treatments-xml/data/42/32/87/423287F433325C20C7E1FC17AEC8E4AA.xml
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<mods:title id="548650B4DC73E0BF7212530B02B4F497">A revision of Palaearctic and Oriental Scymbalium and Micrillus III. New species, new combinations, and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<heading id="916C818E33325C21C7E1FC17AE0FE481" box="[178,584,908,931]" fontSize="8" level="2" pageId="8" pageNumber="985" reason="2">
<emphasis id="F8EFEAF033325C21C7E1FC17AD83E480" bold="true" box="[178,452,908,930]" pageId="8" pageNumber="985">
<taxonomicName id="0D9B4D6133325C21C7E1FC17AD36E480" ID-CoL="9V8WP" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[178,369,908,930]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="985" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="F8EFEAF033325C21C7E1FC17AD36E480" bold="true" box="[178,369,908,930]" italics="true" pageId="8" pageNumber="985">Micrillus hamatus</emphasis>
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</emphasis>
(
<figureCitation id="52A02A6733325C21C682FC16AE06E481" box="[465,577,909,931]" captionStart="Figs 18-24" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[186,1086,213,1490]" captionTargetId="figure@11.[179,1086,206,1491]" captionText="Figs 18-24: Micrillus virgatus (18-19) and M. hamatus (20-24): (18, 21) male sternite VIII; (19) basal portion of aedeagus in lateral view; (20) habitus; (22-23) aedeagus in lateral and in ventral view; (24) apex of ventral process in lateral view. Scale bars: 20: 1.0 mm; 18, 21-23: 0.5 mm; 19, 24: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776478" httpUri="https://zenodo.org/record/3776478/files/figure.png" pageId="8" pageNumber="985">Figs 20-24</figureCitation>
)
</heading>
</paragraph>
<paragraph id="CA2436E233325C21C7E1FC29ACB3E4D7" blockId="8.[178,1085,945,1013]" pageId="8" pageNumber="985">
<materialsCitation id="7AF33CBF33325C21C7E1FC29ACB3E4D7" ID-GBIF-Occurrence="2609445632" collectingDate="2007-06-03" collectorName="Lgt. F. Houska" country="Morocco" determinerName="V. Assing" location="MOROCCO" pageId="8" pageNumber="985" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
T y p e m a t e r i a l:
<typeStatus id="1520884033325C21C6FBFC2AADBAE4E5" box="[424,509,945,967]" pageId="8" pageNumber="985" type="holotype">Holotype</typeStatus>
<specimenCount id="DC9DFD6B33325C21C557FC29AE5FE4E5" box="[516,536,946,967]" pageId="8" pageNumber="985" type="male"></specimenCount>
: &quot;
<collectingCountry id="B28C767233325C21C579FC29AED8E4E5" box="[554,671,946,967]" name="Morocco" pageId="8" pageNumber="985">MOROCCO</collectingCountry>
SE,
<date id="BE25102233325C21C59CFC2AAF67E4E5" box="[719,800,945,967]" pageId="8" pageNumber="985" value="2007-06-03">
<collectingDate id="AE61E9CA33325C21C59CFC2AAF67E4E5" box="[719,800,945,967]" pageId="8" pageNumber="985" value="2007-06-03">3.6.2007</collectingDate>
</date>
, Zagora
<quantity id="0D639B0733325C21C420FC29AFE8E4E5" box="[883,943,945,967]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="985" unit="km" value="30.0">30 km</quantity>
SE, Zaouia-el- Barrahnia, Drâa riv.,
<collectorName id="676E533433325C21C625FC52AE47E4FC" box="[374,512,968,990]" pageId="8" pageNumber="985">Lgt. F. Houška</collectorName>
/
<typeStatus id="1520884033325C21C547FC53AE34E4FC" box="[532,627,968,990]" pageId="8" pageNumber="985" type="holotype">Holotypus</typeStatus>
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<taxonomicName id="0D9B4D6133325C21C5C2FC53AF70E4FF" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[657,823,968,989]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="985" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="F8EFEAF033325C21C5C2FC53AF70E4FF" box="[657,823,968,989]" italics="true" pageId="8" pageNumber="985">Micrillus hamatus</emphasis>
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<taxonomicNameLabel id="E3DC578B33325C21C46DFC52AF36E4FC" box="[830,881,969,990]" pageId="8" pageNumber="985" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="3FF8DA2633325C21C4F0FC52AFB8E4FC" box="[931,1023,968,990]" pageId="8" pageNumber="985">V. Assing</determinerName>
2018&quot; (cAss).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8281656933325C20C7E1FB98AEC8E4AA" lastPageId="9" lastPageNumber="986" pageId="8" pageNumber="985" type="description">
<paragraph id="CA2436E233325C21C7E1FB98AE0EE314" blockId="8.[178,1084,1027,1078]" pageId="8" pageNumber="985">E t y m o l o g y: The specific epithet (Latin, adjective: hooked) alludes to the shape of the apex of the aedeagus in lateral view.</paragraph>
<paragraph id="CA2436E233325C21C7E1FBDDAD03E391" blockId="8.[178,1085,1093,1203]" pageId="8" pageNumber="985">
D e s c r i p t i o n: Body length 7.0 mm; length of forebody
<quantity id="0D639B0733325C21C402FBDEAFE2E379" box="[849,933,1093,1115]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="985" unit="mm" value="3.6">3.6 mm</quantity>
. Habitus as in
<figureCitation id="52A02A6733325C21C7E1FBF8ACBBE35A" box="[178,252,1123,1145]" captionStart="Figs 18-24" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[186,1086,213,1490]" captionTargetId="figure@11.[179,1086,206,1491]" captionText="Figs 18-24: Micrillus virgatus (18-19) and M. hamatus (20-24): (18, 21) male sternite VIII; (19) basal portion of aedeagus in lateral view; (20) habitus; (22-23) aedeagus in lateral and in ventral view; (24) apex of ventral process in lateral view. Scale bars: 20: 1.0 mm; 18, 21-23: 0.5 mm; 19, 24: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776478" httpUri="https://zenodo.org/record/3776478/files/figure.png" pageId="8" pageNumber="985">Fig. 20</figureCitation>
. Coloration: head blackish-brown with the anterior margin of the frons yellowishbrown; pronotum, elytra, and abdomen yellowish-red; legs yellowish; antennae yellowish-red.
</paragraph>
<paragraph id="CA2436E233325C21C7E1FB58AE4CE249" blockId="8.[178,1085,1219,1387]" pageId="8" pageNumber="985">Head 1.1 times as long as broad; punctation fine and moderately dense; median dorsal portion extensively impunctate; integument with extremely fine transverse microsculpture visible only at high magnification (100 x). Eyes weakly convex, not protruding from lateral contours of head, slightly shorter than postocular portion in dorsal view. Antennae slender; antennomere IV approximately twice as long as broad; antennomere X approximately 1.5 times as long as broad.</paragraph>
<paragraph id="CA2436E233325C21C7E1FAE0AFBDE2E9" blockId="8.[178,1085,1402,1483]" pageId="8" pageNumber="985">Pronotum 1.25 times as long as broad and as broad as head, of trapezoid shape, broadest at anterior angles and tapering posteriad; posterior margin weakly concave; punctation fine and moderately dense; midline impunctate; interstices without microsculpture.</paragraph>
<paragraph id="CA2436E233325C21C7E1FA41AEC5E108" blockId="8.[178,1085,1498,1579]" pageId="8" pageNumber="985">Elytra nearly as long as pronotum; punctation very dense and very fine. Hind wings present. Protarsomeres I-IV undilated; metatarsomere I very long, slightly longer than the combined length of metatarsomeres II-IV.</paragraph>
<paragraph id="CA2436E233335C20C7E1FF40AE49E62C" blockId="9.[178,1084,219,270]" pageId="9" pageNumber="986">Abdomen approximately as broad as elytra; punctation dense and fine; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="CA2436E233335C20C7E1FE85AFDEE6A9" blockId="9.[178,1085,285,395]" pageId="9" pageNumber="986">
<specimenCount id="DC9DFD6B33335C20C7E1FE85AC8FE611" box="[178,200,286,307]" pageId="9" pageNumber="986" type="male"></specimenCount>
: sternite VIII (
<figureCitation id="52A02A6733335C20C63AFE86ADF1E611" box="[361,438,285,307]" captionStart="Figs 18-24" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[186,1086,213,1490]" captionTargetId="figure@11.[179,1086,206,1491]" captionText="Figs 18-24: Micrillus virgatus (18-19) and M. hamatus (20-24): (18, 21) male sternite VIII; (19) basal portion of aedeagus in lateral view; (20) habitus; (22-23) aedeagus in lateral and in ventral view; (24) apex of ventral process in lateral view. Scale bars: 20: 1.0 mm; 18, 21-23: 0.5 mm; 19, 24: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776478" httpUri="https://zenodo.org/record/3776478/files/figure.png" pageId="9" pageNumber="986">Fig. 21</figureCitation>
) distinctly oblong, its posterior excision broadly and weakly concave; posterolateral processes of segment IX (valvulae) long and slender; aedeagus (
<figureCitation id="52A02A6733335C20C7EAFEC3AD6BE64C" box="[185,300,344,366]" captionStart="Figs 18-24" captionStartId="11.[178,218,1506,1528]" captionTargetBox="[186,1086,213,1490]" captionTargetId="figure@11.[179,1086,206,1491]" captionText="Figs 18-24: Micrillus virgatus (18-19) and M. hamatus (20-24): (18, 21) male sternite VIII; (19) basal portion of aedeagus in lateral view; (20) habitus; (22-23) aedeagus in lateral and in ventral view; (24) apex of ventral process in lateral view. Scale bars: 20: 1.0 mm; 18, 21-23: 0.5 mm; 19, 24: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3776478" httpUri="https://zenodo.org/record/3776478/files/figure.png" pageId="9" pageNumber="986">Figs 22-24</figureCitation>
)
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long, symmetric, and weakly sclerotized; ventral process hookshaped in lateral view; internal sac with long dark membranous structure.
</paragraph>
<paragraph id="CA2436E233335C20C7E1FE00AE5EE561" blockId="9.[178,1085,410,579]" pageId="9" pageNumber="986">
C o m p a r a t i v e n o t e s: Based on the long and slender valvulae IX and on the male primary and secondary characters,
<taxonomicName id="0D9B4D6133335C20C50BFE22AE94E6EF" authorityName="Assing" authorityYear="2018" box="[600,723,440,462]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="F8EFEAF033335C20C50BFE22AE94E6EF" box="[600,723,440,462]" italics="true" pageId="9" pageNumber="986">M. hamatus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="0D9B4D6133335C20C42BFE22AFBEE6EC" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[888,1017,441,462]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="F8EFEAF033335C20C42BFE22AFBEE6EC" box="[888,1017,441,462]" italics="true" pageId="9" pageNumber="986">M. testaceus</emphasis>
</taxonomicName>
group (see
<bibRefCitation id="AE0A4B1333335C20C7B7FE4EAD37E6C8" author="ASSING V" box="[228,368,469,490]" journalOrPublisher="Neue" pageId="9" pageNumber="986" pagination="301 - 344" part="1" refId="ref9555" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stu ttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." title="A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stu ttgarter Beitrage zur Naturkunde" type="journal article" year="2008">ASSING 2008</bibRefCitation>
,
<bibRefCitation id="AE0A4B1333335C20C62DFE4EADF2E6C9" author="ASSING V" box="[382,437,469,491]" journalOrPublisher="Linzer Biologische Beitrage" pageId="9" pageNumber="986" pagination="1479 - 1520" part="45" refId="ref9612" refString="ASSING V. (2013): On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 45 (2): 1479 - 1520." title="On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)" type="journal article" year="2013">2013</bibRefCitation>
). It is distinguished from all the species of this group by the conspicuously hook-shaped apex and the strongly sclerotized internal structures of the aedeagus. It is additionally distinguished from the three
<taxonomicName id="0D9B4D6133335C20C446FD8BAF35E507" box="[789,882,528,549]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="genus">
<emphasis id="F8EFEAF033335C20C446FD8BAF35E507" box="[789,882,528,549]" italics="true" pageId="9" pageNumber="986">Micrillus</emphasis>
</taxonomicName>
species previously recorded from
<collectingCountry id="B28C767233335C20C61BFDB6ADE4E560" box="[328,419,557,578]" name="Morocco" pageId="9" pageNumber="986">Morocco</collectingCountry>
as follows:
</paragraph>
<paragraph id="CA2436E233335C20C7E1FDC9AE67E581" blockId="9.[178,1085,594,675]" pageId="9" pageNumber="986">
from the widespread
<taxonomicName id="0D9B4D6133335C20C6C3FDC8AE56E54A" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[400,529,595,616]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="F8EFEAF033335C20C6C3FDC8AE56E54A" box="[400,529,595,616]" italics="true" pageId="9" pageNumber="986">M. testaceus</emphasis>
</taxonomicName>
(ERICHSON, 1840) by the coloration, a less flat forebody, longer antennae with more oblong antennomeres IV-X, a more slender and more oblong pronotum, and longer elytra;
</paragraph>
<paragraph id="CA2436E233335C20C7E1FD29AFABE402" blockId="9.[178,1086,690,800]" pageId="9" pageNumber="986">
from
<taxonomicName id="0D9B4D6133335C20C7BAFD28AD36E5E5" baseAuthorityName="Fauvel" baseAuthorityYear="1875" box="[233,369,690,712]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="F8EFEAF033335C20C7BAFD28AD36E5E5" box="[233,369,690,712]" italics="true" pageId="9" pageNumber="986">M. scabrosus</emphasis>
</taxonomicName>
(FAUVEL, 1875) and
<taxonomicName id="0D9B4D6133335C20C518FD28AEA5E5EA" authorityName="Assing" authorityYear="2008" box="[587,738,691,712]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="tenuipennis">
<emphasis id="F8EFEAF033335C20C518FD28AEA5E5EA" box="[587,738,691,712]" italics="true" pageId="9" pageNumber="986">M. tenuipennis</emphasis>
</taxonomicName>
<bibRefCitation id="AE0A4B1333335C20C5BBFD29AF33E5E5" author="ASSING V" box="[744,884,690,711]" journalOrPublisher="Neue" pageId="9" pageNumber="986" pagination="301 - 344" part="1" refId="ref9555" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stu ttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." title="A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stu ttgarter Beitrage zur Naturkunde" type="journal article" year="2008">ASSING, 2008</bibRefCitation>
(both distributed in North
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) by a much paler and distinctly bicoloured body, a less flat forebody, a relatively smaller head, much larger eyes (strongly reduced in
<taxonomicName id="0D9B4D6133335C20C409FD76AFA1E420" baseAuthorityName="Fauvel" baseAuthorityYear="1875" box="[858,998,749,770]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="F8EFEAF033335C20C409FD76AFA1E420" box="[858,998,749,770]" italics="true" pageId="9" pageNumber="986">M. scabrosus</emphasis>
</taxonomicName>
and
<taxonomicName id="0D9B4D6133335C20C372FD76AD60E402" authorityName="Assing" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="tenuipennis">
<emphasis id="F8EFEAF033335C20C372FD76AD60E402" italics="true" pageId="9" pageNumber="986">M. tenuipennis</emphasis>
</taxonomicName>
), a posterior less strongly tapering pronotum, and much longer elytra.
</paragraph>
<paragraph id="CA2436E233335C20C7E1FCABA858E467" blockId="9.[178,1055,815,837]" box="[178,1055,815,837]" pageId="9" pageNumber="986">
For illustrations of
<taxonomicName id="0D9B4D6133335C20C626FCABADB4E467" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[373,499,816,837]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="F8EFEAF033335C20C626FCABADB4E467" box="[373,499,816,837]" italics="true" pageId="9" pageNumber="986">M. testaceus</emphasis>
</taxonomicName>
,
<taxonomicName id="0D9B4D6133335C20C553FCABAEC0E466" baseAuthorityName="Fauvel" baseAuthorityYear="1875" box="[512,647,815,837]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="F8EFEAF033335C20C553FCABAEC0E466" box="[512,647,815,837]" italics="true" pageId="9" pageNumber="986">M. scabrosus</emphasis>
</taxonomicName>
, and
<taxonomicName id="0D9B4D6133335C20C5EDFCABAF12E467" authorityName="Assing" authorityYear="2008" box="[702,853,816,837]" class="Insecta" family="Staphylinidae" genus="Micrillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="986" phylum="Arthropoda" rank="species" species="tenuipennis">
<emphasis id="F8EFEAF033335C20C5EDFCABAF12E467" box="[702,853,816,837]" italics="true" pageId="9" pageNumber="986">M. tenuipennis</emphasis>
</taxonomicName>
see
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.
</paragraph>
<paragraph id="CA2436E233335C20C7E1FCCEAEC8E4AA" blockId="9.[178,1085,852,904]" pageId="9" pageNumber="986">
D i s t r i b u t i o n: The
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locality is situated to the southeast of Zagora, South
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, not far from the border with
<collectingCountry id="B28C767233335C20C56FFCE9AECCE4AA" box="[572,651,882,904]" name="Algeria" pageId="9" pageNumber="986">Algeria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>