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<document id="D2DB16344B696CA8E53659DEDF19AE4C" ID-DOI="10.5281/zenodo.12520892" ID-ISSN="1407-8619" ID-Zenodo-Dep="12520892" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719251696330" checkinUser="felipe" docAuthor="Shavrin, Alexey V." docDate="2020" docId="423DE34C9A061849FF321B24520DFBD7" docLanguage="en" docName="BalticJColeopterol.20.1.29-33.pdf" docOrigin="Baltic Journal of Coleopterology 20 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.3:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="3" docTitle="Anthobium golovatchi Shavrin, 2020, sp.n." docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="BE049B349A07184AFFD11A26516CFFC0" masterDocTitle="A new species of the genus Anthobium Leach, 1819 from Cuba (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="33" masterPageNumber="29" pageNumber="29" updateTime="1719253310998" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="15D61CECD9A13B29EB78DF5E6AD17CD9">A new species of the genus Anthobium Leach, 1819 from Cuba (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="8488E1028EBAB2327DB44362F7683D74">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="0189C74F228682ABC93C5CD10D6862CE">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="2C318F439364799FB04BBFF4E822E462">2020</mods:date>
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<treatment id="423DE34C9A061849FF321B24520DFBD7" LSID="urn:lsid:plazi:treatment:423DE34C9A061849FF321B24520DFBD7" httpUri="http://treatment.plazi.org/id/423DE34C9A061849FF321B24520DFBD7" lastPageId="3" pageId="1">
<subSubSection id="828E01D19A06184BFF321B245376FEDA" box="[227,538,258,282]" pageId="1" type="nomenclature">
<paragraph id="CA2B525A9A06184BFF321B245376FEDA" blockId="1.[227,538,258,314]" box="[227,538,258,282]" pageId="1">
<heading id="9163E5369A06184BFF321B245376FEDA" bold="true" box="[227,538,258,282]" fontSize="10" level="2" pageId="1" reason="4">
<emphasis id="F8E08E489A06184BFF321B245376FEDA" bold="true" box="[227,538,258,282]" pageId="1">
<taxonomicName id="0D9429D99A06184BFF321B2450B6FEDA" ID-CoL="8XTT6" authorityName="Shavrin" authorityYear="2020" box="[227,474,258,282]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="F8E08E489A06184BFF321B2450B6FEDA" bold="true" box="[227,474,258,282]" italics="true" pageId="1">Anthobium golovatchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3D333339A06184BFE341B245376FEDA" box="[485,538,258,282]" pageId="1" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="828E01D19A06184BFF321B045028FEFA" box="[227,324,290,314]" pageId="1" type="description">
<paragraph id="CA2B525A9A06184BFF321B045028FEFA" blockId="1.[227,538,258,314]" box="[227,324,290,314]" pageId="1">
<figureCitation id="52AF4EDF9A06184BFF321B045028FEFA" box="[227,324,290,314]" captionStart="Fig" captionStartId="2.[303,343,1521,1545]" captionText="Fig. 16. Anthobium golovatchi: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26)." pageId="1">Figs. 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="828E01D19A06184BFF321B445055FDDA" pageId="1" type="materials_examined">
<paragraph id="CA2B525A9A06184BFF321B445055FDDA" blockId="1.[227,737,354,538]" pageId="1">
<emphasis id="F8E08E489A06184BFF321B445012FEBA" bold="true" box="[227,382,354,378]" pageId="1">Type material.</emphasis>
<typeStatus id="152FECF89A06184BFE551B44508EFEBA" box="[388,482,354,378]" pageId="1" type="holotype">Holotype</typeStatus>
, male:
<collectingCountry id="B28312CA9A06184BFDE11B445305FEBA" box="[560,617,354,378]" name="Cuba" pageId="1">Cuba</collectingCountry>
: Prov. Pinar del | Rio, Parque Nac. “La | Gūira”, oligodominant | tropical rainforest, | sifted litter,
<date id="BE2A749A9A06184BFDE01B8453C7FE7A" box="[561,683,418,442]" pageId="1" value="1981-06-12">06.12.1981</date>
. | S.I. Golovatch leg.,
<typeStatus id="152FECF89A06184BFE761BE45357FE1A" box="[423,571,450,474]" pageId="1" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="0D9429D99A06184BFD871BE453A3FE1A" authorityName="Leach" authorityYear="1819" box="[598,719,450,474]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="F8E08E489A06184BFD871BE453A3FE1A" box="[598,719,450,474]" italics="true" pageId="1">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="0D9429D99A06184BFF321BC45039FE3A" authorityName="Shavrin" authorityYear="2020" box="[227,341,482,506]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
<emphasis id="F8E08E489A06184BFF321BC45039FE3A" box="[227,341,482,506]" italics="true" pageId="1">golovatchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3D333339A06184BFE8B1BC550E2FE3A" box="[346,398,483,506]" pageId="1" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.
<collectionCode id="AC85CA9F9A06184BFDC11BC55349FE3B" box="[528,549,483,507]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" type="Museum">V</collectionCode>
. des. 2020 &lt;red&gt; (ZMM).
</paragraph>
</subSubSection>
<subSubSection id="828E01D19A061849FF3218645219FDB7" lastPageId="3" pageId="1" type="description">
<paragraph id="CA2B525A9A06184BFF3218645039FBFA" blockId="1.[227,737,578,1082]" pageId="1">
<emphasis id="F8E08E489A06184BFF321864501CFD9A" bold="true" box="[227,368,578,602]" pageId="1">Description.</emphasis>
Measurements of the
<typeStatus id="152FECF89A06184BFD8E186453D0FD9A" box="[607,700,578,602]" pageId="1" type="holotype">holotype</typeStatus>
(in mm): maximum width of head including eyes: 0.67; length of head (from base of labrum to neck constriction along head midline): 0.42; length of antenna: 1.20; ocular length (longitudinal): 0.15; length of pronotum: 0.62; maximum width of pronotum: 1.15; sutural length of elytra (length of elytra from the apex of scutellum to the posterior margin of sutural angle): 1.25; maximum width of elytra: 1.32; maximum width of abdomen: 1.05; length of metatibia: 0.67; length of metatarsus: 0.29 (metatarsomeres 14: 0.17; metatarsomere 5: 0.12); length of aedeagus (from base of the median lobe to apex of parameres): 0.82; total length (from anterior margin of clypeus to apex of abdomen): 3.37.
</paragraph>
<paragraph id="CA2B525A9A06184BFF321E445598FCD7" blockId="1.[227,737,1122,1723]" lastBlockId="1.[774,1286,191,791]" pageId="1">
Body, antennomeres 411 and legs yelow-brown (head and middle portion of pronotum slightly darker); mouthparts, antennomeres 13 and tarsi yellow. Body shiny; pronotum, scutellum and elytra without microsculpture; apical portion of head with distinct, regular, transverse meshes, middle portion with indistinct, irregular and diagonal microreticulation, anterio-lateral portions of head between antennal insertion and apical margin of eye with dense, diagonal and rugose sculpture, median and posterior portions of infraorbital ridges with indistinct transverse meshes; neck with indistinct, isodiametric sculpture; abdomen with very dense isodiametric microsculpture. Head with irregular, sparse and fine punctation, finer in anterior portion, denser and deeper in middle and on infraorbital ridges, with two diagonal impunctated portions between grooves and median part; anterior portion of neck with very dense and deep punctation, slightly larger on posterior portions of infraorbital ridges, basal portion of head with very sparse, fine punctation; pronotum with dense, moderately large and deep punctation, finer in middle, sparser in mediobasal, larger and deeper in lateral portions; scutellum with several fine and sparsely distributed punctures; punctation of elytra as that on pronotum, finer in parascutellar area, sparser in middle, larger and deeper in mediobasal and apical portions; middle portions of each elytron forming seven indistinct rows of vague and tangled punctures; abdominal tergites without visible punctures. Body glabrous, apical portion of head with several long tactile setae; surface of clypeus with very sparse, short setae; abdominal tergites with regular, short and fine setae, and with additional longer and sparsely distributed erect setation. Habitus as in
<figureCitation id="52AF4EDF9A06184BFB7E18D95582FCD7" box="[1199,1262,767,791]" captionStart="Fig" captionStartId="2.[303,343,1521,1545]" captionText="Fig. 16. Anthobium golovatchi: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26)." pageId="1">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="CA2B525A9A06184BFCD719195468F977" blockId="1.[774,1286,831,1719]" pageId="1">Head 1.5 times as wide as long; middle portion slightly elevated, with deep and long grooves in front of ocelli, reaching middle level of eyes; postocular ridge acute, with surface between it and posterior margin of eyes as long as two nearest ommatidia; anteriobasal portions of clypeus with distinct, diagonal impressions; basal portion between ocelli without transverse impression; anterior portion between antennal insertion and anterior margin of eye with very deep, semicircular notch. Eyes moderately small, convex. Ocelli large, situated at level slightly behind postocular ridges; distance between ocelli about as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palpus about two and half times as long as preceding segment, slightly widened in middle and gradually narrowing toward acute apex; preapical segment narrow, one and half times as long as wide. Antenna exceeding apical third of elytra when reclined, antennomeres 411 covered by dense pubescence; basal antennomere distinctly more than twice as long as wide, antennomere 2 suboval, distinctly narrower than basal antennomere, about twice as wide as long, 3 distinctly narrower and longer than 2, 47 about as long as 3, but distinctly wider, 8 distinctly shorter than 7, 9 slightly shorter than 8, 10 transverse,</paragraph>
<caption id="9EEB02D29A051848FEFE1FD753EFF989" pageId="2" startId="2.[303,343,1521,1545]" targetType="figure">
<paragraph id="CA2B525A9A051848FEFE1FD753EFF989" blockId="2.[303,1362,1521,1609]" pageId="2">
Fig. 16.
<taxonomicName id="0D9429D99A051848FE491FD753E7F9C9" authorityName="Shavrin" authorityYear="2020" box="[408,651,1521,1545]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="F8E08E489A051848FE491FD753E7F9C9" box="[408,651,1521,1545]" italics="true" pageId="2">Anthobium golovatchi</emphasis>
</taxonomicName>
: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26).
</paragraph>
</caption>
<paragraph id="CA2B525A9A041849FD671AB7525EFF64" blockId="3.[694,818,145,164]" box="[694,818,145,164]" pageId="3">
<emphasis id="F8E08E489A041849FD671AB7525EFF64" box="[694,818,145,164]" italics="true" pageId="3">Shavrin A. V.</emphasis>
</paragraph>
<paragraph id="CA2B525A9A041849FF321A9950B1FED7" blockId="3.[227,738,191,279]" pageId="3">about as long as wide, apical antennomere about 1.3 times as long as 10, from apical third sharply narrowing toward apex.</paragraph>
<paragraph id="CA2B525A9A041849FF321B665343FD77" blockId="3.[227,737,319,695]" pageId="3">Pronotum transverse, 1.8 times as wide as long, 1.7 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad, with distinctly concaved laterobasal margins in front of acute posterior angles; apical angles widely rounded, slightly protruded anteriad; apical margin somewhat straight, markedly narrower than posterior margin; middle portion elevated, with very indistinct mediobasal transverse impression; lateral margins without crenulation; lateral portions moderately wide, each with deep, small pit slightly above middle.</paragraph>
<paragraph id="CA2B525A9A041849FF3218F9530FFC17" blockId="3.[227,737,735,983]" pageId="3">Elytra very convex, slightly wider than long, twice as long as pronotum, slightly widened toward apical third, reaching basal margin of abdominal tergite V; apical margin somewhat straight; surface of elytra without elevations; lateral portions moderately narrow, indistinctly explanate; lateroapical margins with indistinct, smooth crenulation. Wings fully developed.</paragraph>
<paragraph id="CA2B525A9A041849FF321E2653A1FB77" blockId="3.[227,738,1023,1207]" pageId="3">Legs moderately long, slender; tibiae covered by sparse, sclerotized and elongate setae, mixed with very dense, long pubescence; apical margins of each tibia with long, sclerotized spines; mesotibiae indistinctly narrowed in middle; metatarsus more than twice shorter than metatibia.</paragraph>
<paragraph id="CA2B525A9A041849FF321EF953C9FA97" blockId="3.[227,737,1247,1367]" pageId="3">Abdomen distinctly narrower than elytra, convex, without visible tomentose spots in middle of abdominal tergite V; apical margin of abdominal tergite VII with narrow palisade fringe.</paragraph>
<paragraph id="CA2B525A9A041849FF321F5952B5FF37" blockId="3.[227,737,1407,1719]" lastBlockId="3.[774,1284,191,247]" pageId="3">
Male. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII slightly concaved. Aedeagus with wide basal portion, gradually narrowing toward truncated apex (
<figureCitation id="52AF4EDF9A041849FEF11FD950EDF9D7" box="[288,385,1535,1559]" captionStart="Fig" captionStartId="2.[303,343,1521,1545]" captionText="Fig. 16. Anthobium golovatchi: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26)." pageId="3">Figs. 2, 4</figureCitation>
); parameres moderately wide, distinctly exceeding apex of median lobe, slightly widened apically, with two long apical and two short preapical setae (
<figureCitation id="52AF4EDF9A041849FE1F1C79537DF9B7" box="[462,529,1631,1655]" captionStart="Fig" captionStartId="2.[303,343,1521,1545]" captionText="Fig. 16. Anthobium golovatchi: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26)." pageId="3">Fig. 4</figureCitation>
); internal sac complicated, long and moderately wide, with field of long sclerotized spines in middle and with large spines in apical portions (
<figureCitation id="52AF4EDF9A041849FBCF1A9955E4FF17" box="[1054,1160,191,215]" captionStart="Fig" captionStartId="2.[303,343,1521,1545]" captionText="Fig. 16. Anthobium golovatchi: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26)." pageId="3">Figs. 56</figureCitation>
). Aedeagus laterally as in
<figureCitation id="52AF4EDF9A041849FC461AF952BAFF37" box="[919,982,223,247]" captionStart="Fig" captionStartId="2.[303,343,1521,1545]" captionText="Fig. 16. Anthobium golovatchi: 1habitus, 2, 4aedeagus (parameral view), 3aedeagus (lateral view), 56apical portion of median lobe (Fig. 5: parameral view, Fig. 6: lateral view). Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 26)." pageId="3">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="CA2B525A9A041849FCD71B3952AFFEF7" blockId="3.[774,1285,287,631]" box="[774,963,287,311]" pageId="3">Female unknown.</paragraph>
<paragraph id="CA2B525A9A041849FCD71B195219FDB7" blockId="3.[774,1285,287,631]" pageId="3">
<emphasis id="F8E08E489A041849FCD71B1952B1FE97" bold="true" box="[774,989,319,343]" pageId="3">Comparative notes.</emphasis>
Regarding the general shape of the forebody and proportions of antennomeres,
<taxonomicName id="0D9429D99A041849FC7C1BA65553FE57" authorityName="Shavrin" authorityYear="2020" box="[941,1087,383,407]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="golovatchi" status="sp.n.">
<emphasis id="F8E08E489A041849FC7C1BA652D1FE57" box="[941,957,384,407]" italics="true" pageId="3">A</emphasis>
.
<emphasis id="F8E08E489A041849FC1A1B595553FE57" box="[971,1087,383,407]" italics="true" pageId="3">golovatchi</emphasis>
</taxonomicName>
<emphasis id="F8E08E489A041849FB971B595510FE57" bold="true" box="[1094,1148,383,407]" pageId="3">
<taxonomicNameLabel id="E3D333339A041849FB971B595510FE57" box="[1094,1148,383,407]" pageId="3" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to Mexican
<taxonomicName id="0D9429D99A041849FCB71B86551FFE77" authority="Thayer, 2003" authorityName="Thayer" authorityYear="2003" box="[870,1139,415,439]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="amicorum">
<emphasis id="F8E08E489A041849FCB71B86521AFE77" box="[870,886,416,439]" italics="true" pageId="3">A</emphasis>
.
<emphasis id="F8E08E489A041849FC501B865286FE77" box="[897,1002,416,439]" italics="true" pageId="3">amicorum</emphasis>
<bibRefCitation id="AE052FAB9A041849FC3C1BB9551FFE77" author="Thayer M. K." box="[1005,1139,415,439]" pageId="3" pagination="311 - 358" refId="ref2167" refString="Thayer M. K. 2003. Omaliinae of Mexico: New species, combinations, and records (Coleoptera: Staphylinidae). In: Cuccodoro G., Leschen R. A. B. (eds.), Systematics of Coleoptera: Papers Celebrating the Retirement of Ivan Lobl, 17. Memoirs on Entomology, International, pp. 311 - 358." type="book chapter" year="2003">Thayer, 2003</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0D9429D99A041849FB701B8652FAFE17" authority="Thayer, 2003" authorityName="Thayer" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="tapatio">
<emphasis id="F8E08E489A041849FB701B8655DDFE77" box="[1185,1201,416,439]" italics="true" pageId="3">A</emphasis>
.
<emphasis id="F8E08E489A041849FB6D1B865469FE77" box="[1212,1285,416,439]" italics="true" pageId="3">tapatio</emphasis>
<bibRefCitation id="AE052FAB9A041849FCD71B9952FAFE17" author="Thayer M. K." box="[774,918,447,471]" pageId="3" pagination="311 - 358" refId="ref2167" refString="Thayer M. K. 2003. Omaliinae of Mexico: New species, combinations, and records (Coleoptera: Staphylinidae). In: Cuccodoro G., Leschen R. A. B. (eds.), Systematics of Coleoptera: Papers Celebrating the Retirement of Ivan Lobl, 17. Memoirs on Entomology, International, pp. 311 - 358." type="book chapter" year="2003">Thayer, 2003</bibRefCitation>
</taxonomicName>
, from which it differs by the larger body, paler coloration, by the shape of laterobasal portions of the pronotum (concaved laterobasal margin in front of acute posterior angles), and different external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="828E01D19A041849FCD718B95571FCB7" pageId="3" type="distribution">
<paragraph id="CA2B525A9A041849FCD718B952CBFCD7" blockId="3.[774,1285,671,791]" pageId="3">
<emphasis id="F8E08E489A041849FCD718B952F6FD77" bold="true" box="[774,922,671,695]" pageId="3">Distribution.</emphasis>
The species is known only from the
<typeStatus id="152FECF89A041849FCFA18E65239FD18" box="[811,853,704,728]" pageId="3">type</typeStatus>
locality in Parque Nacional La Güira (=La Güira National Park) in
<collectingRegion id="08509CB89A041849FBDA18F9546CFD37" box="[1035,1280,735,759]" country="Cuba" name="Pinar del Rio" pageId="3">Pinar del Rio Province</collectingRegion>
, Western
<collectingCountry id="B28312CA9A041849FCB718D952CEFCD7" box="[870,930,767,791]" name="Cuba" pageId="3">Cuba</collectingCountry>
.
</paragraph>
<paragraph id="CA2B525A9A041849FCD719195571FCB7" blockId="3.[774,1285,831,887]" pageId="3">
<emphasis id="F8E08E489A041849FCD7191952E9FC97" bold="true" box="[774,901,831,855]" pageId="3">Bionomics.</emphasis>
The specimen were taken by sifting litter in tropical rainforest.
</paragraph>
</subSubSection>
<subSubSection id="828E01D19A041849FCD719B9520DFBD7" pageId="3" type="etymology">
<paragraph id="CA2B525A9A041849FCD719B9520DFBD7" blockId="3.[774,1285,927,1047]" pageId="3">
<emphasis id="F8E08E489A041849FCD719B952E4FC77" bold="true" box="[774,904,927,951]" pageId="3">Etymology.</emphasis>
Eponymic, the species is named to honour the Russian entomologist Sergei I. Golovatch (
<collectingRegion id="08509CB89A041849FC5619F95284FC37" box="[903,1000,991,1015]" country="Russia" name="Moscow" pageId="3">Moscow</collectingRegion>
), the collector of the
<typeStatus id="152FECF89A041849FB0619C65469FC38" box="[1239,1285,992,1016]" pageId="3">type</typeStatus>
material.
</paragraph>
</subSubSection>
</treatment>
</document>