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<document id="2E7760D6159C0B44926DCDBE3F6F3EFE" ID-CLB-Dataset="30483" ID-DOI="10.11646/zootaxa.4397.1.1" ID-GBIF-Dataset="b44266be-2443-4676-abd8-015e1e069156" ID-ISSN="1175-5326" ID-Zenodo-Dep="1202783" ID-ZooBank="DAB14765-7C9C-41FF-9ECF-563B82B9D258" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521441517859" checkinUser="plazi" docAuthor="Amr, Zuhair S., Abu, Mohammad A., Qumsiyeh, Mazin &amp; Eid, Ehab" docDate="2018" docId="C32887CBFFA6BA54FF3DFE08FB3BEFC9" docLanguage="en" docName="zootaxa.4397.1.1.pdf" docOrigin="Zootaxa 4397 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Psammomys obesus Cretzschmar 1828" docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="3F11FFB3FF90BA63FFAAFFF7FFEDEA22" masterDocTitle="Systematics, distribution and ecological analysis of rodents in Jordan" masterLastPageNumber="94" masterPageNumber="1" pageNumber="55" updateTime="1698532151927" updateUser="ExternalLinkService">
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<mods:title id="1A5AE0D893A4CE096C64600A14F525A3">Systematics, distribution and ecological analysis of rodents in Jordan</mods:title>
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<treatment id="C32887CBFFA6BA54FF3DFE08FB3BEFC9" ID-DOI="http://doi.org/10.5281/zenodo.5991832" ID-GBIF-Taxon="142340525" ID-Zenodo-Dep="5991832" LSID="urn:lsid:plazi:treatment:C32887CBFFA6BA54FF3DFE08FB3BEFC9" httpUri="http://treatment.plazi.org/id/C32887CBFFA6BA54FF3DFE08FB3BEFC9" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
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<paragraph id="4B3E36DDFFA6BA55FF3DFE08FD95E838" blockId="54.[151,632,511,571]" box="[151,632,511,538]" pageId="54" pageNumber="55">
<heading id="107681B1FFA6BA55FF3DFE08FD95E838" bold="true" box="[151,632,511,538]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<taxonomicName id="8C814D5EFFA6BA55FF3DFE08FD95E838" ID-CoL="4NCRG" authority="Cretzschmar, 1828" authorityName="Cretzschmar" authorityYear="1828" box="[151,632,511,538]" class="Mammalia" family="Muridae" genus="Psammomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="obesus">
<emphasis id="79F5EACFFFA6BA55FF3DFE08FD95E838" bold="true" box="[151,632,511,538]" pageId="54" pageNumber="55">
<emphasis id="79F5EACFFFA6BA55FF3DFE08FE6AE83B" bold="true" box="[151,391,511,537]" italics="true" pageId="54" pageNumber="55">Psammomys obesus</emphasis>
Cretzschmar, 1828
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4B3E36DDFFA6BA55FF3DFDD5FE0AE819" blockId="54.[151,632,511,571]" box="[151,487,546,571]" pageId="54" pageNumber="55">
<emphasis id="79F5EACFFFA6BA55FF3DFDD5FEBAE819" bold="true" box="[151,343,546,571]" pageId="54" pageNumber="55">Common name:</emphasis>
Fat sand rat.
</paragraph>
</subSubSection>
<subSubSection id="039B6556FFA6BA55FF3DFD9DFB5CE981" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="4B3E36DDFFA6BA55FF3DFD9DFD66E931" blockId="54.[151,1436,618,1147]" pageId="54" pageNumber="55">
<emphasis id="79F5EACFFFA6BA55FF3DFD9DFEE7E8A1" bold="true" box="[151,266,618,643]" pageId="54" pageNumber="55">Diagnosis</emphasis>
: Large rodent. Fur dorsally sandy to yellowish in color, ventral side grayish white, line of demarcation not very distinct (
<figureCitation id="D3BA2A58FFA6BA55FEF6FD79FE23E885" box="[348,462,654,679]" captionStart="FIGURE 62" captionStartId="54.[151,250,1960,1982]" captionTargetBox="[264,1323,1199,1938]" captionTargetId="figure@54.[264,1323,1199,1938]" captionTargetPageId="54" captionText="FIGURE 62. The fat sand rat, Psammomys obesus, from Swaqa area." httpUri="https://zenodo.org/record/1202899/files/figure.png" pageId="54" pageNumber="55">Figure 62</figureCitation>
). No white patches behind ears or above eyes. Ears densely haired and short. Feet with black claws. Long hair on hindfeet soles, with naked heel. Tail short and thick terminating with a black tuft. Skull robust and very angular with well-developed ridges. Upper incisors without grooves, else the crown structure of molars very similar to
<taxonomicName id="8C814D5EFFA6BA55FE3EFD0BFE13E931" box="[404,510,764,787]" class="Mammalia" family="Muridae" genus="Meriones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="79F5EACFFFA6BA55FE3EFD0BFE13E931" box="[404,510,764,787]" italics="true" pageId="54" pageNumber="55">Meriones</emphasis>
</taxonomicName>
(
<figureCitation id="D3BA2A58FFA6BA55FDA7FD0DFD93E931" box="[525,638,762,787]" captionStart="FIGURE 63" captionStartId="55.[151,250,1285,1307]" captionTargetBox="[264,1323,193,1264]" captionTargetId="figure@55.[264,1323,193,1264]" captionTargetPageId="55" captionText="FIGURE 63. A. Dorsal,B. Ventral, C. Lateral views of the skull, and D. Upper maxillary teeth for Psammomys obesus." httpUri="https://zenodo.org/record/1202901/files/figure.png" pageId="54" pageNumber="55">Figure 63</figureCitation>
).
</paragraph>
<paragraph id="4B3E36DDFFA6BA55FF6DFCE9FB5CE981" blockId="54.[151,1436,618,1147]" pageId="54" pageNumber="55">
<emphasis id="79F5EACFFFA6BA55FF6DFCE9FDF6E915" bold="true" box="[199,539,798,823]" pageId="54" pageNumber="55">Localities: Previous records.</emphasis>
Al Muwaqqar (
<bibRefCitation id="2F104B2CFFA6BA55FD64FCE8FC54E915" box="[718,953,798,823]" pageId="54" pageNumber="55" refId="ref31164" refString="Amr, Z. S. &amp; Saliba, E. K. (1986) Ecological observations on the Fat Jird, Psammomys obesus dianae, in the Mowaqqar area of Jordan. Dirasat, 13 (7), 155 - 161." type="journal article">Amr &amp; Saliba, 1986</bibRefCitation>
); Al Muwaqqar, Qaşr al Ḩallābāt,
<quantity id="8C799B38FFA6BA55FA94FCE9FA68E915" box="[1342,1413,798,823]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="54" pageNumber="55" unit="km" value="60.0">60 km</quantity>
S Ammān (
<bibRefCitation id="2F104B2CFFA6BA55FEA0FCB4FE0CE979" box="[266,481,834,859]" pageId="54" pageNumber="55" refId="ref31125" refString="Amr, Z. S. &amp; Disi, A. M. (1988) Jordanian Mammals Acquired by the Jordan University Natural History Museum. Publication of the University of Jordan, Amman, 32 pp." type="book">Amr &amp; Disi, 1988</bibRefCitation>
);
<quantity id="8C799B38FFA6BA55FE5CFCB4FDD2E979" box="[502,575,834,859]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="54" pageNumber="55" unit="km" value="20.0">20 km</quantity>
NW El Quweira, Wādī Ramm (Abu Baker &amp; Amr, 2004); Ḑānā Biosphere Reserve (Yousef &amp;
<bibRefCitation id="2F104B2CFFA6BA55FED2FC90FE1AE95D" box="[376,503,870,895]" pageId="54" pageNumber="55" refId="ref30412" refString="Abu Baker, M., Qarqaz, M. &amp; Amr, Z. (2005) Small mammal remains recovered from pellets of the Barn Owl (Tyto alba) at Shawmari Wildlife Reserve, eastern Jordan. Casopis Narodniho Muzea, Rada prirodovedna, 174 (1 - 4), 12 7 - 129." type="journal article">Amr, 2005</bibRefCitation>
).
<emphasis id="79F5EACFFFA6BA55FDA1FC91FD44E95D" bold="true" box="[523,681,870,895]" pageId="54" pageNumber="55">New records.</emphasis>
Al-Hassa, Al Jafr, Al Lajjun, Al Qaţrānah, Ash Shawmarī Nature Reserve, Aqaba, Azraq, Dhabaah, Jabal Masuda, Ma'an, Swaqa, Umm ar Rasas (
<figureCitation id="D3BA2A58FFA6BA55FB99FC7DFB48E981" box="[1075,1189,906,931]" captionStart="FIGURE 64" captionStartId="56.[151,250,1423,1445]" captionTargetBox="[275,1307,228,1368]" captionTargetId="figure@56.[264,1323,193,1401]" captionTargetPageId="56" captionText="FIGURE 64. Distribution of the fat sand rat, Psammomys obesus." httpUri="https://zenodo.org/record/1202903/files/figure.png" pageId="54" pageNumber="55">Figure 64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="039B6556FFA6BA55FF6DFC59FC8EEE59" pageId="54" pageNumber="55" type="description">
<paragraph id="4B3E36DDFFA6BA55FF6DFC59FC8EEE59" blockId="54.[151,1436,618,1147]" pageId="54" pageNumber="55">
<emphasis id="79F5EACFFFA6BA55FF6DFC59FEC1E9E5" bold="true" box="[199,300,942,967]" pageId="54" pageNumber="55">Habitat:</emphasis>
The ecology of this species was studied by
<bibRefCitation id="2F104B2CFFA6BA55FCB5FC58FBFCE9E5" box="[799,1041,942,967]" pageId="54" pageNumber="55" refId="ref31164" refString="Amr, Z. S. &amp; Saliba, E. K. (1986) Ecological observations on the Fat Jird, Psammomys obesus dianae, in the Mowaqqar area of Jordan. Dirasat, 13 (7), 155 - 161." type="journal article">Amr &amp; Saliba (1986)</bibRefCitation>
, where they reported on its diurnal activity, feeding habits, burrow system and association with other animals. The fat sand rat is a colonial species forming large colonies constructed close to
<emphasis id="79F5EACFFFA6BA55FD3FFC01FD11EE2D" box="[661,764,1014,1039]" italics="true" pageId="54" pageNumber="55">Anabasis</emphasis>
sp. shrubs. It was found to share burrow with the Syrian hamster. In some areas, previously known to harbor dense colonies of this jird disappeared entirely; where as new colonies are established in new areas away from the original foci. Distribution of this species is associated with the presence of chenopods (i.e.
<emphasis id="79F5EACFFFA6BA55FE65FB95FD46EE59" box="[463,683,1122,1147]" italics="true" pageId="54" pageNumber="55">
Anabasis
<taxonomicName id="8C814D5EFFA6BA55FD97FB95FD46EE59" box="[573,683,1122,1147]" class="Mammalia" family="Muridae" genus="Acomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="articulata">articulata</taxonomicName>
</emphasis>
,
<taxonomicName id="8C814D5EFFA6BA55FD13FB94FCC1EE59" box="[697,812,1123,1147]" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="syriaca">
<emphasis id="79F5EACFFFA6BA55FD13FB94FCC1EE59" box="[697,812,1123,1147]" italics="true" pageId="54" pageNumber="55">A. syriaca</emphasis>
</taxonomicName>
etc).
</paragraph>
</subSubSection>
<caption id="1FFE6655FFA6BA55FF3DF85FFCB4ED9C" box="[151,857,1960,1982]" httpUri="https://zenodo.org/record/1202899/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1960,1982]" targetBox="[264,1323,1199,1938]" targetPageId="54">
<paragraph id="4B3E36DDFFA6BA55FF3DF85FFCB4ED9C" blockId="54.[151,857,1960,1982]" box="[151,857,1960,1982]" pageId="54" pageNumber="55">
<emphasis id="79F5EACFFFA6BA55FF3DF85FFEF2ED9F" bold="true" box="[151,287,1960,1982]" pageId="54" pageNumber="55">FIGURE 62.</emphasis>
The fat sand rat,
<taxonomicName id="8C814D5EFFA6BA55FE7AF85EFD77ED9C" authorityName="Cretzschmar" authorityYear="1828" box="[464,666,1960,1982]" class="Mammalia" family="Muridae" genus="Psammomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="obesus">
<emphasis id="79F5EACFFFA6BA55FE7AF85EFD77ED9C" box="[464,666,1960,1982]" italics="true" pageId="54" pageNumber="55">Psammomys obesus</emphasis>
</taxonomicName>
, from Swaqa area.
</paragraph>
</caption>
<caption id="1FFE6655FFA7BA54FF3DFAF2FABAEF39" box="[151,1367,1285,1307]" httpUri="https://zenodo.org/record/1202901/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1285,1307]" targetBox="[264,1323,193,1264]" targetPageId="55">
<paragraph id="4B3E36DDFFA7BA54FF3DFAF2FABAEF39" blockId="55.[151,1367,1285,1307]" box="[151,1367,1285,1307]" pageId="55" pageNumber="56">
<emphasis id="79F5EACFFFA7BA54FF3DFAF2FED3EF38" bold="true" box="[151,318,1285,1307]" pageId="55" pageNumber="56">FIGURE 63. A.</emphasis>
Dorsal,
<emphasis id="79F5EACFFFA7BA54FE3EFAF1FE41EF39" bold="true" box="[404,428,1286,1307]" pageId="55" pageNumber="56">B.</emphasis>
Ventral,
<emphasis id="79F5EACFFFA7BA54FDA2FAF2FDCDEF39" bold="true" box="[520,544,1285,1307]" pageId="55" pageNumber="56">C.</emphasis>
Lateral views of the skull, and
<emphasis id="79F5EACFFFA7BA54FCCAFAF1FC9FEF39" bold="true" box="[864,882,1286,1307]" pageId="55" pageNumber="56">D</emphasis>
. Upper maxillary teeth for
<taxonomicName id="8C814D5EFFA7BA54FB2DFAF1FABCEF39" authorityName="Cretzschmar" authorityYear="1828" box="[1159,1361,1285,1307]" class="Mammalia" family="Muridae" genus="Psammomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="obesus">
<emphasis id="79F5EACFFFA7BA54FB2DFAF1FABCEF39" box="[1159,1361,1285,1307]" italics="true" pageId="55" pageNumber="56">Psammomys obesus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="039B6556FFA7BA54FF6DFAB5FF32EF5D" pageId="55" pageNumber="56" type="biology_ecology">
<paragraph id="4B3E36DDFFA7BA54FF6DFAB5FF32EF5D" blockId="55.[151,1436,1346,1515]" pageId="55" pageNumber="56">
<emphasis id="79F5EACFFFA7BA54FF6DFAB5FECCEF79" bold="true" box="[199,289,1346,1371]" pageId="55" pageNumber="56">Biology</emphasis>
: Gestation period may last for 2325 days, and females give birth to 28 young (Osborn &amp; Helmy, 1980).
</paragraph>
</subSubSection>
<subSubSection id="039B6556FFA7BA54FF6DFA7DFB3BEFC9" pageId="55" pageNumber="56" type="discussion">
<paragraph id="4B3E36DDFFA7BA54FF6DFA7DFB3BEFC9" blockId="55.[151,1436,1346,1515]" pageId="55" pageNumber="56">
<emphasis id="79F5EACFFFA7BA54FF6DFA7DFED9EF81" bold="true" box="[199,308,1418,1443]" pageId="55" pageNumber="56">Remarks</emphasis>
: This is another important reservoir animal for human leishmaniasis, a disease known in
<collectingCountry id="3396764DFFA7BA54FA88FA7DFA80EF81" box="[1314,1389,1418,1443]" name="Jordan" pageId="55" pageNumber="56">Jordan</collectingCountry>
and associated with areas colonized by this jird. The karyotype for specimens collected from
<collectingCountry id="3396764DFFA7BA54FB2CFA59FB3CEFE5" box="[1158,1233,1454,1479]" name="Jordan" pageId="55" pageNumber="56">Jordan</collectingCountry>
yeiled 2n=48 and FN=74, 75 with 18 acrocentric or telocentric and 28 biarmed autosomes (Qumsiyeh
<emphasis id="79F5EACFFFA7BA54FBE2FA23FB94EFC9" box="[1096,1145,1490,1515]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
., 1986).
</paragraph>
</subSubSection>
</treatment>
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