treatments-xml/data/42/44/43/424443F5846857F6A98F13BACE092ADF.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.15.3084" ID-PMC="PMC3492929" ID-Pensoft-Pub="1314-2003-15-1" ID-Pensoft-UUID="FF9AC356FFDAFB26FF95FFE3FFA1520C" ID-PubMed="23185125" ID-Zenodo-Dep="576134" ModsDocID="1314-2003-15-1" checkinTime="1451251993896" checkinUser="pensoft" docAuthor="Soreng, Robert J. &amp; Peterson, Paul M." docDate="2012" docId="424443F5846857F6A98F13BACE092ADF" docLanguage="en" docName="PhytoKeys 15: 1-104" docOrigin="PhytoKeys 15" docPubDate="2012-08-06" docSource="http://dx.doi.org/10.3897/phytokeys.15.3084" docTitle="Poa palmeri Soreng &amp; P. M. Peterson 2012, sp. nov." docType="treatment" docVersion="3" id="FF9AC356FFDAFB26FF95FFE3FFA1520C" lastPageNumber="43" masterDocId="FF9AC356FFDAFB26FF95FFE3FFA1520C" masterDocTitle="Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: new records, re-evaluation of P. ruprechtii, and two new species, P. palmeri and P. wendtii" masterLastPageNumber="104" masterPageNumber="1" pageNumber="41" updateTime="1668139833991" updateUser="ExternalLinkService">
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<mods:title>Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: new records, re-evaluation of P. ruprechtii, and two new species, P. palmeri and P. wendtii</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soreng, Robert J.</mods:namePart>
<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peterson, Paul M.</mods:namePart>
<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-08-06</mods:number>
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<mods:detail type="volume">
<mods:number>15</mods:number>
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<mods:start>1</mods:start>
<mods:end>104</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.15.3084</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.15.3084</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-15-1</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576134</mods:identifier>
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<treatment ID-GBIF-Taxon="182229300" LSID="urn:lsid:plazi:treatment:424443F5846857F6A98F13BACE092ADF" httpUri="http://treatment.plazi.org/id/424443F5846857F6A98F13BACE092ADF" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection pageId="40" pageNumber="41" type="nomenclature">
<paragraph pageId="40" pageNumber="41">
15. 
<taxonomicName LSID="424443F5-8468-57F6-A98F-13BACE092ADF" authority="Soreng &amp; P. M. Peterson" authorityName="Soreng &amp; P. M. Peterson" authorityYear="2012" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa palmeri" order="Poales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="palmeri" status="sp. nov.">Poa palmeri Soreng &amp; P.M. Peterson</taxonomicName>
<taxonomicNameLabel pageId="40" pageNumber="41">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-E Poa gymnantha Pilg. A spikelet B lemma and palea C palea D staminode and lodicules (pistillate-flower) E pistil (pistillate-flower) F-I Poa chamaeclinos Pilg. F spikelet G floret H palea I pistil (pistillate-flower) J Poa palmeri Soreng &amp; P. M. Peterson J spikelet K-M Poa strictiramea Hitchc. K spikelet L floret M palea N-P Poa calycina var. mathewsii (Ball) Refulio N spikelet O floret P palea. A-E drawn from Peterson 12863 et al. from Peru F-I drawn from Soreng 3315 &amp; Soreng; J drawn from Peterson 18790 &amp; Valdes-Reyna K-M drawn from Soreng 3204 &amp; Spellenberg N-P drawn from Beaman 1732." httpUri="https://binary.pensoft.net/fig/10141" pageId="40" pageNumber="41">Figs 6J</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Poa palmeri Soreng &amp; P. M. Peterson. A habit B inflorescence C sheath, ligule, blade D spikelet E glumes F floret G callus of floret H lemma dorsal view I palea dorsal and ventral views J stamen K pistil from perfect-flowered plant, with lodicules attached to palea L rudimentary pistil from staminate plant, with lodicules. A-J and L drawn from holotype collection (Pringle 10212) K drawn from Palmer 1366." httpUri="https://binary.pensoft.net/fig/10150" pageId="40" pageNumber="41">15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="type">
<paragraph pageId="40" pageNumber="41">Type:</paragraph>
<paragraph pageId="40" pageNumber="41">
Mexico. Nuevo 
<normalizedToken originalValue="León">Leon</normalizedToken>
, Sierra Madre [Oriental] above Monterey [ca 
<geoCoordinate degrees="25" direction="north" minutes="38" orientation="latitude" precision="925" value="25.633333">25°38'N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="west" minutes="38" orientation="longitude" precision="925" value="-100.63333">100°38'W</geoCoordinate>
], calcareous ledges, 3000 ft [misprint for 8000 ft; 2440 m], 31 Mar 1906, 
<emphasis italics="true" pageId="40" pageNumber="41">C.G.Pringle 10212</emphasis>
(holotype: US-462254!♂; isotypes; GH!, MEXU-539725!♂, MO-1770420!♀, MSC, NMC!, NY-431382!, WYAC).
</paragraph>
</subSubSection>
<subSubSection lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
<paragraph pageId="40" pageNumber="41">Diagnosis.</paragraph>
<paragraph lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
Differing from 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="40" pageNumber="41">Poa ruprechtii</emphasis>
</taxonomicName>
Peyr. by leaf collar margins flared (versus not flared), ligules of upper culm leaves truncate (versus obtuse to acute), flag leaf sheaths fused by an invaginated hyaline membrane for much of the upper fusion length (versus abruptly fused by firmly chartaceous margins), spikelet pedicles elongated and moderately to densely scabrous (versus shorter and usually densely to more coarsely scabrous), distal rachilla internodes usually 1-2 mm long (versus less than 1 mm), lemmas thinly chartaceous, intermediate veins faint to moderately 
<pageBreakToken pageId="41" pageNumber="42" start="start">prominent</pageBreakToken>
, not extending to near the upper margin, margin and apex fairly broadly scarious-hyaline (versus lemmas chartaceous, intermediate veins prominent, extending nearly to the margin, margin and apex narrowly scarious), anthers 1.6-2.8 (versus 0.7-1.05[-1.25]) mm.
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph pageId="41" pageNumber="42">Description.</paragraph>
<paragraph lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
Trioecious (mostly hermaphroditic, but some pistillate or staminate). 
<emphasis bold="true" pageId="41" pageNumber="42">Perennials</emphasis>
; tufted, sometimes sub-rhizomatous to rhizomatous, tufts dense to loose, generally medium girth and height, green or bluish-grey-green; tillers extravaginal (basally cataphyllous), and/or mainly intravaginal (each subtended by a single elongated, 2-keeled longitudinally split prophyll), with lateral or downward tending, cataphyllous shoots, or without them. 
<emphasis bold="true" pageId="41" pageNumber="42">Culms</emphasis>
(26-)50-70 cm tall, erect or bases slightly decumbent, fairly slender, not branching above the base, leafy, terete or weakly compressed, smooth; nodes 3-4, terete or slightly compressed, 1-2 exerted. 
<emphasis bold="true" pageId="41" pageNumber="42">Leaf</emphasis>
sheaths slightly compressed, smooth or sparsely to infrequently densely scabrous, glabrous or rarely retrorsely hirtellous to strigulose; butt sheaths papery, smooth, glabrous; flag leaf sheaths (5-)7-18 cm long, margins fused 34-65% the length, fused by an invaginated hyaline margin for much of that length, 1.1-3.1 
<normalizedToken originalValue="×">x</normalizedToken>
longer than its blade; collar margins of lower and mid culm leaves usually flared, smooth or scabrous, glabrous or more often ciliate; ligules 0.8-3 mm long, scarious to hyaline, adaxially scabrous or puberulent, upper margins often ciliolate, apices truncate and entire below to obtuse and irregularly dentate/lacerate above, sterile shoot ligules like those of the culm leaves; blades 2-30 cm long, (1.5-)2-3 mm wide, flat and slightly lax to involute and moderately firm, abaxially and adaxially smooth or lightly scabrous over the veins, margins scabrous, narrowly prow-tipped; lower mid-cauline blades the longest, 10-30 cm long, shorter upward, flag leaf blades 27-50 (90)% longer than their sheaths, flag leaf blade 2.3-10.8 cm long; sterile shoot blades similar to cauline blades or more involute. 
<emphasis bold="true" pageId="41" pageNumber="42">Panicles</emphasis>
6.5-20 cm long, erect, open, usually trapezoidal to pyramidal, sparse to moderately congested, with (18-)40-100 spikelets, proximal internode (2-)3-5 cm long; rachis with (1-)2-4(-5) branches per node; primary branches spreading to eventually reflexed, fairly flexuous, terete to weakly angled, sparsely to moderately scabrous; lateral pedicels 1/2 to equaling the spikelets, moderately to densely scabrous, prickles fine to moderately coarse; longest branches 4-10 cm, with 3-34 spikelets in distal 1/2, loosely arranged. 
<emphasis bold="true" pageId="41" pageNumber="42">Spikelets</emphasis>
3-8 mm long, to 3.5 
<normalizedToken originalValue="×">x</normalizedToken>
long as wide, lanceolate, laterally compressed (sexually dimorphic - staminate spikelets with more florets, up to 7-9); spikelets infrequently bulbiferous; florets (2-)3-6(-9), pistillate, staminate, or hermaphroditic; rachilla internodes terete, distal internodes terete, exceeding 1 mm long, smooth or lightly scabrous, glabrous; glumes narrow lanceolate, distinctly keeled; lower glumes 1.6-3.5 mm long, 1/2-2/3 as long as adjacent lemmas, 1-veined; upper glumes 2.2-4.9 mm long, 3-veined; calluses dorsally webbed, web scant to distinct, hairs 1-2 mm long, woolly; lemmas 2.6-5 mm long, lanceolate, 5-veined, body thinly chartaceous, distinctly keeled, keels to 1/3-2/3 and marginal veins to 1/5-1/3 sparsely short to long villous, intermediate veins smooth or sparsely scabrous, glabrous, between veins smooth, glabrous, intermediate veins obscure to moderately prominent, not extending to near the margin, 
<pageBreakToken pageId="42" pageNumber="43" start="start">margins</pageBreakToken>
smooth, broadly scarious-hyaline, smooth, apices acute; paleas scabrous, medially rarely softly puberulent over the keels. between keels narrow (0.3-0.4 mm), muriculate, scabrous to sparsely puberulent. 
<emphasis bold="true" pageId="42" pageNumber="43">Flowers</emphasis>
chasmogamous; lodicules 0.5 mm long, lanceolate, with a narrow lateral lobe; anthers 1.6-2.8 mm long, sometimes late aborted, infrequently vestigial throughout individual plants. 
<emphasis bold="true" pageId="42" pageNumber="43">Caryopses</emphasis>
1.8-2.1 mm long, elliptical in side-view, sulcus broad and shallow, brown, hilum 0.2 mm long, round, grain adherent to the palea. 2 
<emphasis italics="true" pageId="42" pageNumber="43">n</emphasis>
= unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="distribution">
<paragraph pageId="42" pageNumber="43">Distribution.</paragraph>
<paragraph pageId="42" pageNumber="43">
The species is endemic to the Sierra Madre Oriental and is found in Coahuila and Nuevo 
<normalizedToken originalValue="León">Leon</normalizedToken>
, Mexico.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="ecology">
<paragraph pageId="42" pageNumber="43">Ecology.</paragraph>
<paragraph pageId="42" pageNumber="43">
This species is found on rocky calcareous substrates in shaded and open forests associated with 
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pinales" pageId="42" pageNumber="43" phylum="Coniferophyta" rank="genus">
<emphasis italics="true" pageId="42" pageNumber="43">Pinus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fagales" pageId="42" pageNumber="43" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="42" pageNumber="43">Quercus</emphasis>
</taxonomicName>
, and 
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pinales" pageId="42" pageNumber="43" phylum="Coniferophyta" rank="genus">
<emphasis italics="true" pageId="42" pageNumber="43">Abies</emphasis>
</taxonomicName>
; between 1750-3760 m. Flowering April through October.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="conservation status">
<paragraph pageId="42" pageNumber="43">Conservation status.</paragraph>
<paragraph pageId="42" pageNumber="43">The species is a regional endemic, known from only 15 collections over 1400 km2.</paragraph>
<paragraph pageId="42" pageNumber="43">
<emphasis bold="true" pageId="42" pageNumber="43">Etymology.</emphasis>
The new species is named for Dr. Edward Palmer (1829-1911), an important early collector for the U.S. Department of Agriculture, known particularly for his work in southwestern USA and northern Mexico, who, in 1880, was the first to collect this plant. We selected the 
<taxonomicName genus="Pringle" lsidName="" pageId="42" pageNumber="43" rank="genus">
<emphasis italics="true" pageId="42" pageNumber="43">Pringle</emphasis>
</taxonomicName>
collection as type because it is widely distributed.
</paragraph>
<paragraph pageId="42" pageNumber="43">
<emphasis bold="true" pageId="42" pageNumber="43">Specimen examined.</emphasis>
Mexico. 
<emphasis bold="true" pageId="42" pageNumber="43">Coahuila:</emphasis>
mountains 6 mi east of Saltillo, Feb to Oct 1880, E.Palmer 1366 (GH, NY, US-924987⚥). Sierra de Los Lirios, 2800 m, Jul to Aug 1942 [error for 1924?], E.Lyonnet 3691 (MEXU-379081⚥, 379082⚥). 22 km SE of San Antonio de las Alazanas, southeast of Saltillo, 20 Jul 1963, F.W.Gould 10538A &amp; D. Watson (TAES). 
<normalizedToken originalValue="Rincòn">Rincon</normalizedToken>
de 
<normalizedToken originalValue="María">Maria</normalizedToken>
, on Hacienda La Babia, ca. 70 road miles northwest from 
<normalizedToken originalValue="Múzquiz">Muzquiz</normalizedToken>
, 1750 m, 27 Apr 1975, T.Wendt 883 &amp; D.Riskind (LL). Cima de Sierra la Viga, ca. 3600 m, 24 Oct 1984, A.McDonald 1172 &amp; Gomez (TAES⚥). ceja y ladera s de Sierra La Viga, 3700 m, 22 Aug 1986, A.McDonald 2104 (TEX, TEX). Las Vigas, 
<normalizedToken originalValue="Cañón">Canon</normalizedToken>
de la Carbonera, Sierra de Arteaga, 2100-2600 m, 5 Jun 1987, J.A.Villarreal &amp; M.A.Carranza s.n. (MEXU-469611). Sierra La Marta, east of Cerro Moro, 3400 m, 22 Jul 1985, S.Ginzbarg 148, A.Whittemore &amp; A.McDonald (MEXU-666069♀). Sierra Coahuilon, ca. 3400-3500 m, 18 Jun 1985, A.McDonald 1485 &amp; M.H.Cervera (MEXU-1072101⚥). SE of San Antonio de las Alazanas, SE of Saltillo, at end of road near Summit of Coahuilon, 
<geoCoordinate degrees="25.2203" direction="north" orientation="latitude" precision="5" value="25.2203">25.2203°N</geoCoordinate>
, 
<geoCoordinate degrees="100.3305" direction="west" orientation="longitude" precision="5" value="-100.3305">100.3305°W</geoCoordinate>
, 3120 m, 17 Oct 1989, P.M.Peterson 8390, J.Valdes-Reyna, &amp; J.Villarreal (US-3518353⚥). 51.6 km southeast of Saltillo and 13 km southeast of Jame on road to Sierra La Viga, 
<geoCoordinate degrees="25.33" direction="north" orientation="latitude" precision="555" value="25.33">25.33°N</geoCoordinate>
, 
<geoCoordinate degrees="100.579" direction="west" orientation="longitude" precision="55" value="-100.579">100.579°W</geoCoordinate>
, 3240 m, 26 Sep 1990, P.M.Peterson 10053, C.R.Annable &amp; J.Valdes-Reyna (US). Sierra 
<normalizedToken originalValue="Zapalinamé">Zapaliname</normalizedToken>
, east of Saltillo, 
<geoCoordinate degrees="25.3468" direction="north" orientation="latitude" precision="5" value="25.3468">25.3468°N</geoCoordinate>
, 
<geoCoordinate degrees="100.9016" direction="west" orientation="longitude" precision="5" value="-100.9016">100.9016°W</geoCoordinate>
, 2700 m, 2 Sep 2005, P.M.Peterson 18787 &amp; J.Valdes-Reyna (US-3496160 (DNA voucher, unpublished, distributed as &quot; 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
&quot;); ditto, E of Saltillo, &quot;El Penitente&quot;, 
<geoCoordinate degrees="25.3468" direction="north" orientation="latitude" precision="5" value="25.3468">25.3468°N</geoCoordinate>
, 
<geoCoordinate degrees="100.9016" direction="west" orientation="longitude" precision="5" value="-100.9016">100.9016°W</geoCoordinate>
, 3070 m, 2 Sep 2005, P.M.Peterson 18790 &amp; J.Valdes-Reyna (US-3496160); ditto, along trail from El Cuatro to El Penitente, 
<geoCoordinate degrees="25.3383" direction="north" orientation="latitude" precision="5" value="25.3383">25.3383°N</geoCoordinate>
, 
<geoCoordinate degrees="100.8882" direction="west" orientation="longitude" precision="5" value="-100.8882">100.8882°W</geoCoordinate>
, 2925 m, 28 Sep 2007, P.M.Peterson 21127, J.M.Saarela, &amp; S.G. 
<normalizedToken originalValue="Gómez-Pérez">Gomez-Perez</normalizedToken>
(US; DNA voucher, unpublished). 
<emphasis bold="true" pageId="42" pageNumber="43">
Nuevo 
<normalizedToken originalValue="León">Leon</normalizedToken>
:
</emphasis>
Cerro 
<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
, east slope, 2135 m, 9 Jul 1963, R.L.McGregor 385, L.J.Harms, A.J.Robinson, R.delRosario, R.Segal (US-2454930⚥).
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="discussion">
<paragraph pageId="42" pageNumber="43">Discussion.</paragraph>
<paragraph pageId="42" pageNumber="43">
A provisional key to 11 species of 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="42" pageNumber="43">Poa</emphasis>
</taxonomicName>
in northern Mexico (excluding Baja California) is provided in 
<bibRefCitation author="Peterson, PM" journalOrPublisher="Sida" pageId="69" pageNumber="70" pagination="905 - 914" refId="B36" refString="Peterson, PM, Soreng, RJ, Herrera-Arrieta, Y, 2006. Poa matri-occidentalis (Poaceae: Pooideae: Poeae: Poinae), a new species from Mexico. Sida 22 (2): 905 - 914" title="Poa matri-occidentalis (Poaceae: Pooideae: Poeae: Poinae), a new species from Mexico." volume="22" year="2006">Peterson et al. (2006)</bibRefCitation>
. In that paper we concluded that 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
, as widely applied, was heterogeneous. However, until we could examine the type of 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="42" pageNumber="43">C. Heller 312</emphasis>
) we could not be sure of the application of the name. Here we emend the description of 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
Peyr. s.s., and treat the material of northeastern Mexico as a new species. 
<taxonomicName authorityName="Soreng &amp; P. M. Peterson" authorityYear="2012" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis italics="true" pageId="42" pageNumber="43">Poa palmeri</emphasis>
</taxonomicName>
has long been confused with 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
(
<bibRefCitation author="Fournier, E" journalOrPublisher="Euphytica" pageId="69" pageNumber="70" refId="B13" refString="Fournier, E, 1886. Mexicanas plantas. Pars secunda: Graminee, Ex Typographeo Reipublicae, Paris 2: 1-160." title="Mexicanas plantas. Pars secunda: Graminee, Ex Typographeo Reipublicae, Paris 2: 1 - 160." year="1886">Fournier 1886</bibRefCitation>
, 
<bibRefCitation author="Hitchcock, AS" journalOrPublisher="Australian Systematic Botany" pageId="69" pageNumber="70" refId="B25" refString="Hitchcock, AS, 1913. Mexican grasses in the United States National Herbarium. Contributions from the United States National Herbarium 17(3): 181-389." title="Mexican grasses in the United States National Herbarium. Contributions from the United States National Herbarium 17 (3): 181 - 389." year="1913">Hitchcock 1913</bibRefCitation>
, Beetle et al. 1999, 
<bibRefCitation author="Espejo Serna, A" journalOrPublisher="Euphytica" pageId="69" pageNumber="70" refId="B12" refString="Espejo Serna, A, Lopez-Ferrari, AR, Valdes-Reyna, J, 2000. Poaceae.In: A. Espejo Serna A, Lopez-Ferrari AR (Eds) Las Monocotiledoneas Mexicanas: una Sinopsis Floristica, Partes IX-XI. Consejo Nacional de la Flora de Mexico, A.C., Universidad Autonoma Metropolitana-Iztapalapa, and Comision Nacional para el Conocimiento y uso de la Biodiversidad, Mexico, D.F. 10: 8-236." title="Poaceae. In: A. Espejo Serna A, Lopez-Ferrari AR (Eds) Las Monocotiledoneas Mexicanas: una Sinopsis Floristica, Partes IX-XI. Consejo Nacional de la Flora de Mexico, A. C., Universidad Autonoma Metropolitana-Iztapalapa, and Comision Nacional para el Conocimiento y uso de la Biodiversidad, Mexico, D. F. 10: 8 - 236." year="2000">Espejo Serna et al. 2000</bibRefCitation>
, 
<bibRefCitation author="Davila Aranda, P" journalOrPublisher="Euphytica" pageId="69" pageNumber="70" refId="B11" refString="Davila Aranda, P, Mejia-Saules, MT, Gomez-Sanchez, M, Valdes-Reyna, J, Ortiz, JJ, Morin, C, Castrejon, J, Ocampo, A, 2006. Catalogo de las Gramineas de Mexico. i-xii, 1-671 + errata." title="Catalogo de las Gramineas de Mexico. i-xii, 1 - 671 + errata." year="2006">
<normalizedToken originalValue="Dávila">Davila</normalizedToken>
Aranda et al. 2006
</bibRefCitation>
). 
<taxonomicName authorityName="Soreng &amp; P. M. Peterson" authorityYear="2012" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis italics="true" pageId="42" pageNumber="43">Poa palmeri</emphasis>
</taxonomicName>
differs in several leaf, panicle, and spikelet characters, and has longer anthers, and the two species do not overlap in geographic range. There is a note on 
<normalizedToken originalValue="Palmer’s">Palmer's</normalizedToken>
label at US, &quot; 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
Peyr, so named at Kew, S.W. (RJS-presumably Sereno Watson), but scarcely distinct from 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis italics="true" pageId="42" pageNumber="43">Poa flexuosa</emphasis>
</taxonomicName>
Muhl.&quot; (= 
<taxonomicName authorityName="Muhl ex Elliott" authorityYear="1816" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="autumnalis">
<emphasis italics="true" pageId="42" pageNumber="43">Poa autumnalis</emphasis>
</taxonomicName>
Muhl. ex Elliott of lowlands from the central and southern Appalachians), and a further note; &quot;related to 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis italics="true" pageId="42" pageNumber="43">Poa flexuosa</emphasis>
</taxonomicName>
&quot; apparently written by A.S. Hitchcock. Interestingly, this name was also applied to the type of 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
at W (
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Poa ruprechtii Peyr. Photo of holotype collection (Heller 312)." httpUri="https://binary.pensoft.net/fig/10151" pageId="42" pageNumber="43">Fig. 16</figureCitation>
). This presumably stems from 
<normalizedToken originalValue="Peyritsch’s">Peyritsch's</normalizedToken>
final statement (p. 8) that his new species is like what is now called 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="cuspidata">
<emphasis italics="true" pageId="42" pageNumber="43">Poa cuspidata</emphasis>
</taxonomicName>
Nutt. (also of the Appalachians), which has sometimes been confused in herbaria with 
<taxonomicName authorityName="Muhl ex Elliott" authorityYear="1816" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="autumnalis">
<emphasis italics="true" pageId="42" pageNumber="43">Poa autumnalis</emphasis>
</taxonomicName>
. Peyritsch stated, &quot;Scheint mit 
<taxonomicName authorityName="Schult" authorityYear="1824" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="brachyphylla">
<emphasis italics="true" pageId="42" pageNumber="43">Poa brachyphylla</emphasis>
</taxonomicName>
Schult. (
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="brevifolia">
<emphasis italics="true" pageId="42" pageNumber="43">Poa brevifolia</emphasis>
</taxonomicName>
Muhl.) verwant zu sein.&quot; The latter two names are synonyms of 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="cuspidata">
<emphasis italics="true" pageId="42" pageNumber="43">Poa cuspidata</emphasis>
</taxonomicName>
Nutt.
</paragraph>
<paragraph pageId="42" pageNumber="43">
<normalizedToken originalValue="Pringle’s">Pringle's</normalizedToken>
label on the type collection can be read as 3000 ft, but we wonder if this is a printing error as the 3 looks more like an 8 when inspected closely, without the serifs of the 3 that are present in the date and other numbers. Whatever the case is, 3000 ft seems far too low for this species.
</paragraph>
<paragraph pageId="42" pageNumber="43">
The breeding system of 
<taxonomicName authorityName="Soreng &amp; P. M. Peterson" authorityYear="2012" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis italics="true" pageId="42" pageNumber="43">Poa palmeri</emphasis>
</taxonomicName>
is not clear. We tentatively identify it as trioecious because a few specimens are staminate throughout, a few are pistillate, most are hermaphroditic, but some of the later have late aborted stamens. Possibly sex-expression varies within some individuals between late and early season panicles, as seen in sequential gynomonoecism (
<bibRefCitation author="Soreng, RJ" journalOrPublisher="Madrono" pageId="69" pageNumber="70" pagination="300 - 306" refId="B59" refString="Soreng, RJ, Keil, DJ, 2003. Sequentially adjusted sex-ratios in gynomonoecism, and Poa diaboli (Poaceae), a new species from California. Madrono 50 (4): 300 - 306" title="Sequentially adjusted sex-ratios in gynomonoecism, and Poa diaboli (Poaceae), a new species from California." volume="50" year="2003">Soreng and Keil 2003</bibRefCitation>
).
</paragraph>
<paragraph pageId="42" pageNumber="43">
It is most easily differentiated from 
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="strictiramea">
<emphasis italics="true" pageId="42" pageNumber="43">Poa strictiramea</emphasis>
</taxonomicName>
by having smooth foliage, more developed lemma and callus pubescence, and smoother lemma surfaces.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10150" pageId="42" pageNumber="43" start="Figure 15" startId="F15">
<paragraph pageId="42" pageNumber="43">
<emphasis bold="true" pageId="42" pageNumber="43">Figure 15.</emphasis>
<taxonomicName authorityName="Soreng &amp; P. M. Peterson" authorityYear="2012" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="palmeri">
<emphasis italics="true" pageId="42" pageNumber="43">Poa palmeri</emphasis>
</taxonomicName>
Soreng &amp; P.M. Peterson. 
<emphasis bold="true" pageId="42" pageNumber="43">A</emphasis>
habit 
<emphasis bold="true" pageId="42" pageNumber="43">B</emphasis>
inflorescence 
<emphasis bold="true" pageId="42" pageNumber="43">C</emphasis>
sheath, ligule, blade 
<emphasis bold="true" pageId="42" pageNumber="43">D</emphasis>
spikelet 
<emphasis bold="true" pageId="42" pageNumber="43">E</emphasis>
glumes 
<emphasis bold="true" pageId="42" pageNumber="43">F</emphasis>
floret 
<emphasis bold="true" pageId="42" pageNumber="43">G</emphasis>
callus of floret 
<emphasis bold="true" pageId="42" pageNumber="43">H</emphasis>
lemma dorsal view 
<emphasis bold="true" pageId="42" pageNumber="43">I</emphasis>
palea dorsal and ventral views 
<emphasis bold="true" pageId="42" pageNumber="43">J</emphasis>
stamen 
<emphasis bold="true" pageId="42" pageNumber="43">K</emphasis>
pistil from perfect-flowered plant, with lodicules attached to palea 
<emphasis bold="true" pageId="42" pageNumber="43">L</emphasis>
rudimentary pistil from staminate plant, with lodicules. 
<emphasis bold="true" pageId="42" pageNumber="43">A-J</emphasis>
and 
<emphasis bold="true" pageId="42" pageNumber="43">L</emphasis>
drawn from holotype collection (
<emphasis italics="true" pageId="42" pageNumber="43">Pringle 10212</emphasis>
) 
<emphasis bold="true" pageId="42" pageNumber="43">K</emphasis>
drawn from 
<emphasis italics="true" pageId="42" pageNumber="43">Palmer 1366</emphasis>
.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/10151" pageId="42" pageNumber="43" start="Figure 16" startId="F16">
<paragraph pageId="42" pageNumber="43">
<emphasis bold="true" pageId="42" pageNumber="43">Figure 16.</emphasis>
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ruprechtii">
<emphasis italics="true" pageId="42" pageNumber="43">Poa ruprechtii</emphasis>
</taxonomicName>
Peyr. Photo of holotype collection (
<emphasis italics="true" pageId="42" pageNumber="43">Heller 312</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>