by several tree species that have been described from this zone in the last years, stressing the biological importance of this area. Some examples are:
<taxonomicName id="8CC44D74FF9524581A2BFEF808DB9E0A" authority="Anderson (2007: 97)" authorityName="Anderson" authorityPageNumber="97" authorityYear="2007" box="[523,1004,302,328]" class="Magnoliopsida" family="Malpighiaceae" genus="Byrsonima" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="homeieri">
<emphasis id="79B0EAE5FF9524581A2BFEF809C59E0A" box="[523,754,302,328]" italics="true" pageId="3" pageNumber="204">Byrsonima homeieri</emphasis>
<bibRefCitation id="2F554B06FF9524581ADAFEF908DB9E0A" author="Anderson, W. R." box="[762,1004,302,328]" pageId="3" pageNumber="204" pagination="95 - 111" refId="ref2681" refString="Anderson, W. R. (2007) Notes on Neotropical Malpighiaceae - IX. Contributions from the University of Michigan Herbarium 25: 95 - 111." type="journal article" year="2007">Anderson (2007: 97)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF9524581BDAFEF80AAF9E2E" authority="Haro-Carrion &amp; Robinson (2008: 17)" authorityName="Haro-Carrion &amp; Robinson" authorityPageNumber="17" authorityYear="2008" class="Magnoliopsida" family="Asteraceae" genus="Critoniopsis" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="zamorensis">
<emphasis id="79B0EAE5FF9524581BDAFEF80E3C9E0A" box="[1018,1291,303,329]" italics="true" pageId="3" pageNumber="204">Critoniopsis zamorensis</emphasis>
<bibRefCitation id="2F554B06FF9524581D35FEF90AAF9E2E" author="Haro-Carrion, X. &amp; Robinson, H." pageId="3" pageNumber="204" pagination="1 - 18" refId="ref2975" refString="Haro-Carrion, X. &amp; Robinson, H. (2008) A review of the genus Critoniopsis in Ecuador (Vernonieae: Asteraceae). Proceedings of the Biological Society of Washington 121 (1): 1 - 18. https: // doi. org / 10.2988 / 07 - 18.1" type="journal article" year="2008">Haro-Carrión &amp; Robinson (2008: 17)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF9524581987FE8508F39E2E" authority="Cornejo (2009: 93)" authorityName="Cornejo" authorityPageNumber="93" authorityYear="2009" box="[423,964,338,364]" class="Magnoliopsida" family="Sabiaceae" genus="Meliosma" kingdom="Plantae" order="Proteales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="longepedicellata">
<emphasis id="79B0EAE5FF9524581987FE8509EC9E2E" box="[423,731,338,364]" italics="true" pageId="3" pageNumber="204">Meliosma longepedicellata</emphasis>
<bibRefCitation id="2F554B06FF9524581AC4FE8508F39E2E" author="Cornejo, X." box="[740,964,338,364]" pageId="3" pageNumber="204" pagination="93 - 96" refId="ref2879" refString="Cornejo, X. (2009) Meliosma longepedicellata (Sabiaceae): a new species from southeastern Ecuador. Harvard Papers in Botany 14: 93 - 96. https: // doi. org / 10.3100 / 025.014.0216" type="journal article" year="2009">Cornejo (2009: 93)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF9524581BF2FE850B899ED3" authority="Arroyo et al. (2013: 507)" authorityName="Arroyo" authorityPageNumber="507" authorityYear="2013" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="zamorana">
<emphasis id="79B0EAE5FF9524581BF2FE850F8E9E2E" box="[978,1209,338,364]" italics="true" pageId="3" pageNumber="204">Magnolia zamorana</emphasis>
<bibRefCitation id="2F554B06FF9524581CE2FE850B899ED3" author="Arroyo, F. &amp; Perez Castaneda, A. J. &amp; Vazquez Garcia, J. A." pageId="3" pageNumber="204" pagination="497 - 508" refId="ref2710" refString="Arroyo, F., Perez Castaneda, A. J. &amp; Vazquez Garcia, J. A. (2013) Six new threatened tree species of Magnolia (Magnoliaceae) from Ecuador and Peru. 2. In: Salcedo Perez, E., Hernandez Alvarez, E., Vazquez Garcia, J. A., Escoto Garcia, T. &amp; Diaz Echavarria, N. (eds.) Recursos Forestales en el Occidente de Mexico. Universidad de Guadalajara, Guadalajara, pp. 497 - 508." type="book chapter" year="2013">
Arroyo
<emphasis id="79B0EAE5FF9524581D3DFE840E6F9E2E" box="[1309,1368,338,364]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
(2013: 507)
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF95245818E9FEA0086E9ED2" authority="Ulloa Ulloa &amp; Homeier (2008: 383)" authorityName="Ulloa Ulloa &amp; Homeier" authorityPageNumber="383" authorityYear="2008" box="[201,857,374,400]" class="Magnoliopsida" family="Melastomataceae" genus="Meriania" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="franciscana">
<emphasis id="79B0EAE5FF95245818E9FEA00A8A9ED2" box="[201,445,374,400]" italics="true" pageId="3" pageNumber="204">Meriania franciscana</emphasis>
<bibRefCitation id="2F554B06FF95245819E4FEA1086E9ED2" author="Ulloa Ulloa, C. &amp; Homeier, J." box="[452,857,374,400]" pageId="3" pageNumber="204" pagination="383 - 387" refId="ref3822" refString="Ulloa Ulloa, C. &amp; Homeier, J. (2008) Meriania franciscana (Melastomataceae), una especie nueva de los Andes sur orientales del Ecuador. Anales del Jardin Botanico de Madrid 65: 383 - 387." type="journal article" year="2008">Ulloa Ulloa &amp; Homeier (2008: 383)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF9524581B45FEA10E909ED2" authority="Berg &amp; Homeier (2010: 197)" authorityName="Berg &amp; Homeier" authorityPageNumber="197" authorityYear="2010" box="[869,1447,374,401]" class="Magnoliopsida" family="Moraceae" genus="Naucleopsis" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="francisci">
<emphasis id="79B0EAE5FF9524581B45FEA10F609ED2" box="[869,1111,374,400]" italics="true" pageId="3" pageNumber="204">Naucleopsis francisci</emphasis>
<bibRefCitation id="2F554B06FF9524581C7EFEA00E909ED2" author="Berg, C. C. &amp; Homeier, J." box="[1118,1447,374,401]" pageId="3" pageNumber="204" pagination="196 - 200" refId="ref2838" refString="Berg, C. C. &amp; Homeier, J. (2010) Three new species of South American Moraceae. Blumea 55: 196 - 200. https: // doi. org / 10.3767 / 000651910 X 527707" type="journal article" year="2010">Berg &amp; Homeier (2010: 197)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF95245818A8FE4C09B19EF6" authority="van der Werff (2020: 210)" authorityName="van der Werff" authorityPageNumber="210" authorityYear="2020" box="[136,646,410,437]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="homeieri">
<emphasis id="79B0EAE5FF95245818A8FE4C0A7F9EF6" box="[136,328,410,437]" italics="true" pageId="3" pageNumber="204">Ocotea homeieri</emphasis>
<bibRefCitation id="2F554B06FF9524581972FE4C09B19EF6" author="van der Werff, H." box="[338,646,410,437]" pageId="3" pageNumber="204" pagination="192 - 230" refId="ref3864" refString="van der Werff, H. (2020) Studies in Andean Ocotea (Lauraceae) V. Species with unisexual flowers and sparsely appressed-pubescent or glabrous leaves without domatia occurring above 1000 m in altitude. Novon 28: 192 - 230. https: // doi. org / 10.3417 / 2020450" type="journal article" year="2020">van der Werff (2020: 210)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8CC44D74FF9524581AE3FE4C0E239EF7" authority="Schneider &amp; Zizka (2012: 265)" authorityName="Schneider &amp; Zizka" authorityPageNumber="265" authorityYear="2012" box="[707,1300,410,437]" class="Magnoliopsida" family="Quiinaceae" genus="Quiina" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="zamorensis">
<emphasis id="79B0EAE5FF9524581AE3FE4C08AE9EF6" box="[707,921,411,437]" italics="true" pageId="3" pageNumber="204">Quiina zamorensis</emphasis>
<bibRefCitation id="2F554B06FF9524581B84FE4D0E239EF7" author="Schneider, J. V. &amp; Zizka, G." box="[932,1300,410,437]" pageId="3" pageNumber="204" pagination="261 - 267" refId="ref3559" refString="Schneider, J. V. &amp; Zizka, G. (2012) Quiina maracaensis J. V. Schneid. &amp; Zizka and Quiina zamorensis J. V. Schneid. &amp; Zizka (Quiinaceae), two new species of Quiina Aubl. from the Neotropics. Candollea 67: 261 - 267. https: // doi. org / 10.15553 / c 2012 v 672 a 7" type="journal article" year="2012">Schneider &amp; Zizka (2012: 265)</bibRefCitation>
</taxonomicName>
. All of these aforementioned species have a narrow pattern of distribution that is restricted to the particularly diverse Amotape-Huancabamba zone (
<bibRefCitation id="2F554B06FF9524581953FE35092D9EBE" author="Weigend, M." box="[371,538,482,508]" pageId="3" pageNumber="204" pagination="38 - 54" refId="ref3918" refString="Weigend, M. (2002) Observations of the Biogeography of the Amotape-Huancabamba Zone in Northern Peru. Botanical Review 68: 38 - 54. https: // doi. org / 10.1663 / 0006 - 8101 (2002) 068 [0038: OOTBOT] 2.0. CO; 2" type="journal article" year="2002">Weigend 2002</bibRefCitation>
,
<bibRefCitation id="2F554B06FF9524581A05FE3509569EBE" author="Weigend, M." box="[549,609,482,508]" pageId="3" pageNumber="204" pagination="127 - 134" refId="ref3971" refString="Weigend, M. (2004) Additional observations on the biogeography of the Amotape Huancabamba zone in Northern Peru: Defining the South-Eastern limits. Revista Peruana de Biologia 11: 127 - 134. https: // doi. org / 10.15381 / rpb. v 11 i 2.2447" type="journal article" year="2004">2004</bibRefCitation>
), south of the Girón-Paute dry valley in southern
<collectingCountry id="33D37667FF9524581CAEFE350FDC9EBE" box="[1166,1259,482,508]" name="Ecuador" pageId="3" pageNumber="204">Ecuador</collectingCountry>
(
<bibRefCitation id="2F554B06FF9524581CDCFE340BF39D62" author="Jorgensen, P. M. &amp; Leon, S." pageId="3" pageNumber="204" pagination="1 - 118" refId="ref3391" refString="Jorgensen, P. M. &amp; Leon, S. (eds.) (1999) Catalogue of the Vascular Plants of Ecuador. Monographs in Systematic Botany from the Missouri Botanical Garden 75: 1 - 118." type="journal article" year="1999">
Jørgensen
<emphasis id="79B0EAE5FF9524581D53FE340E9B9EBE" box="[1395,1452,482,508]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
1999
</bibRefCitation>
).
</paragraph>
<paragraph id="4B7B36F7FF952458189DFDFD0F099D2A" blockId="3.[136,1452,158,1625]" pageId="3" pageNumber="204">
With its relatively high-density wood (
<quantity id="8C3C9B12FF9524581AACFDFD09EE9D06" box="[652,729,554,580]" metricMagnitude="-4" metricUnit="kg" metricValue="7.199999999999999" pageId="3" pageNumber="204" unit="g" value="0.72">0.72 g</quantity>
/cm
<superScript id="BCB19BBFFF9524581B24FDFD083B9D7A" attach="left" box="[772,780,554,568]" fontSize="6" pageId="3" pageNumber="204">3</superScript>
), large fruits and slow diameter growth rates,
<taxonomicName id="8CC44D74FF9524581D63FDFC0A7A9D2A" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF9524581D63FDFC0A7A9D2A" italics="true" pageId="3" pageNumber="204">Heisteria austroecuadorica</emphasis>
</taxonomicName>
combines characteristics typical of a late successional tree species.
</paragraph>
<paragraph id="4B7B36F7FF952458189DFDA50E099DF2" blockId="3.[136,1452,158,1625]" pageId="3" pageNumber="204">
<emphasis id="79B0EAE5FF952458189DFDA50A749DCE" bold="true" box="[189,323,626,652]" pageId="3" pageNumber="204">Phenology:</emphasis>
—Flowering was observed in June, September, October and December; fruiting material was collected in April, September and October. There are no existing observations on flower pollination or fruit dispersal.
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF952458189DFD6D0BE89CEE" pageId="3" pageNumber="204" type="conservation">
<paragraph id="4B7B36F7FF952458189DFD6D0BE89CEE" blockId="3.[136,1452,158,1625]" pageId="3" pageNumber="204">
<emphasis id="79B0EAE5FF952458189DFD6D0A839D96" bold="true" box="[189,436,698,724]" pageId="3" pageNumber="204">Conservation status:</emphasis>
—This species is uncommon in the Reserva San Francisco: Homeier has only found a total of six trees (out of almost 4000) within three recently established one-hectare plots. At a second location, outside the Reserve, in El
<collectingCountry id="33D37667FF952458190BFCD50A4A9C5E" box="[299,381,770,796]" name="Lebanon" pageId="3" pageNumber="204">Líbano</collectingCountry>
, only one tree has been seen. A third locality is represented by one collection only in the Jamboe river basin just further down at slightly lower elevation. Based on the very small area of occurrence (
<quantity id="8C3C9B12FF9524581CC1FCF10E1F9C02" box="[1249,1320,806,832]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="204" unit="km" value="12.0">12 km</quantity>
<superScript id="BCB19BBFFF9524581D08FCF10E079C76" attach="left" box="[1320,1328,806,820]" fontSize="6" pageId="3" pageNumber="204">2</superScript>
), an extent of occurrence of
<quantity id="8C3C9B12FF952458196EFC9D0A9A9C26" box="[334,429,842,868]" metricMagnitude="4" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="204" unit="km" value="10.3">10.3 km</quantity>
<superScript id="BCB19BBFFF952458198DFC9D0A829C1A" attach="left" box="[429,437,842,856]" fontSize="6" pageId="3" pageNumber="204">2</superScript>
, the existence of only three localities, only one of which is protected within the private Reserva San Francisco, we assign a provisional category of Endangered (EN B2a(iv)), following the
<bibRefCitation id="2F554B06FF9524581D33FCB90E9B9CCA" author="IUCN Standards &amp; Petitions Committee" box="[1299,1452,878,904]" pageId="3" pageNumber="204" refId="ref3250" refString="IUCN Standards and Petitions Committee (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 1 July 2022)." type="book" year="2022">IUCN (2022)</bibRefCitation>
criteria.
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF952458189DFC61090E9B8E" pageId="3" pageNumber="204" type="materials_examined">
<paragraph id="4B7B36F7FF952458189DFC61090E9B8E" blockId="3.[136,1452,158,1625]" pageId="3" pageNumber="204">
<materialsCitation id="FBAC3CAAFF952458189DFC610E359CB6" ID-GBIF-Occurrence="4127574304" collectingDate="2005-06" collectorName="O. Cabrera &amp; N. Leon" country="Ecuador" elevation="1925" latitude="-3.9666667" location="Loja-Zamora" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Zamora Chinchipe" typeStatus="paratype">
<emphasis id="79B0EAE5FF952458189DFC610A089C92" bold="true" box="[189,319,950,976]" pageId="3" pageNumber="204">
<typeStatus id="947F8855FF952458189DFC610A0E9C92" box="[189,313,950,976]" pageId="3" pageNumber="204" type="paratype">Paratypes</typeStatus>
:
</emphasis>
—
<collectingCountry id="33D37667FF952458197AFC610A8B9C92" box="[346,444,950,976]" name="Ecuador" pageId="3" pageNumber="204">Ecuador</collectingCountry>
:
<collectingRegion id="8900F815FF95245819EBFC6009929C92" box="[459,677,950,977]" country="Ecuador" name="Zamora-Chinchipe" pageId="3" pageNumber="204">Zamora Chinchipe</collectingRegion>
:
<collectingMunicipality id="AB1FAC8DFF9524581A93FC60088A9C92" box="[691,957,950,977]" pageId="3" pageNumber="204">Reserva San Francisco</collectingMunicipality>
, road
<location id="4E1B602CFF9524581C29FC610F979C92" LSID="urn:lsid:plazi:treatment:C36D87E1FF97245818A8FBE30E7E991A:4E1B602CFF9524581C29FC610F979C92" box="[1033,1184,950,976]" country="Ecuador" latitude="-3.9666667" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" name="Loja-Zamora" pageId="3" pageNumber="204" stateProvince="Zamora Chinchipe">Loja-Zamora</location>
, ca.
<quantity id="8C3C9B12FF9524581CFAFC610E129C92" box="[1242,1317,950,976]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="204" unit="km" value="30.0">
<locationDeviation id="5AC92FD5FF9524581CFAFC610E129C92" box="[1242,1317,950,976]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="204" unit="km" value="30.0">30 km</locationDeviation>
</quantity>
from Loja,
<geoCoordinate id="2EF05030FF95245818A8FC0D0BE19CB7" box="[136,214,986,1013]" degrees="3" direction="south" minutes="58" orientation="latitude" pageId="3" pageNumber="204" precision="925" value="-3.9666667">3°58’S</geoCoordinate>
<geoCoordinate id="2EF05030FF95245818FCFC0D0A759CB6" box="[220,322,986,1012]" degrees="79" direction="west" minutes="04" orientation="longitude" pageId="3" pageNumber="204" precision="925" value="-79.066666">79°04’W</geoCoordinate>
,
<quantity id="8C3C9B12FF952458196DFC0D0ADF9CB7" box="[333,488,986,1013]" metricMagnitude="3" metricUnit="m" metricValue="1.925" metricValueMax="2.05" metricValueMin="1.8" pageId="3" pageNumber="204" unit="m" value="1925.0" valueMax="2050.0" valueMin="1800.0">
<elevation id="C0E9D1C4FF952458196DFC0D0ADF9CB7" box="[333,488,986,1013]" metricMagnitude="3" metricUnit="m" metricValue="1.925" metricValueMax="2.05" metricValueMin="1.8" pageId="3" pageNumber="204" unit="m" value="1925.0" valueMax="2050.0" valueMin="1800.0">1800–2050 m</elevation>
</quantity>
,
<date id="3F7A1037FF95245819D2FC0C09549CB7" box="[498,611,986,1013]" pageId="3" pageNumber="204" value="2005-06">
<collectingDate id="2F3EE9DFFF95245819D2FC0C09549CB7" box="[498,611,986,1013]" pageId="3" pageNumber="204" value="2005-06">June 2005</collectingDate>
</date>
(fl.),
<emphasis id="79B0EAE5FF9524581A80FC0C08F19CB6" box="[672,966,986,1013]" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF9524581A80FC0C08289CB6" box="[672,799,986,1013]" pageId="3" pageNumber="204">O. Cabrera</collectorName>
&amp;
<collectorName id="E6315321FF9524581B1FFC0C08A09CB7" box="[831,919,987,1013]" pageId="3" pageNumber="204">N. León</collectorName>
869
</emphasis>
(HUTPL, fragm. in GUAY)
</materialsCitation>
;
<materialsCitation id="FBAC3CAAFF9524581D2BFC0C08CF9B5A" ID-GBIF-Occurrence="4127574303" collectingDate="2001-10-20" collectionCode="LOJA, QCNE, MO, GOET" collectorName="J. Homeier" country="Ecuador" elevation="1840" latitude="-3.9666667" location="Loja-Zamora" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
same location,
<quantity id="8C3C9B12FF95245818A8FC290BD79B5B" box="[136,224,1022,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="3" pageNumber="204" unit="m" value="1840.0">
<elevation id="C0E9D1C4FF95245818A8FC290BD79B5B" box="[136,224,1022,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="3" pageNumber="204" unit="m" value="1840.0">1840 m</elevation>
</quantity>
,
<date id="3F7A1037FF95245818CCFC290A9E9B5A" box="[236,425,1022,1048]" pageId="3" pageNumber="204" value="2001-10-20">
<collectingDate id="2F3EE9DFFF95245818CCFC290A9E9B5A" box="[236,425,1022,1048]" pageId="3" pageNumber="204" value="2001-10-20">20 October 2001</collectingDate>
</date>
(fr.),
<emphasis id="79B0EAE5FF95245819CAFC2809919B5A" box="[490,678,1022,1048]" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF95245819CAFC2809509B5A" box="[490,615,1023,1048]" pageId="3" pageNumber="204">J. Homeier</collectorName>
1025
</emphasis>
(
<collectionCode id="2DD5AE32FF9524581A95FC2909CC9B5A" box="[693,763,1022,1048]" pageId="3" pageNumber="204">LOJA</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581B2AFC2908619B5A" box="[778,854,1022,1048]" pageId="3" pageNumber="204">QCNE</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581B45FC2908A59B5A" box="[869,914,1022,1048]" pageId="3" pageNumber="204">MO</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581B80FC2908C79B5A" box="[928,1008,1022,1048]" pageId="3" pageNumber="204">GOET</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="FBAC3CAAFF9524581C24FC2808B39B7E" ID-GBIF-Occurrence="4127574306" collectingDate="2003-09-11" collectionCode="GOET, MO, QCNE" collectorName="J. Homeier &amp; W. Quizhpe" country="Ecuador" elevation="1910" latitude="-3.9666667" location="Loja-Zamora" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
same location,
<quantity id="8C3C9B12FF9524581C8DFC290E329B5B" box="[1197,1285,1022,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="204" unit="m" value="1910.0">
<elevation id="C0E9D1C4FF9524581C8DFC290E329B5B" box="[1197,1285,1022,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="204" unit="m" value="1910.0">1910 m</elevation>
</quantity>
,
<date id="3F7A1037FF9524581D31FC290BF79B7E" pageId="3" pageNumber="204" value="2003-09-11">
<collectingDate id="2F3EE9DFFF9524581D31FC290BF79B7E" pageId="3" pageNumber="204" value="2003-09-11">11 September 2003</collectingDate>
</date>
(sterile),
<emphasis id="79B0EAE5FF952458190CFBF409BC9B7E" box="[300,651,1058,1085]" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF952458190CFBF40A9F9B7E" box="[300,424,1059,1084]" pageId="3" pageNumber="204">J. Homeier</collectorName>
&amp;
<collectorName id="E6315321FF95245819ECFBF4097B9B7E" box="[460,588,1058,1084]" pageId="3" pageNumber="204">W. Quizhpe</collectorName>
1261
</emphasis>
(
<collectionCode id="2DD5AE32FF9524581AB9FBF509D19B7E" box="[665,742,1058,1084]" pageId="3" pageNumber="204">GOET</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581AD2FBF508289B7E" box="[754,799,1058,1084]" pageId="3" pageNumber="204">MO</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581B0DFBF5084E9B7E" box="[813,889,1058,1084]" pageId="3" pageNumber="204">QCNE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="FBAC3CAAFF9524581BB1FBF4097F9B22" ID-GBIF-Occurrence="4127574307" collectingDate="2013-10-20" collectionCode="HUTPL, QCA, GOET" collectorName="J. Homeier" country="Ecuador" elevation="1910" latitude="-3.9666667" location="Loja-Zamora" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
same location,
<quantity id="8C3C9B12FF9524581C1AFBF50FA69B7F" box="[1082,1169,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="204" unit="m" value="1910.0">
<elevation id="C0E9D1C4FF9524581C1AFBF50FA69B7F" box="[1082,1169,1058,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="204" unit="m" value="1910.0">1910 m</elevation>
</quantity>
,
<date id="3F7A1037FF9524581CBCFBF50E6F9B7E" box="[1180,1368,1058,1084]" pageId="3" pageNumber="204" value="2013-10-20">
<collectingDate id="2F3EE9DFFF9524581CBCFBF50E6F9B7E" box="[1180,1368,1058,1084]" pageId="3" pageNumber="204" value="2013-10-20">20 October 2013</collectingDate>
</date>
(fl.),
<emphasis id="79B0EAE5FF9524581DB8FBF40A1D9B22" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF9524581DB8FBF40BDD9B22" pageId="3" pageNumber="204">J. Homeier</collectorName>
5098
</emphasis>
(
<collectionCode id="2DD5AE32FF952458191AFB910AAE9B22" box="[314,409,1094,1120]" pageId="3" pageNumber="204">HUTPL</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581986FB910AD69B22" box="[422,481,1094,1120]" pageId="3" pageNumber="204">QCA</collectionCode>
,
<collectionCode id="2DD5AE32FF95245819D0FB9109779B22" box="[496,576,1094,1120]" pageId="3" pageNumber="204">GOET</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="FBAC3CAAFF9524581A74FB900A959BC6" ID-GBIF-Occurrence="4127574305" collectingDate="2016-12-05" collectingDateMax="2011-04" collectingDateMin="1999-09" collectionCode="HUTPL, QCA, GOET" collectorName="J. Homeier &amp; J. Pena" country="Ecuador" county="Carate" elevation="2040" latitude="-3.9666667" location="Loja-Zamora" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
same location,
<quantity id="8C3C9B12FF9524581B20FB91086F9B23" box="[768,856,1094,1121]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="3" pageNumber="204" unit="m" value="2040.0">
<elevation id="C0E9D1C4FF9524581B20FB91086F9B23" box="[768,856,1094,1121]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="3" pageNumber="204" unit="m" value="2040.0">2040 m</elevation>
</quantity>
,
<date id="3F7A1037FF9524581B45FB910F189B22" box="[869,1071,1094,1121]" pageId="3" pageNumber="204" value="2016-12-05">
<collectingDate id="2F3EE9DFFF9524581B45FB910F189B22" box="[869,1071,1094,1121]" pageId="3" pageNumber="204" value="2016-12-05">5 December 2016</collectingDate>
</date>
(fl.),
<emphasis id="79B0EAE5FF9524581C51FB900E9C9B22" box="[1137,1451,1094,1121]" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF9524581C51FB900FD89B22" box="[1137,1263,1095,1120]" pageId="3" pageNumber="204">J. Homeier</collectorName>
&amp;
<collectorName id="E6315321FF9524581D36FB900E5C9B22" box="[1302,1387,1095,1120]" pageId="3" pageNumber="204">J. Peña</collectorName>
5820
</emphasis>
(
<collectionCode id="2DD5AE32FF95245818B0FBBD0BD89BC6" box="[144,239,1130,1156]" pageId="3" pageNumber="204">HUTPL</collectionCode>
,
<collectionCode id="2DD5AE32FF95245818DBFBBD0A019BC6" box="[251,310,1130,1156]" pageId="3" pageNumber="204">QCA</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581967FBBD0AA09BC6" box="[327,407,1130,1156]" pageId="3" pageNumber="204">GOET</collectionCode>
).
</materialsCitation>
<materialsCitation id="FBAC3CAAFF952458198AFBBD08359BEA" ID-GBIF-Occurrence="4127574301" collectingDate="2011-04" collectingDateMax="2011-04" collectingDateMin="1999-09" collectionCode="LOJA, QCA, QCNE, GOET" collectorName="J. Homeier &amp; F. A. Werner &amp; D. Carate" country="Lebanon" county="Carate" elevation="1720" latitude="-3.9666667" location="Road Loja-Zamora" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
<location id="4E1B602CFF952458198AFBBD09B29BC6" LSID="urn:lsid:plazi:treatment:C36D87E1FF97245818A8FBE30E7E991A:4E1B602CFF952458198AFBBD09B29BC6" box="[426,645,1130,1156]" country="Lebanon" county="Carate" latitude="-3.9666667" longLatPrecision="1307" longitude="-79.066666" municipality="Reserva San Francisco" name="Road Loja-Zamora" pageId="3" pageNumber="204" stateProvince="Loja">Road Loja-Zamora</location>
, El
<collectingCountry id="33D37667FF9524581A94FBBD08319BC6" box="[692,774,1130,1156]" name="Lebanon" pageId="3" pageNumber="204">Líbano</collectingCountry>
,
<geoCoordinate id="2EF05030FF9524581B33FBBD08649BC7" box="[787,851,1130,1157]" degrees="4" direction="south" minutes="5" orientation="latitude" pageId="3" pageNumber="204" precision="925" value="-4.0833335">4°5’S</geoCoordinate>
<geoCoordinate id="2EF05030FF9524581B7CFBBD08F49BC7" box="[860,963,1130,1157]" degrees="78" direction="west" minutes="59" orientation="longitude" pageId="3" pageNumber="204" precision="925" value="-78.98333">78°59’W</geoCoordinate>
,
<quantity id="8C3C9B12FF9524581BF0FBBD0F1E9BC7" box="[976,1065,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="204" unit="m" value="1720.0">
<elevation id="C0E9D1C4FF9524581BF0FBBD0F1E9BC7" box="[976,1065,1130,1157]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="204" unit="m" value="1720.0">1720 m</elevation>
</quantity>
,
<date id="3F7A1037FF9524581C15FBBD0F879BC6" box="[1077,1200,1130,1156]" pageId="3" pageNumber="204" value="2011-04">
<collectingDate id="2F3EE9DFFF9524581C15FBBD0F879BC6" box="[1077,1200,1130,1156]" pageId="3" pageNumber="204" value="2011-04">April 2011</collectingDate>
</date>
(fr.),
<emphasis id="79B0EAE5FF9524581CD4FBBC0A919BEA" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF9524581CD4FBBC0E459BC6" box="[1268,1394,1131,1156]" pageId="3" pageNumber="204">J. Homeier</collectorName>
,
<collectorName id="E6315321FF9524581D5FFBBC0BEE9BEA" pageId="3" pageNumber="204">F.A. Werner</collectorName>
&amp;
<collectorName id="E6315321FF95245818DAFB580A5E9BEB" box="[250,361,1167,1193]" pageId="3" pageNumber="204">D. Cárate</collectorName>
4713
</emphasis>
(
<collectionCode id="2DD5AE32FF9524581993FB590ACE9BEA" box="[435,505,1166,1192]" pageId="3" pageNumber="204">LOJA</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581A26FB5909769BEA" box="[518,577,1166,1192]" pageId="3" pageNumber="204">QCA</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581A6EFB5909AD9BEA" box="[590,666,1166,1192]" pageId="3" pageNumber="204">QCNE</collectionCode>
,
<collectionCode id="2DD5AE32FF9524581A87FB5909C09BEA" box="[679,759,1166,1192]" pageId="3" pageNumber="204">GOET</collectionCode>
).
</materialsCitation>
<materialsCitation id="FBAC3CAAFF9524581B27FB5909039B8E" ID-GBIF-Occurrence="4127574308" collectingDate="1999-09" collectingDateMax="2011-04" collectingDateMin="1999-09" collectionCode="MO" collectorName="W. Palacios" country="Ecuador" county="Carate" elevation="1000" latitude="-3.9666667" location="rio Jamboe" longLatPrecision="1307" longitude="-79.066666" municipality="Cuenca" pageId="3" pageNumber="204" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
<collectingMunicipality id="AB1FAC8DFF9524581B27FB59086C9BEA" box="[775,859,1166,1192]" pageId="3" pageNumber="204">Cuenca</collectingMunicipality>
del
<location id="4E1B602CFF9524581BA8FB590F329BEA" LSID="urn:lsid:plazi:treatment:C36D87E1FF97245818A8FBE30E7E991A:4E1B602CFF9524581BA8FB590F329BEA" box="[904,1029,1166,1192]" country="Ecuador" county="Carate" latitude="-3.9666667" longLatPrecision="1307" longitude="-79.066666" municipality="Cuenca" name="rio Jamboe" pageId="3" pageNumber="204" stateProvince="Loja">río Jamboe</location>
,
<geoCoordinate id="2EF05030FF9524581C2FFB590F699BEA" box="[1039,1118,1166,1192]" degrees="4" direction="south" minutes="08" orientation="latitude" pageId="3" pageNumber="204" precision="925" value="-4.133333">4°08’S</geoCoordinate>
<geoCoordinate id="2EF05030FF9524581C43FB590FFD9BEB" box="[1123,1226,1166,1193]" degrees="78" direction="west" minutes="56" orientation="longitude" pageId="3" pageNumber="204" precision="925" value="-78.933334">78°56’W</geoCoordinate>
,
<quantity id="8C3C9B12FF9524581CF4FB590E1E9BEB" box="[1236,1321,1166,1193]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="204" unit="m" value="1000.0">
<elevation id="C0E9D1C4FF9524581CF4FB590E1E9BEB" box="[1236,1321,1166,1193]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="204" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="3F7A1037FF9524581D14FB590BF79B8E" pageId="3" pageNumber="204" value="1999-09">
<collectingDate id="2F3EE9DFFF9524581D14FB590BF79B8E" pageId="3" pageNumber="204" value="1999-09">September 1999</collectingDate>
</date>
(fl, fr),
<emphasis id="79B0EAE5FF9524581939FB640ADC9B8E" box="[281,491,1202,1228]" italics="true" pageId="3" pageNumber="204">
<collectorName id="E6315321FF9524581939FB640AA99B8E" box="[281,414,1202,1228]" pageId="3" pageNumber="204">W. Palacios</collectorName>
14881
</emphasis>
(
<collectionCode id="2DD5AE32FF95245819DAFB65091E9B8E" box="[506,553,1202,1228]" pageId="3" pageNumber="204">MO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="03DE657CFF952458189DFB010E7E991A" pageId="3" pageNumber="204" type="discussion">
<paragraph id="4B7B36F7FF952458189DFB010E7E991A" blockId="3.[136,1452,158,1625]" pageId="3" pageNumber="204">
<emphasis id="79B0EAE5FF952458189DFB010A729BB2" bold="true" box="[189,325,1238,1264]" pageId="3" pageNumber="204">Discussion:</emphasis>
—The fruits of
<taxonomicName id="8CC44D74FF9524581A26FB0008769BB2" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[518,833,1238,1264]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF9524581A26FB0008769BB2" box="[518,833,1238,1264]" italics="true" pageId="3" pageNumber="204">Heisteria austroecuadorica</emphasis>
</taxonomicName>
are characterized by the apex presenting a circular depression, a key character in Sleumer’s treatment (1984) that is only found in three other species in tropical America:
<taxonomicName id="8CC44D74FF95245818A8FAC809039A7A" authority="Sleumer (1984: 49)" authorityName="Sleumer" authorityPageNumber="49" authorityYear="1984" box="[136,564,1310,1336]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="amphoricarpa">
<emphasis id="79B0EAE5FF95245818A8FAC80A7B9A7A" box="[136,332,1310,1336]" italics="true" pageId="3" pageNumber="204">H. amphoricarpa</emphasis>
<bibRefCitation id="2F554B06FF9524581975FAC909039A7A" author="Sleumer, H. O." box="[341,564,1310,1336]" pageId="3" pageNumber="204" pagination="1 - 159" refId="ref3668" refString="Sleumer, H. O. (1984) Olacaceae. Flora Neotropica 38: 1 - 159." type="journal article" year="1984">Sleumer (1984: 49)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8CC44D74FF9524581A61FAC809FA9A7A" authorityName="Sleumer" authorityYear="1984" box="[577,717,1310,1336]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="asplundii">
<emphasis id="79B0EAE5FF9524581A61FAC809FA9A7A" box="[577,717,1310,1336]" italics="true" pageId="3" pageNumber="204">H. asplundii</emphasis>
</taxonomicName>
, and
<taxonomicName id="8CC44D74FF9524581B2EFAC80F459A7B" authority="Sleumer (1984: 50)" authorityName="Sleumer" authorityPageNumber="50" authorityYear="1984" box="[782,1138,1310,1337]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="skutchii">
<emphasis id="79B0EAE5FF9524581B2EFAC808BE9A7A" box="[782,905,1310,1336]" italics="true" pageId="3" pageNumber="204">H. skutchii</emphasis>
<bibRefCitation id="2F554B06FF9524581BB2FAC90F459A7B" author="Sleumer, H. O." box="[914,1138,1310,1337]" pageId="3" pageNumber="204" pagination="1 - 159" refId="ref3668" refString="Sleumer, H. O. (1984) Olacaceae. Flora Neotropica 38: 1 - 159." type="journal article" year="1984">Sleumer (1984: 50)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="8CC44D74FF9524581C5EFAC80EB89A7A" authorityName="Sleumer" authorityYear="1984" box="[1150,1423,1310,1336]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="amphoricarpa">
<emphasis id="79B0EAE5FF9524581C5EFAC80EB89A7A" box="[1150,1423,1310,1336]" italics="true" pageId="3" pageNumber="204">Heisteria amphoricarpa</emphasis>
</taxonomicName>
, a Brazilian species, has a unique urn-shaped calyx enveloping most of the drupe and flowers with five stamens, while
<taxonomicName id="8CC44D74FF95245818A8FAB00A359AC2" authorityName="Sleumer" authorityYear="1984" box="[136,258,1382,1408]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="skutchii">
<emphasis id="79B0EAE5FF95245818A8FAB00A359AC2" box="[136,258,1382,1408]" italics="true" pageId="3" pageNumber="204">H. skutchii</emphasis>
</taxonomicName>
, a Costa Rican species with 10 stamens, has much wider, patent fruiting calyx (over
<quantity id="8C3C9B12FF9524581CE3FAB10E249AC3" box="[1219,1299,1382,1409]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="204" unit="mm" value="30.0">30 mm</quantity>
diameter). In
<collectingCountry id="33D37667FF95245818A8FA5D0BD09AE6" box="[136,231,1418,1444]" name="Ecuador" pageId="3" pageNumber="204">Ecuador</collectingCountry>
, the fruits of
<taxonomicName id="8CC44D74FF952458195BFA5C09989AE6" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[379,687,1418,1444]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF952458195BFA5C09989AE6" box="[379,687,1418,1444]" italics="true" pageId="3" pageNumber="204">Heisteria austroecuadorica</emphasis>
</taxonomicName>
are relatively large with a depressed apex like those of
<taxonomicName id="8CC44D74FF9524581D3AFA5C0E939AE6" authorityName="Sleumer" authorityYear="1984" box="[1306,1444,1418,1444]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="asplundii">
<emphasis id="79B0EAE5FF9524581D3AFA5C0E939AE6" box="[1306,1444,1418,1444]" italics="true" pageId="3" pageNumber="204">H. asplundii</emphasis>
</taxonomicName>
, a tree endemic to northwestern
<collectingCountry id="33D37667FF95245819CBFA79097C9A8A" box="[491,587,1454,1480]" name="Ecuador" pageId="3" pageNumber="204">Ecuador</collectingCountry>
. However,
<taxonomicName id="8CC44D74FF9524581AEAFA7808859A8A" authorityName="Cornejo, Homeier &amp; C. Ulloa. A, Inflorescence" authorityYear="2023" box="[714,946,1454,1480]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="austroecuadorica">
<emphasis id="79B0EAE5FF9524581AEAFA7808859A8A" box="[714,946,1454,1480]" italics="true" pageId="3" pageNumber="204">H. austroecuadorica</emphasis>
</taxonomicName>
immediately differs from
<taxonomicName id="8CC44D74FF9524581CFBFA780E529A8A" authorityName="Sleumer" authorityYear="1984" box="[1243,1381,1454,1480]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="asplundii">
<emphasis id="79B0EAE5FF9524581CFBFA780E529A8A" box="[1243,1381,1454,1480]" italics="true" pageId="3" pageNumber="204">H. asplundii</emphasis>
</taxonomicName>
by its narrower leaves, with blades elliptic-lanceolate to narrowly elliptic or narrowly lanceolate,
<quantity id="8C3C9B12FF9524581C5AFA050FFB9AAF" box="[1146,1228,1490,1517]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="3" pageNumber="204" unit="cm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 cm</quantity>
wide (
<emphasis id="79B0EAE5FF9524581D38FA040E579AAE" box="[1304,1376,1491,1516]" italics="true" pageId="3" pageNumber="204">versus</emphasis>
leaves always with broadly elliptic blades,
<quantity id="8C3C9B12FF9524581A3EFA2109BA9953" box="[542,653,1526,1553]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="204" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
wide in
<taxonomicName id="8CC44D74FF9524581ACFFA20084E9952" authorityName="Sleumer" authorityYear="1984" box="[751,889,1526,1552]" class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" kingdom="Plantae" order="Santalales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="asplundii">
<emphasis id="79B0EAE5FF9524581ACFFA20084E9952" box="[751,889,1526,1552]" italics="true" pageId="3" pageNumber="204">H. asplundii</emphasis>
</taxonomicName>
), the flowers with 10 stamens (
<emphasis id="79B0EAE5FF9524581CFDFA200FCC9952" box="[1245,1275,1527,1552]" italics="true" pageId="3" pageNumber="204">vs.</emphasis>
5 stamens), the smaller fruiting calyx, up to
<quantity id="8C3C9B12FF95245819E4F9CD09389977" box="[452,527,1562,1589]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="204" unit="cm" value="2.8">2.8 cm</quantity>
and covering just the base of the drupe (
<emphasis id="79B0EAE5FF9524581BF3F9CC08C69976" box="[979,1009,1563,1588]" italics="true" pageId="3" pageNumber="204">vs.</emphasis>
up to
<quantity id="8C3C9B12FF9524581C16F9CD0FB69977" box="[1078,1153,1562,1589]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="204" unit="cm" value="4.8">4.8 cm</quantity>
and covering the basal 1/3 of the drupe); additionally, these two species are widely separated on opposite sides of the Andes mountains.
</paragraph>
</subSubSection>
</treatment>
</document>
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