292 lines
32 KiB
XML
292 lines
32 KiB
XML
<document ID-DOI="10.11646/zootaxa.4766.1.6" ID-GBIF-Dataset="286b29e4-4dce-4f68-b57a-9479c0764190" ID-ISSN="1175-5334" ID-Zenodo-Dep="3762612" ID-ZooBank="urn:lsid:zoobank.org:pub:3ABD266F-875A-415F-B794-D9562EC8CEE0" checkinTime="1587585237912" checkinUser="carolina" docAuthor="Brugnera, Ricardo, Paim, Marcelo Reginato, Roell, Talita, Dellapé, Gimena & Grazia, Jocelia" docDate="2020" docId="421B8792FFF06C632CC97F7A245B8D8B" docLanguage="en" docName="zootaxa.4766.1.6.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.10:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="10" docTitle="Tylospilus armatusimilis Brugnera, Paim & Grazia 2020" docType="treatment" docVersion="8" lastPageNumber="134" masterDocId="BE22FFEAFFF66C652C5E7B5B261D8A6F" masterDocTitle="Taxonomic modifications in Tylospilus Stål (Hemiptera: Pentatomidae: Asopinae) redescription of T. nigrobinotatus (Berg), description of a new species and an updated identification key for the genus" masterLastPageNumber="138" masterPageNumber="128" pageNumber="134" updateTime="1669135604859" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic modifications in Tylospilus Stål (Hemiptera: Pentatomidae: Asopinae) redescription of T. nigrobinotatus (Berg), description of a new species and an updated identification key for the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Brugnera, Ricardo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Paim, Marcelo Reginato</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Roell, Talita</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dellapé, Gimena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grazia, Jocelia</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-04-17</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4766</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>128</mods:start>
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<mods:end>138</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.4766.1.6</mods:identifier>
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<mods:identifier type="GBIF-Dataset">286b29e4-4dce-4f68-b57a-9479c0764190</mods:identifier>
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<mods:identifier type="ISSN">1175-5334</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3762612</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:3ABD266F-875A-415F-B794-D9562EC8CEE0</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3804197" ID-GBIF-Taxon="163732755" ID-Zenodo-Dep="3804197" LSID="urn:lsid:plazi:treatment:421B8792FFF06C632CC97F7A245B8D8B" httpUri="http://treatment.plazi.org/id/421B8792FFF06C632CC97F7A245B8D8B" lastPageNumber="134" pageId="6" pageNumber="134">
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<subSubSection box="[151,780,1057,1084]" pageId="6" pageNumber="134" type="nomenclature">
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||
<paragraph blockId="6.[151,780,1057,1119]" box="[151,780,1057,1084]" pageId="6" pageNumber="134">
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||
<heading bold="true" box="[151,780,1057,1084]" fontSize="10" level="2" pageId="6" pageNumber="134" reason="2">
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<taxonomicName authority="Brugnera, Paim & Grazia" authorityName="Brugnera, Paim & Grazia" authorityYear="2020" box="[151,780,1057,1084]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="armatusimilis">
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||
<emphasis bold="true" box="[151,780,1057,1084]" pageId="6" pageNumber="134">
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||
<emphasis bold="true" box="[151,447,1057,1084]" italics="true" pageId="6" pageNumber="134">Tylospilus armatusimilis</emphasis>
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Brugnera, Paim & Grazia
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,522,1093,1119]" pageId="6" pageNumber="134" type="description">
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<paragraph blockId="6.[151,780,1057,1119]" box="[151,522,1093,1119]" pageId="6" pageNumber="134">
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<figureCitation box="[151,356,1093,1119]" captionStart="FIGURES 17–24" captionStartId="4.[151,264,1971,1996]" captionTargetBox="[155,1430,529,1935]" captionTargetId="figure@4.[152,1436,525,1946]" captionTargetPageId="4" captionText="FIGURES 17–24. Habitus of Tylospilus nigrobinotatus (17–19, 23) and Tylospilus armatusimilis sp. nov. (20–22, 24). Dorsal (17, 20), lateral (18, 21), ventral (19, 22) and frontal (23, 24) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762616" httpUri="https://zenodo.org/record/3762616/files/figure.png" pageId="6" pageNumber="134">Figures 20–22, 24</figureCitation>
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,
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<figureCitation box="[368,522,1093,1119]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">28–30, 34–36</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="6" pageNumber="134" type="materials_examined">
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||
<paragraph blockId="6.[151,1437,1164,2020]" pageId="6" pageNumber="134">
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<materialsCitation collectingDate="2005-12-30" collectionCode="MNHNM" collectorName="G. J. Wibmer" country="Uruguay" location="Piedra de Los Indios" pageId="6" pageNumber="134" specimenCount="1" specimenCount-female="1" stateProvince="Colonia" typeStatus="holotype">
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||
<emphasis bold="true" box="[151,524,1164,1191]" pageId="6" pageNumber="134">
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||
<typeStatus box="[151,262,1165,1191]" pageId="6" pageNumber="134">Holotype</typeStatus>
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||
.
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||
<collectingCountry box="[275,418,1164,1191]" name="Uruguay" pageId="6" pageNumber="134">URUGUAY</collectingCountry>
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||
:
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||
<collectingRegion box="[431,524,1164,1191]" country="Uruguay" name="Colonia" pageId="6" pageNumber="134">Colonia</collectingRegion>
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||
</emphasis>
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||
<specimenCount box="[532,553,1166,1192]" pageId="6" pageNumber="134" type="female">♀</specimenCount>
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||
,
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||
<locationDeviation box="[569,776,1164,1192]" pageId="6" pageNumber="134">
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||
Ruta
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||
<quantity box="[630,702,1165,1191]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="134" unit="km" value="21.0">21 km</quantity>
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||
184.5
|
||
</locationDeviation>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:421B8792FFF06C632CC97F7A245B8D8B:CF6D605FFFF06C632F4A7FD6221F8EC8" box="[788,1026,1165,1192]" country="Uruguay" name="Piedra de Los Indios" pageId="6" pageNumber="134" stateProvince="Colonia">Piedra de Los Indios</location>
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||
,
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||
<date box="[1038,1177,1164,1191]" pageId="6" pageNumber="134" value="2005-12-30">
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||
<collectingDate box="[1038,1177,1164,1191]" pageId="6" pageNumber="134" value="2005-12-30">30.XII.2005</collectingDate>
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||
</date>
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||
, beating
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||
<taxonomicName authority=", G. J. Wibmer" authorityName="G. J. Wibmer" class="Magnoliopsida" family="Anacardiaceae" genus="Schinus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="134" phylum="Tracheophyta" rank="genus">
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||
<emphasis box="[1281,1368,1165,1191]" italics="true" pageId="6" pageNumber="134">Schinus</emphasis>
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||
,
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||
<collectorName pageId="6" pageNumber="134">G. J. Wibmer</collectorName>
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||
</taxonomicName>
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||
leg. (
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||
<collectionCode box="[305,422,1201,1227]" collectionName="MNHNM" country="Mexico" httpUri="http://grbio.org/cool/nctz-br8x" name="Museo Nacional de Historia Natural, Mexico City" pageId="6" pageNumber="134">MNHNM</collectionCode>
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||
).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph blockId="6.[151,1437,1164,2020]" pageId="6" pageNumber="134">
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||
<emphasis bold="true" box="[199,587,1236,1263]" pageId="6" pageNumber="134">
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||
<typeStatus box="[199,321,1237,1263]" pageId="6" pageNumber="134">Paratypes</typeStatus>
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||
.
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||
<materialsCitation box="[336,1064,1236,1264]" collectingDate="2005-12-30" collectionCode="MNHNM" collectorName="G. J. Wibmer" country="Uruguay" location="Piedra de Los Indios" pageId="6" pageNumber="134" specimenCount="1" specimenCount-male="1" stateProvince="Colonia">
|
||
<collectingCountry box="[336,479,1236,1263]" name="Uruguay" pageId="6" pageNumber="134">URUGUAY</collectingCountry>
|
||
:
|
||
<collectingRegion box="[494,587,1236,1263]" country="Uruguay" name="Colonia" pageId="6" pageNumber="134">Colonia</collectingRegion>
|
||
<specimenCount box="[597,634,1237,1263]" pageId="6" pageNumber="134" type="male">1♂</specimenCount>
|
||
, Same data as Holotype (
|
||
<collectionCode box="[935,1052,1237,1263]" collectionName="MNHNM" country="Mexico" httpUri="http://grbio.org/cool/nctz-br8x" name="Museo Nacional de Historia Natural, Mexico City" pageId="6" pageNumber="134">MNHNM</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
</emphasis>
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||
<materialsCitation collectingDate="2002-12-25" collectionCode="JEE" collectorName="G. J. Wibmer" country="Uruguay" location="Piedra de Los Indios" pageId="6" pageNumber="134" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount box="[1074,1120,1237,1264]" pageId="6" pageNumber="134" type="female">1 ♀</specimenCount>
|
||
,
|
||
<locationDeviation box="[1137,1350,1236,1264]" pageId="6" pageNumber="134">
|
||
Ruta
|
||
<quantity box="[1200,1274,1237,1263]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="134" unit="km" value="21.0">21 km</quantity>
|
||
184.5
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:421B8792FFF06C632CC97F7A245B8D8B:CF6D605FFFF06C63290B7F8E272A8F7C" name="Piedra de Los Indios" pageId="6" pageNumber="134">Piedra de Los Indios</location>
|
||
,
|
||
<date box="[325,464,1272,1299]" pageId="6" pageNumber="134" value="2002-12-25">
|
||
<collectingDate box="[325,464,1272,1299]" pageId="6" pageNumber="134" value="2002-12-25">25.XII.2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[478,632,1273,1300]" pageId="6" pageNumber="134">G. J. Wibmer</collectorName>
|
||
leg. (
|
||
<collectionCode box="[700,751,1273,1299]" collectionName="JEE" pageId="6" pageNumber="134">JEE</collectionCode>
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||
);
|
||
</materialsCitation>
|
||
<materialsCitation collectingDate="2002-12-23" collectionCode="MZUSP" collectorName="G. J. Wibmer" country="Uruguay" location="Piedra de Los Indios" pageId="6" pageNumber="134" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount box="[773,808,1273,1299]" pageId="6" pageNumber="134" type="female">1♀</specimenCount>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:421B8792FFF06C632CC97F7A245B8D8B:CF6D605FFFF06C632F667FA222348F7C" box="[824,1065,1273,1300]" name="Piedra de Los Indios" pageId="6" pageNumber="134">Piedra de Los Indios</location>
|
||
,
|
||
<date box="[1078,1217,1273,1299]" pageId="6" pageNumber="134" value="2002-12-23">
|
||
<collectingDate box="[1078,1217,1273,1299]" pageId="6" pageNumber="134" value="2002-12-23">23.XII.2002</collectingDate>
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||
</date>
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||
,
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||
<collectorName box="[1232,1386,1273,1300]" pageId="6" pageNumber="134">G. J. Wibmer</collectorName>
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||
leg. (
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||
<collectionCode box="[159,259,1309,1335]" collectionName="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="134">MZUSP</collectionCode>
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||
);
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||
</materialsCitation>
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||
<materialsCitation box="[278,1436,1309,1336]" collectingDate="1989-02-09" collectionCode="UFRG" collectorName="C. W. & L. O'Brien & G. Wibmer" country="Uruguay" location="Arroyo San Pedro" pageId="6" pageNumber="134" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
|
||
<specimenCount box="[278,320,1309,1336]" pageId="6" pageNumber="134" type="female">3 ♀</specimenCount>
|
||
<specimenCount box="[328,364,1309,1335]" pageId="6" pageNumber="134" type="male">1♂</specimenCount>
|
||
,
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||
<locationDeviation box="[377,566,1309,1336]" pageId="6" pageNumber="134">Ruta 21, km 194</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:421B8792FFF06C632CC97F7A245B8D8B:CF6D605FFFF06C632E1F7E4625138F58" box="[577,782,1309,1335]" name="Arroyo San Pedro" pageId="6" pageNumber="134">Arroyo San Pedro</location>
|
||
,
|
||
<date box="[794,898,1309,1335]" pageId="6" pageNumber="134" value="1989-02-09">
|
||
<collectingDate box="[794,898,1309,1335]" pageId="6" pageNumber="134" value="1989-02-09">9.II.1989</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[910,1125,1309,1336]" pageId="6" pageNumber="134">C. W. & L.O’Brien</collectorName>
|
||
&
|
||
<collectorName box="[1162,1287,1309,1335]" pageId="6" pageNumber="134">G. Wibmer</collectorName>
|
||
leg (
|
||
<collectionCode box="[1343,1424,1309,1335]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="6" pageNumber="134">UFRG</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation box="[151,1024,1345,1372]" collectingDate="1991-12-28" collectionCode="JEE" collectorName="G. Wibmer" country="Uruguay" location="Arroyo San Pedro" pageId="6" pageNumber="134" specimenCount="2" specimenCount-female="2">
|
||
<specimenCount box="[151,193,1345,1372]" pageId="6" pageNumber="134" type="female">2 ♀</specimenCount>
|
||
,
|
||
<locationDeviation box="[207,396,1345,1372]" pageId="6" pageNumber="134">Ruta 21, km 194</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:421B8792FFF06C632CC97F7A245B8D8B:CF6D605FFFF06C632DC87E1A247E8F34" box="[406,611,1345,1371]" country="Saint Pierre and Miquelon" county="Piedra de Los Indios" municipality="Piedra de Los Indios" name="Arroyo San Pedro" pageId="6" pageNumber="134">Arroyo San Pedro</location>
|
||
,
|
||
<date box="[622,762,1345,1371]" pageId="6" pageNumber="134" value="1991-12-28">
|
||
<collectingDate box="[622,762,1345,1371]" pageId="6" pageNumber="134" value="1991-12-28">28.XII.1991</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[773,898,1345,1371]" pageId="6" pageNumber="134">G. Wibmer</collectorName>
|
||
leg. (
|
||
<collectionCode box="[961,1012,1345,1371]" collectionName="JEE" pageId="6" pageNumber="134">JEE</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="134" type="diagnosis">
|
||
<paragraph blockId="6.[151,1437,1164,2020]" pageId="6" pageNumber="134">
|
||
<emphasis bold="true" box="[199,322,1380,1407]" pageId="6" pageNumber="134">Diagnosis.</emphasis>
|
||
Similar to
|
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<taxonomicName authorityName=": Thomas" authorityYear="1992" box="[452,645,1381,1407]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="nigrobinotatus">
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||
<emphasis box="[452,645,1381,1407]" italics="true" pageId="6" pageNumber="134">T. nigrobinotatus</emphasis>
|
||
</taxonomicName>
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||
in general aspects,
|
||
<taxonomicName authorityName="Brugnera, Paim & Grazia" authorityYear="2020" box="[871,1055,1381,1407]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="armatusimilis">
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||
<emphasis box="[871,1055,1381,1407]" italics="true" pageId="6" pageNumber="134">T. armatusimilis</emphasis>
|
||
</taxonomicName>
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||
can be recognized by the anterolateral margins of the pronotum concolorous with the pronotal disc; humeral angles shorter and slender, laterally directed (
|
||
<figureCitation box="[256,429,1453,1479]" captionStart="FIGURES 17–24" captionStartId="4.[151,264,1971,1996]" captionTargetBox="[155,1430,529,1935]" captionTargetId="figure@4.[152,1436,525,1946]" captionTargetPageId="4" captionText="FIGURES 17–24. Habitus of Tylospilus nigrobinotatus (17–19, 23) and Tylospilus armatusimilis sp. nov. (20–22, 24). Dorsal (17, 20), lateral (18, 21), ventral (19, 22) and frontal (23, 24) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762616" httpUri="https://zenodo.org/record/3762616/files/figure.png" pageId="6" pageNumber="134">Figs 20–22, 24</figureCitation>
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); black spots on the apices of radial veins distinctly smaller than in
|
||
<taxonomicName authorityName=": Thomas" authorityYear="1992" box="[1184,1376,1453,1479]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="nigrobinotatus">
|
||
<emphasis box="[1184,1376,1453,1479]" italics="true" pageId="6" pageNumber="134">T. nigrobinotatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="FIGURES 17–24" captionStartId="4.[151,264,1971,1996]" captionTargetBox="[155,1430,529,1935]" captionTargetId="figure@4.[152,1436,525,1946]" captionTargetPageId="4" captionText="FIGURES 17–24. Habitus of Tylospilus nigrobinotatus (17–19, 23) and Tylospilus armatusimilis sp. nov. (20–22, 24). Dorsal (17, 20), lateral (18, 21), ventral (19, 22) and frontal (23, 24) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762616" httpUri="https://zenodo.org/record/3762616/files/figure.png" pageId="6" pageNumber="134">Fig. 20</figureCitation>
|
||
), and lateral row of black spots on the urosternites restricted to the anterior margin of each urosternite (
|
||
<figureCitation box="[1339,1423,1489,1515]" captionStart="FIGURES 17–24" captionStartId="4.[151,264,1971,1996]" captionTargetBox="[155,1430,529,1935]" captionTargetId="figure@4.[152,1436,525,1946]" captionTargetPageId="4" captionText="FIGURES 17–24. Habitus of Tylospilus nigrobinotatus (17–19, 23) and Tylospilus armatusimilis sp. nov. (20–22, 24). Dorsal (17, 20), lateral (18, 21), ventral (19, 22) and frontal (23, 24) views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762616" httpUri="https://zenodo.org/record/3762616/files/figure.png" pageId="6" pageNumber="134">Fig. 22</figureCitation>
|
||
); females have a well-defined rounded projection on gonocoxites XIII (
|
||
<figureCitation box="[957,1092,1525,1551]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">Figs 28, 29</figureCitation>
|
||
), secondary thickening of gonapophyses IX sickle-like (
|
||
<figureCitation box="[457,540,1561,1587]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">Fig. 30</figureCitation>
|
||
), anterior projection of thickening of vaginal intima v-shaped (
|
||
<figureCitation box="[1243,1324,1561,1587]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">Fig. 30</figureCitation>
|
||
) and
|
||
<emphasis italics="true" pageId="6" pageNumber="134">capsula seminalis</emphasis>
|
||
elliptical (
|
||
<figureCitation box="[428,511,1597,1623]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">Fig. 30</figureCitation>
|
||
); in males, the mesial portion of ventral rim slightly concave (
|
||
<figureCitation box="[1196,1279,1597,1623]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">Fig. 33</figureCitation>
|
||
), and superior process of dorsal rim of pygophore subrectangular (
|
||
<figureCitation box="[728,811,1632,1659]" captionStart="FIGURES 25–36" captionStartId="5.[151,264,1783,1808]" captionTargetBox="[202,1385,189,1757]" captionTargetId="figure@5.[200,1388,181,1757]" captionTargetPageId="5" captionText="FIGURES 25–36. Genitalia of Tylospilus nigrobinotatus (25–27, 31–33) and Tylospilus armatusimilis sp. nov. (28–30, 34–36). Female (25–30). Male (31–36). Genital plates in posterior view (25, 28). Gonocoxites VIII in lateral view (26, 29). Internal genitalia (27, 30). Pygophore in dorsal (31, 34), posterior (32, 35) and ventral (33, 36) views. Abbreviations: ao = anterior opening; cs = capsula seminalis; dpdr = distal part of ductus receptaculi; dr = dorsal rim; gcVIII = gonocoxites VIII; gcIX = gonocoxites IX; laVIII = laterotergites VIII; laIX = laterotergites IX; par = paramere; pi = pars intermedialis; ppdr = proximal part of ductus receptaculi; spdr = superior process of dorsal rim; st = secondary thickening of gonapophyses IX; tvi = thickening of vaginal intima; va = vesicular area; vr = ventral rim; x = abdominal segment X. Scale bars: 25, 28 = 0.5 mm; 26–27, 29–36 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762618" httpUri="https://zenodo.org/record/3762618/files/figure.png" pageId="6" pageNumber="134">Fig. 35</figureCitation>
|
||
). A differential diagnosis between
|
||
<taxonomicName authorityName=": Thomas" authorityYear="1992" box="[1198,1389,1633,1659]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="nigrobinotatus">
|
||
<emphasis box="[1198,1389,1633,1659]" italics="true" pageId="6" pageNumber="134">T. nigrobinotatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Brugnera & Paim & Roell & Dellapé & Grazia, 2020" authorityName="Brugnera & Paim & Roell & Dellapé & Grazia" authorityYear="2020" box="[151,333,1669,1695]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="armatusimilis" status="sp. nov.">
|
||
<emphasis box="[151,333,1669,1695]" italics="true" pageId="6" pageNumber="134">T. armatusimilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[340,429,1669,1695]" pageId="6" pageNumber="134">
|
||
<taxonomicNameLabel box="[340,429,1669,1695]" pageId="6" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is presented on table 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="134" type="description">
|
||
<paragraph blockId="6.[151,1437,1164,2020]" pageId="6" pageNumber="134">
|
||
<emphasis bold="true" box="[199,384,1705,1731]" pageId="6" pageNumber="134">Measurements:</emphasis>
|
||
(n=10). Head length: 1.2 ± 0.10 (1.1–1.5), width: 1.6 ± 0.07 (1.5–1.7); pronotum length: 1.7 ± 0.15 (1.4–1.9), width: 5.55 ± 0.43 (4.7–6.0); scutellum length: 2.7 ± 0.19 (2.3–2.9), width: 2.45 ± 0.18 (2.4–2.8); antennomeres length: I 0.22 ± 0.05 (0.2–0.3), II 0.82 ± 0.5 (0.8–0.9), III 0.65 ± 0.5 (0.6–0.7), IV 0.6 ± 0.0 (0.6–0.6); V 0.5 ± 0.0 (0.5–0.5); labial segments length: I 0.75 ± 0.05 (0.7–0.8), II 0.85 ± 0.04 (0.8–0.9), III 0.58 ± 0.03 (0.5–0.6), IV 0.59 ± 0.08 (0.5–0.6); abdominal length: 3.97 ± 1.07 (3.2–5.8), width: 3.8 ± 0.95 (3.1–5.3); total length: 7.5 ± 0.50 (7.0–8.4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="134" type="etymology">
|
||
<paragraph blockId="6.[151,1437,1164,2020]" pageId="6" pageNumber="134">
|
||
<emphasis bold="true" box="[199,334,1921,1947]" pageId="6" pageNumber="134">Etymology.</emphasis>
|
||
From
|
||
<taxonomicName authority="(Latin)" baseAuthorityName="Latin" box="[410,587,1921,1948]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis box="[410,502,1922,1947]" italics="true" pageId="6" pageNumber="134">armatus</emphasis>
|
||
(Latin)
|
||
</taxonomicName>
|
||
and
|
||
<emphasis box="[643,716,1921,1947]" italics="true" pageId="6" pageNumber="134">similis</emphasis>
|
||
(Latin) in reference to
|
||
<taxonomicName box="[979,1099,1922,1947]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis box="[979,1099,1922,1947]" italics="true" pageId="6" pageNumber="134">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation author="Thomas, D. B." box="[1257,1431,1921,1947]" pageId="6" pageNumber="134" refId="ref7830" refString="Thomas, D. B. (1992) Taxonomic synopsis of the asopine Pentatomidae (Heteroptera) of the Western Hemisphere. Entomological Society of America, Lanham, Maryland, 156 pp." type="book" year="1992">Thomas (1992)</bibRefCitation>
|
||
, here considered to be a junior synonym of
|
||
<emphasis box="[630,829,1957,1983]" italics="true" pageId="6" pageNumber="134">
|
||
<taxonomicName authorityName=": Thomas" authorityYear="1992" box="[630,826,1957,1983]" class="Insecta" family="Pentatomidae" genus="Tylospilus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="134" phylum="Arthropoda" rank="species" species="nigrobinotatus">T. nigrobinotatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,582,1992,2020]" pageId="6" pageNumber="134" type="distribution">
|
||
<paragraph blockId="6.[151,1437,1164,2020]" box="[199,582,1992,2020]" pageId="6" pageNumber="134">
|
||
<emphasis bold="true" box="[199,355,1992,2019]" pageId="6" pageNumber="134">Distribution:</emphasis>
|
||
<collectingCountry box="[362,460,1994,2020]" name="Uruguay" pageId="6" pageNumber="134">Uruguay</collectingCountry>
|
||
(
|
||
<collectingRegion box="[475,570,1993,2020]" country="Uruguay" name="Colonia" pageId="6" pageNumber="134">Colonia</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |