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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.90667" ID-Pensoft-Pub="1313-2970-1145-181" ID-Pensoft-UUID="51133424FE065812A1E6434518904D50" ID-ZooBank="60D183EBA5A349AB8A6CA2BA9B395E41" ModsDocID="1313-2970-1145-181" checkinTime="1675506015366" checkinUser="pensoft" docAuthor="D'Alessandro, Paola &amp; Biondi, Maurizio" docDate="2023" docId="421B5B245870506F97848FB162B43101" docLanguage="en" docName="ZooKeys 1145: 181-189" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.90667" docTitle="Procalus maculipennis D'Alessandro &amp; Biondi 2023, comb. nov." docType="treatment" docVersion="1" id="51133424FE065812A1E6434518904D50" lastPageNumber="181" masterDocId="51133424FE065812A1E6434518904D50" masterDocTitle="Generic placement of the African flea beetle Polyclada maculipennis Bryant and the possible occurrence of the genus Procalus Clark in sub-Saharan Africa (Coleoptera, Chrysomelidae, Galerucinae, Alticini)" masterLastPageNumber="189" masterPageNumber="181" pageNumber="181" updateTime="1675506015366" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Generic placement of the African flea beetle Polyclada maculipennis Bryant and the possible occurrence of the genus Procalus Clark in sub-Saharan Africa (Coleoptera, Chrysomelidae, Galerucinae, Alticini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>D'Alessandro, Paola</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4481-9152</mods:nameIdentifier>
<mods:affiliation>Department of Life, Health &amp; Environmental Sciences, University of L'Aquila, Via Vetoio-Coppito, 67100 L'Aquila, Italy</mods:affiliation>
<mods:nameIdentifier type="email">paola.dalessandro@univaq.it</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Biondi, Maurizio</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2190-7376</mods:nameIdentifier>
<mods:affiliation>Department of Life, Health &amp; Environmental Sciences, University of L'Aquila, Via Vetoio-Coppito, 67100 L'Aquila, Italy</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1145</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>181</mods:start>
<mods:end>189</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1145.90667</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.90667</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1145-181</mods:identifier>
<mods:identifier type="ZooBank">60D183EBA5A349AB8A6CA2BA9B395E41</mods:identifier>
<mods:identifier type="Pensoft-UUID">51133424FE065812A1E6434518904D50</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:421B5B245870506F97848FB162B43101" httpUri="http://treatment.plazi.org/id/421B5B245870506F97848FB162B43101" lastPageNumber="181" pageId="0" pageNumber="181">
<subSubSection pageId="0" pageNumber="181" type="nomenclature">
<paragraph pageId="0" pageNumber="181">
<taxonomicName LSID="421B5B24-5870-506F-9784-8FB162B43101" authority="(Bryant, 1942)" authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="Bryant" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis" status="comb. nov.">Procalus maculipennis (Bryant, 1942)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="181">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="description">
<paragraph pageId="0" pageNumber="181">
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Lectotype of Polyclada maculipennis Bryant, 1942 1 habitus, dorsal view 2 ibid., labels 3 ibid., habitus, lateral view 4 Polyclada bohemani Baly, male, Kenya (BAQ), habitus." figureDoi="10.3897/zookeys.1145.90667.figures1-4" httpUri="https://binary.pensoft.net/fig/805298" pageId="0" pageNumber="181">Figs 1-3</figureCitation>
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">, 5-8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="reference_group">
<paragraph pageId="0" pageNumber="181">
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="D'Alessandro &amp; Biondi" baseAuthorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis" status="comb. nov.">Polyclada maculipennis</taxonomicName>
Bryant, 1942: 164.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="type material">
<paragraph pageId="0" pageNumber="181">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="181">
Lectotype of
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="D'Alessandro &amp; Biondi" baseAuthorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada maculipennis</emphasis>
</taxonomicName>
♂: &quot;Kamerun Conradt // Coll. Kraatz // Pres. By Imp. Inst. Ent. B.N. 1933-468 //
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="D'Alessandro &amp; Biondi" baseAuthorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada maculipennis</emphasis>
</taxonomicName>
Bryant / Det. G.E. Bryant&quot; (NHMUK) (here designated by M. Biondi and P.
<normalizedToken originalValue="D’Alessandro">D'Alessandro</normalizedToken>
) (Figs
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Lectotype of Polyclada maculipennis Bryant, 1942 1 habitus, dorsal view 2 ibid., labels 3 ibid., habitus, lateral view 4 Polyclada bohemani Baly, male, Kenya (BAQ), habitus." figureDoi="10.3897/zookeys.1145.90667.figures1-4" httpUri="https://binary.pensoft.net/fig/805298" pageId="0" pageNumber="181">1-3</figureCitation>
). Paralectotypes of
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="D'Alessandro &amp; Biondi" baseAuthorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada maculipennis</emphasis>
</taxonomicName>
, 2♀♀: same data as for lectotype (NHMUK).
</paragraph>
<caption doi="10.3897/zookeys.1145.90667.figures1-4" httpUri="https://binary.pensoft.net/fig/805298" pageId="0" pageNumber="181" start="Figures 1–4" startId="F1">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figures 1-4.</emphasis>
Lectotype of
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="D'Alessandro &amp; Biondi" baseAuthorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada maculipennis</emphasis>
</taxonomicName>
Bryant, 1942
<emphasis bold="true" pageId="0" pageNumber="181">1</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="181">2</emphasis>
ibid., labels
<emphasis bold="true" pageId="0" pageNumber="181">3</emphasis>
ibid., habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="181">4</emphasis>
<taxonomicName authorityName="Baly" authorityYear="1861" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada bohemani" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="bohemani">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada bohemani</emphasis>
</taxonomicName>
Baly, male, Kenya (BAQ), habitus.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="materials_examined">
<paragraph pageId="0" pageNumber="181">
Material of
<taxonomicName authorityName="Clark" authorityYear="1865" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus</emphasis>
</taxonomicName>
species examined for comparison.
</paragraph>
<paragraph pageId="0" pageNumber="181">
<materialsCitation collectingDate="1899-05" collectingDateMax="1986-11-20" collectingDateMin="1899-05" collectorName="Coll. Nickerl, Det. J. Bechyne, D. Ed. Varas Arangua, F. Silvestri, P. Herbst" country="Chile" determinerName="Det. J. Bechyne" location="Vina del Mar" specimenCount="14" specimenCount-female="1" stateProvince="Valparaiso" typeStatus="syntype">
<collectingCountry name="Chile">Chile</collectingCountry>
:
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus reduplicatus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="reduplicatus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus reduplicatus</emphasis>
</taxonomicName>
,
<typeStatus>syntype</typeStatus>
,
<specimenCount type="female">1♀</specimenCount>
, &quot;
<collectingCountry name="Chile">Chile</collectingCountry>
//
<collectorName>Coll. Nickerl</collectorName>
/ Mus. Pragense //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus mutans" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mutans">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus mutans</emphasis>
</taxonomicName>
Blchd. // Typus //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus reduplicatus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="reduplicatus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus reduplicatus</emphasis>
</taxonomicName>
n. sp. Type
<specimenCount>♀</specimenCount>
/ 1948
<determinerName>
<collectorName>
Det. J.
<normalizedToken originalValue="Bechyně”">Bechyne&quot;</normalizedToken>
</collectorName>
</determinerName>
(NMPC);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus viridis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="viridis">Procalus cf. viridis</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
, El Tabo,
<collectingRegion country="Chile" name="Valparaiso">Valparaiso</collectingRegion>
,
<collectingDate value="1986-09-29">29 Sept. 1986</collectingDate>
(NHMB);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus lenzi" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="lenzi">Procalus cf. lenzi</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
,
<collectingRegion country="Chile" name="Region Metropolitana">Santiago</collectingRegion>
, Pudahuel,
<collectingDate value="1986-11-20">20 Nov. 1986</collectingDate>
(NHMB);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus malaisei" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="malaisei">Procalus cf. malaisei</taxonomicName>
,
<specimenCount type="generic">2 specimens</specimenCount>
,
<collectorName>D. Ed. Varas Arangua</collectorName>
leg., 1921 (MSNG);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus lenzi" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="lenzi">Procalus cf. lenzi</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
, ibid (MSNG);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus viridis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="viridis">Procalus cf. viridis</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:421B5B245870506F97848FB162B43101:481849A7C5921F6325E8C03C2DC69A7E" country="Chile" name="Vina del Mar" stateProvince="Valparaiso">
<normalizedToken originalValue="Viña">Vina</normalizedToken>
del Mar
</location>
,
<collectingRegion country="Chile" name="Valparaiso">Valparaiso</collectingRegion>
,
<collectingDate value="1899-05">May 1899</collectingDate>
,
<collectorName>F. Silvestri</collectorName>
leg. (MSNG);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus silvai" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="silvai">Procalus cf. silvai</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
, Concepcion, 1903,
<collectorName>P. Herbst</collectorName>
leg. (MSNG);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus reduplicatus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="reduplicatus">Procalus cf. reduplicatus</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
, ibid (MSNG);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus mutans" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mutans">Procalus cf. mutans</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
, Concepcion,
<collectingDate value="1903-09">Sept. 1903</collectingDate>
,
<collectorName>P. Herbst</collectorName>
leg. (MSNG);
<taxonomicName authorityName="Clark" authorityYear="1865" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus</emphasis>
</taxonomicName>
sp.,
<specimenCount type="generic">2 specimens</specimenCount>
, ibid (MSNG);
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus reduplicatus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="reduplicatus">Procalus cf. reduplicatus</taxonomicName>
,
<specimenCount type="generic">1 specimen</specimenCount>
, Concepcion, 1904,
<collectorName>P. Herbst</collectorName>
leg. (MSNG);
<taxonomicName authorityName="Clark" authorityYear="1865" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus</emphasis>
</taxonomicName>
sp.,
<specimenCount type="generic">1 specimen</specimenCount>
, ibid (MSNG)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="181" type="remarks">
<paragraph pageId="0" pageNumber="181">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="181">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada</emphasis>
</taxonomicName>
can be immediately distinguished from similar genera by the antennae, longer than half body length, with antennomeres 4-10 pectinate or flabellate in male and serrate in female (Fig.
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Lectotype of Polyclada maculipennis Bryant, 1942 1 habitus, dorsal view 2 ibid., labels 3 ibid., habitus, lateral view 4 Polyclada bohemani Baly, male, Kenya (BAQ), habitus." figureDoi="10.3897/zookeys.1145.90667.figures1-4" httpUri="https://binary.pensoft.net/fig/805298" pageId="0" pageNumber="181">4</figureCitation>
). Characters of the antenna along with other morphological features, such as procoxal cavities open posteriorly, antennomere 4 at least double length of antennomere 3, as well as elytral punctation always confused, densely and uniformly impressed, permits identification of the genus within the
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Blepharida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Blepharida" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Blepharida</emphasis>
</taxonomicName>
group in the Afrotropical region (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.253.3414" author="Biondi, M" journalOrPublisher="ZooKeys" pageId="0" pageNumber="181" pagination="1 - 158" refId="B5" refString="Biondi, M, D'Alessandro, P, 2012. Afrotropical flea beetle genera: A key to their identification, updated catalogue and biogeographical analysis (Coleoptera, Chrysomelidae, Galerucinae, Alticini). ZooKeys 253: 1 - 158, DOI: https://doi.org/10.3897/zookeys.253.3414" title="Afrotropical flea beetle genera: A key to their identification, updated catalogue and biogeographical analysis (Coleoptera, Chrysomelidae, Galerucinae, Alticini)." url="https://doi.org/10.3897/zookeys.253.3414" volume="253" year="2012">
Biondi and
<normalizedToken originalValue="D’Alessandro">D'Alessandro</normalizedToken>
2012
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1163/1876312X-48022152" author="Biondi, M" journalOrPublisher="Insect Systematics &amp; Evolution" pageId="0" pageNumber="181" pagination="97 - 155" refId="B6" refString="Biondi, M, Frasca, R, Grobbelaar, E, D'Alessandro, P, 2017. Supraspecific taxonomy of the flea beetle genus Blepharida Chevrolat, 1836 (Coleoptera: Chrysomelidae) in the Afrotropical Region and description of Afroblepharida subgen. nov. Insect Systematics &amp; Evolution 48 (2): 97 - 155, DOI: https://doi.org/10.1163/1876312X-48022152" title="Supraspecific taxonomy of the flea beetle genus Blepharida Chevrolat, 1836 (Coleoptera: Chrysomelidae) in the Afrotropical Region and description of Afroblepharida subgen. nov." url="https://doi.org/10.1163/1876312X-48022152" volume="48" year="2017">Biondi et al. 2017</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="181">
The type material of
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="D'Alessandro &amp; Biondi" baseAuthorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada maculipennis</emphasis>
</taxonomicName>
has all the diagnostic characters of
<taxonomicName authorityName="Clark" authorityYear="1865" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1080/00222936508679421" author="Clark, H" journalOrPublisher="Librairie Mequignon-Marvis Pere et Fils, Paris" pageId="0" pageNumber="181" refId="B14" refString="Clark, H, 1865. An examination of the Dejeanian genus Coelomera (Coleoptera-Phytophaga) and its affinities. The Annals and Magazine of Natural History (Series 3) 16: 256-268. https://doi.org/10.1080/00222936508679421" title="An examination of the Dejeanian genus Coelomera (Coleoptera-Phytophaga) and its affinities. The Annals and Magazine of Natural History (Series 3) 16: 256 - 268." url="https://doi.org/10.1080/00222936508679421" year="1865">Clark 1865</bibRefCitation>
;
<bibRefCitation author="Jerez, V" journalOrPublisher="Zoologia" pageId="0" pageNumber="181" pagination="109 - 125" publicationUrl="http://www.biodiversitylibrary.org/item/89079" refId="B28" refString="Jerez, V, 1992. Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana. Zoologia 56 (3-4): 109 - 125, http://www.biodiversitylibrary.org/item/89079" title="Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana." url="http://www.biodiversitylibrary.org/item/89079" volume="56" year="1992">Jerez 1992</bibRefCitation>
), none of which occur in any
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Polyclada" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Polyclada" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Polyclada</emphasis>
</taxonomicName>
species: antennae with antennomeres 1-5 flattened compared to 6-11, especially in male, and antennomere 1 clearly distally enlarged, and strikingly serrated in male (Fig.
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Lectotype of Polyclada maculipennis Bryant, 1942 1 habitus, dorsal view 2 ibid., labels 3 ibid., habitus, lateral view 4 Polyclada bohemani Baly, male, Kenya (BAQ), habitus." figureDoi="10.3897/zookeys.1145.90667.figures1-4" httpUri="https://binary.pensoft.net/fig/805298" pageId="0" pageNumber="181">1</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.1080/00222936508679421" author="Clark, H" journalOrPublisher="Librairie Mequignon-Marvis Pere et Fils, Paris" pageId="0" pageNumber="181" refId="B14" refString="Clark, H, 1865. An examination of the Dejeanian genus Coelomera (Coleoptera-Phytophaga) and its affinities. The Annals and Magazine of Natural History (Series 3) 16: 256-268. https://doi.org/10.1080/00222936508679421" title="An examination of the Dejeanian genus Coelomera (Coleoptera-Phytophaga) and its affinities. The Annals and Magazine of Natural History (Series 3) 16: 256 - 268." url="https://doi.org/10.1080/00222936508679421" year="1865">Clark 1865</bibRefCitation>
;
<bibRefCitation author="Jerez, V" journalOrPublisher="Zoologia" pageId="0" pageNumber="181" pagination="109 - 125" publicationUrl="http://www.biodiversitylibrary.org/item/89079" refId="B28" refString="Jerez, V, 1992. Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana. Zoologia 56 (3-4): 109 - 125, http://www.biodiversitylibrary.org/item/89079" title="Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana." url="http://www.biodiversitylibrary.org/item/89079" volume="56" year="1992">Jerez 1992</bibRefCitation>
: figs 4A, 5A, 6A, 7A, 8A, 1995: figs 1, 2); fifth abdominal sternite in male with a wide, deep, oval depression (Fig.
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">5</figureCitation>
;
<bibRefCitation author="Jerez, V" journalOrPublisher="Zoologia" pageId="0" pageNumber="181" pagination="109 - 125" publicationUrl="http://www.biodiversitylibrary.org/item/89079" refId="B28" refString="Jerez, V, 1992. Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana. Zoologia 56 (3-4): 109 - 125, http://www.biodiversitylibrary.org/item/89079" title="Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana." url="http://www.biodiversitylibrary.org/item/89079" volume="56" year="1992">Jerez 1992</bibRefCitation>
: figs 5E, 6D, 7C, 8F, 1995: fig. 2); metafemoral extensor tendon simplified, very slender (Fig.
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">6</figureCitation>
;
<bibRefCitation author="Furth, DG" editor="Furth, DG" journalOrPublisher="Backhuys Publishers, Leiden" pageId="0" pageNumber="181" pagination="116 - 135" refId="B23" refString="Furth, DG, Suzuki, K, 1994. Character correlation studies of problematic genera of Alticinae in relation to Galerucinae (Coleoptera: Chrysomelidae). In: Furth, DG, Ed., Proceedings of the Third International Symposium on the Chrysomelidae, Beijing, 1992. Backhuys Publishers, Leiden: 116 - 135" title="Character correlation studies of problematic genera of Alticinae in relation to Galerucinae (Coleoptera: Chrysomelidae)." volumeTitle="Proceedings of the Third International Symposium on the Chrysomelidae, Beijing, 1992." year="1994">Furth and Suzuki 1994</bibRefCitation>
: fig. 6b;
<bibRefCitation author="Jerez, V" journalOrPublisher="Zoologia" pageId="0" pageNumber="181" pagination="109 - 125" publicationUrl="http://www.biodiversitylibrary.org/item/89079" refId="B28" refString="Jerez, V, 1992. Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana. Zoologia 56 (3-4): 109 - 125, http://www.biodiversitylibrary.org/item/89079" title="Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana." url="http://www.biodiversitylibrary.org/item/89079" volume="56" year="1992">Jerez 1992</bibRefCitation>
: figs 2, 3D): dorsal lobe straight, with very elongate extended arm; central furrow very wide; ventral lobe subtriangular; recurved flange short, poorly sclerotized. Additionally, the median lobe of aedeagus and spermatheca (Figs
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">7</figureCitation>
,
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">8</figureCitation>
) are typical of the
<taxonomicName authorityName="Clark" authorityYear="1865" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus</emphasis>
</taxonomicName>
species (median lobe of aedeagus:
<bibRefCitation author="Furth, DG" editor="Furth, DG" journalOrPublisher="Backhuys Publishers, Leiden" pageId="0" pageNumber="181" pagination="116 - 135" refId="B23" refString="Furth, DG, Suzuki, K, 1994. Character correlation studies of problematic genera of Alticinae in relation to Galerucinae (Coleoptera: Chrysomelidae). In: Furth, DG, Ed., Proceedings of the Third International Symposium on the Chrysomelidae, Beijing, 1992. Backhuys Publishers, Leiden: 116 - 135" title="Character correlation studies of problematic genera of Alticinae in relation to Galerucinae (Coleoptera: Chrysomelidae)." volumeTitle="Proceedings of the Third International Symposium on the Chrysomelidae, Beijing, 1992." year="1994">Furth and Suzuki 1994</bibRefCitation>
: fig. 6a;
<bibRefCitation author="Jerez, V" journalOrPublisher="Zoologia" pageId="0" pageNumber="181" pagination="109 - 125" publicationUrl="http://www.biodiversitylibrary.org/item/89079" refId="B28" refString="Jerez, V, 1992. Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana. Zoologia 56 (3-4): 109 - 125, http://www.biodiversitylibrary.org/item/89079" title="Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana." url="http://www.biodiversitylibrary.org/item/89079" volume="56" year="1992">Jerez 1992</bibRefCitation>
: figs 4E, 5C, 6E, 7B, 8G, 1995: fig. 10; spermatheca:
<bibRefCitation author="Furth, DG" editor="Furth, DG" journalOrPublisher="Backhuys Publishers, Leiden" pageId="0" pageNumber="181" pagination="116 - 135" refId="B23" refString="Furth, DG, Suzuki, K, 1994. Character correlation studies of problematic genera of Alticinae in relation to Galerucinae (Coleoptera: Chrysomelidae). In: Furth, DG, Ed., Proceedings of the Third International Symposium on the Chrysomelidae, Beijing, 1992. Backhuys Publishers, Leiden: 116 - 135" title="Character correlation studies of problematic genera of Alticinae in relation to Galerucinae (Coleoptera: Chrysomelidae)." volumeTitle="Proceedings of the Third International Symposium on the Chrysomelidae, Beijing, 1992." year="1994">Furth and Suzuki 1994</bibRefCitation>
: fig. 6c;
<bibRefCitation author="Jerez, V" journalOrPublisher="Zoologia" pageId="0" pageNumber="181" pagination="109 - 125" publicationUrl="http://www.biodiversitylibrary.org/item/89079" refId="B28" refString="Jerez, V, 1992. Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana. Zoologia 56 (3-4): 109 - 125, http://www.biodiversitylibrary.org/item/89079" title="Revision taxonomica del genero Procalus Clark, 1865 (Chrysomelidae - Alticinae). Gayana." url="http://www.biodiversitylibrary.org/item/89079" volume="56" year="1992">Jerez 1992</bibRefCitation>
: figs 4D, 5D, 6B, 7D, 8E, 1995: fig. 11). Median lobe of the aedeagus of the lectotype here designated (Fig.
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">7</figureCitation>
): thickset and smooth; in ventral view tapering towards the apex, and slightly narrowing subapically; apex subtriangular, widely obtuse, with a small median tooth; in lateral view, clearly curved. Spermatheca of the paralectotypes (Fig.
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Procalus maculipennis Bryant, 1942 comb. nov. 5 ventral parts in male 6 metafemoral extensor tendon 7 median lobe of aedeagus in ventral, dorsal, and lateral views (a-c, respectively) 8 spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe." figureDoi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181">8</figureCitation>
): subcylindrical and elongate basally, curved towards the ductus attachment; distal part distinctly bent and about 1/3 the basal part in length; ductus basally inserted, short, uncoiled, moderately thickset. We therefore propose the new combination
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="Bryant" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus maculipennis</emphasis>
</taxonomicName>
(Bryant, 1942), comb. nov.
</paragraph>
<caption doi="10.3897/zookeys.1145.90667.figures5-8" httpUri="https://binary.pensoft.net/fig/805299" pageId="0" pageNumber="181" start="Figures 5–8" startId="F2">
<paragraph pageId="0" pageNumber="181">
<emphasis bold="true" pageId="0" pageNumber="181">Figures 5-8.</emphasis>
<taxonomicName authorityName="D'Alessandro &amp; Biondi" authorityYear="2023" baseAuthorityName="Bryant" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Procalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Procalus maculipennis" order="Coleoptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis italics="true" pageId="0" pageNumber="181">Procalus maculipennis</emphasis>
</taxonomicName>
Bryant, 1942 comb. nov.
<emphasis bold="true" pageId="0" pageNumber="181">5</emphasis>
ventral parts in male
<emphasis bold="true" pageId="0" pageNumber="181">6</emphasis>
metafemoral extensor tendon
<emphasis bold="true" pageId="0" pageNumber="181">7</emphasis>
median lobe of aedeagus in ventral, dorsal, and lateral views (
<emphasis bold="true" pageId="0" pageNumber="181">a-c</emphasis>
, respectively)
<emphasis bold="true" pageId="0" pageNumber="181">8</emphasis>
spermatheca. Abbreviations: cf: central furrow; dl: dorsal lobe; ea: extended arm; rf: recurved flange; vl: ventral lobe.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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