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<document ID-DOI="http://doi.org/10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" approvalRequired="37" approvalRequired_for_taxonomicNames="8" approvalRequired_for_treatments="29" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FFCA6A75FF8B9D6A142BBD49" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Nyctophilus heran Kitchener, How &amp; Maharadatunkamsi 1991" docType="treatment" docVersion="10" lastPageNumber="807" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="807" updateTime="1658412944878" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Vespertilionidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>716</mods:start>
<mods:end>981</mods:end>
</mods:extent>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6397752</mods:identifier>
<mods:identifier type="GBIF-Dataset">45351c32-25dd-422c-bdb2-00e73deb4943</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">6397752</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6578437" ID-GBIF-Taxon="195628043" ID-Zenodo-Dep="6578437" LSID="urn:lsid:plazi:treatment:4C3D87E8FFCA6A75FF8B9D6A142BBD49" httpUri="http://treatment.plazi.org/id/4C3D87E8FFCA6A75FF8B9D6A142BBD49" lastPageNumber="807" pageId="49" pageNumber="807">
<subSubSection box="[119,176,2867,2917]" pageId="49" pageNumber="807" type="multiple">
<paragraph blockId="49.[117,1122,2867,2997]" box="[119,176,2867,2917]" pageId="49" pageNumber="807">
<heading box="[119,176,2867,2917]" pageId="49" pageNumber="807">
<figureCitation box="[119,176,2867,2917]" captionStart="Plate 58: Vespertilionidae" captionStartId="43.[119,149,3359,3384]" captionTargetBox="[11,2759,17,3657]" captionTargetPageId="42" captionText="78. New Guinea Big-eared Bat (Pharotis imogene), 79. Western Long-eared Bat (Nyctophilus major), 80. Corbens Long-eared Bat (Nyctophilus corbeni), 81. Pallid Long-eared Bat (Nyctophilus daedalus), 82. Goulds Long-eared Bat (Nyctophilus gouldi), 83. Tasmanian Long-eared Bat (Nyctophilus sherrini), 84. New Caledonian Long-eared Bat (Nyctophilus nebulosus), 85. Papuan Long-eared Bat (Nyctophilus macrotis), 86. Pygmy Long-eared Bat (Nyctophilus walkeri), 87. Eastern Long-eared Bat (Nyctophilus bifax), 88. Northern Long-eared Bat (Nyctophilus arnhemensis), 89. Mount Missim Long-eared Bat (Nyctophilus shirleyae), 90. Small-toothed Long-eared Bat (Nyctophilus microdon), 91. Lesser Long-eared Bat (Nyctophilus geoffroyi), 92. Sunda Long-eared Bat (Nyctophilus heran), 93. Lord Howe Long-eared Bat (Nyctophilus howensis)" figureDoi="http://doi.org/10.5281/zenodo.6398510" httpUri="https://zenodo.org/record/6398510/files/figure.png" pageId="49" pageNumber="807">92.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[194,663,2867,2917]" pageId="49" pageNumber="807" type="vernacular_names">
<paragraph blockId="49.[117,1122,2867,2997]" box="[194,663,2867,2917]" pageId="49" pageNumber="807">
<heading box="[194,663,2867,2917]" pageId="49" pageNumber="807">
<vernacularName box="[194,663,2867,2917]" pageId="49" pageNumber="807">Sunda Long-eared Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[731,1051,2867,2917]" pageId="49" pageNumber="807" type="nomenclature">
<paragraph blockId="49.[117,1122,2867,2997]" box="[731,1051,2867,2917]" pageId="49" pageNumber="807">
<heading box="[731,1051,2867,2917]" pageId="49" pageNumber="807">
<taxonomicName authorityName="Kitchener, How &amp; Maharadatunkamsi" authorityYear="1991" box="[731,1051,2867,2917]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="heran">
<emphasis box="[731,1051,2867,2917]" italics="true" pageId="49" pageNumber="807">Nyctophilus heran</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="vernacular_names">
<paragraph blockId="49.[117,1122,2867,2997]" box="[119,1120,2935,2956]" pageId="49" pageNumber="807">
<heading box="[119,1120,2935,2956]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[119,195,2935,2956]" pageId="49" pageNumber="807">French:</emphasis>
<vernacularName box="[204,434,2935,2956]" pageId="49" pageNumber="807">Nyctophile de la Sonde</vernacularName>
/
<emphasis bold="true" box="[455,545,2935,2956]" pageId="49" pageNumber="807">German:</emphasis>
<vernacularName box="[555,818,2935,2956]" pageId="49" pageNumber="807">Sunda-Langohrfledermaus</vernacularName>
/
<emphasis bold="true" box="[839,930,2935,2956]" pageId="49" pageNumber="807">Spanish:</emphasis>
<vernacularName box="[940,1120,2935,2956]" pageId="49" pageNumber="807">Nictofila de Sonda</vernacularName>
</heading>
</paragraph>
<paragraph blockId="49.[117,1122,2867,2997]" box="[118,609,2974,2995]" pageId="49" pageNumber="807">
<heading box="[118,609,2974,2995]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[118,366,2974,2995]" pageId="49" pageNumber="807">Other common names:</emphasis>
<vernacularName box="[375,609,2974,2995]" pageId="49" pageNumber="807">Sundan Long-eared Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="reference_group">
<paragraph blockId="49.[730,1324,3047,3470]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[731,881,3047,3076]" pageId="49" pageNumber="807">Taxonomy.</emphasis>
<taxonomicName authority="Kitchener, How &amp; Maharadatunkamsi, 1991" authorityName="Kitchener, How &amp; Maharadatunkamsi" authorityYear="1991" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="heran">Nyctophilus heran Kitchener, How &amp; Maharadatunkamsi, 1991</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="materials_examined">
<paragraph blockId="49.[730,1324,3047,3470]" pageId="49" pageNumber="807">
<materialsCitation ID-GBIF-Occurrence="3783319328" latitude="-8.366667" longitude="31.0" pageId="49" pageNumber="807">
“Kampong Merdeka, Desa Hadakewa, Pulau Lembata,
<collectingRegion box="[731,933,3161,3194]" country="Indonesia" name="Nusa Tenggara Timur" pageId="49" pageNumber="807">Nusa Tenggara</collectingRegion>
[=
<collectingRegion box="[978,1264,3161,3194]" country="Indonesia" name="Nusa Tenggara Timur" pageId="49" pageNumber="807">Lesser Sunda Islands</collectingRegion>
],
<collectingCountry name="Indonesia" pageId="49" pageNumber="807">Indonesia</collectingCountry>
(
<geoCoordinate box="[856,959,3204,3233]" degrees="08" direction="south" minutes="22" orientation="latitude" pageId="49" pageNumber="807" precision="925" value="-8.366667">08°22'S</geoCoordinate>
,
<geoCoordinate box="[977,1096,3204,3233]" degrees="123" direction="east" minutes="31" orientation="longitude" pageId="49" pageNumber="807" precision="55555" value="31.0">123°31'E</geoCoordinate>
) ... at sea level.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="discussion">
<paragraph blockId="49.[730,1324,3047,3470]" lastBlockId="49.[1398,2609,284,1663]" pageId="49" pageNumber="807">
Placement of
<taxonomicName authorityName="Kitchener, How &amp; Maharadatunkamsi" authorityYear="1991" box="[926,1152,3240,3273]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="heran">Nyctophilus heran</taxonomicName>
in a species group is uncertain. It is most similar to
<taxonomicName authorityName="Leach" authorityYear="1821" box="[731,870,3319,3352]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="geoffroyi">N. geoffroyi</taxonomicName>
and is tentatively included in the
<taxonomicName authorityName="Leach" authorityYear="1821" box="[730,831,3358,3391]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="geoffroyi">geoffroyi</taxonomicName>
species group. There were apparently recently obtained specimens of
<taxonomicName authorityName="Leach" authorityYear="1821" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="genus">Nyctophilus</taxonomicName>
on Timor that apparently resembled
<taxonomicName authorityName="Kitchener, How &amp; Maharadatunkamsi" authorityYear="1991" box="[1475,1589,284,317]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="heran">N. heran</taxonomicName>
, but a comparison of these samples to other
<taxonomicName authorityName="Leach" authorityYear="1821" box="[2224,2369,284,317]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="genus">Nyctophilus</taxonomicName>
has not yet been undertaken, and the name
<taxonomicName baseAuthorityName="E. Geoffroy" baseAuthorityYear="1806" box="[1784,1914,324,357]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="timoriensis">timoriensis</taxonomicName>
will continue to go unused for now. Monotypic.
</paragraph>
</subSubSection>
<subSubSection box="[1403,2452,363,396]" pageId="49" pageNumber="807" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6397962" ID-Zenodo-Dep="6397962" box="[1403,2452,363,396]" httpUri="https://zenodo.org/record/6397962/files/figure.png" inLine="true" pageId="49" pageNumber="807" targetBox="[114,706,3047,3461]" targetPageId="49">
<paragraph blockId="49.[1398,2609,284,1663]" box="[1403,2452,363,396]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1403,1579,363,396]" pageId="49" pageNumber="807">Distribution.</emphasis>
Known only from type locality on Lembata I, Lesser Sunda Is.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="description">
<paragraph blockId="49.[1398,2609,284,1663]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1403,1647,407,436]" pageId="49" pageNumber="807">Descriptive notes.</emphasis>
Head-body 51-5 mm, tail 40-7 mm, ear 23-4 mm, hindfoot 6-4 mm, forearm 39-3 mm; weight 7-6 g (type specimen). The Sunda Long-eared Bat has very large ears and unique simple noseleaf consisting of two ridges, one further on muzzle and another immediately above nostrils, with vertical groove in middle and furred trough between them. Dorsal pelage is light drab brown; venter is paler. Face, ears, and wing membranes are light buff. Rostrum is short and blunt, with ridge across muzzle over nostrils that is well developed and high, with paired mounds joined medially by a conspicuous membrane that expands distally to form a Y-shaped groove. Ears are very large and broad, with bluntly roundedtip, horizontal ribbing on inner surfaces, inward curved anterior edges, and smooth posterior edges (ears can fold back at top ofthick part of anterior edge); large and furred interauricular band crosses forehead between ears; tragus is moderately long and bluntly rounded at tip, being practically straight on anterior margin. Glans penisis slender and lacks very prominent distal lobe; urethral lobe is ventrally U-shaped;it differs from the Lesser Long-eared Bat (
<taxonomicName authorityName="Leach" authorityYear="1821" box="[2129,2271,915,948]" class="Mammalia" family="Vespertilionidae" genus="Nyctophilus" kingdom="Animalia" order="Chiroptera" pageId="49" pageNumber="807" phylum="Chordata" rank="species" species="geoffroyi">N. geoffroyi</taxonomicName>
) in lacking pronounced dorsal crest and distal lobe that overshoots ventral mound. Baculum has moderately thin shaft in dorsal view, tapering to non-bifurcated tip, base being strongly bifurcated; in lateral view, baculum is curved downward at base, but shaftis straight to pointed tip. Skull is generally similar to the Lesser Long-eared Bat,sagittal crest is weak, rostrum is slightly more depressed and median sulcusis slightly deeper than in the Lesser Long-eared Bat, tympanic bullae are relatively inflated, and M” and M, are moderately reduced.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="biology_ecology">
<paragraph blockId="49.[1398,2609,284,1663]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1400,1512,1192,1225]" pageId="49" pageNumber="807">Habitat.</emphasis>
Type specimen was collected over a small stagnant poolin a small dry stream
<quantity box="[1401,1486,1232,1265]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="807" unit="m" value="200.0">200 m</quantity>
from mangrovesat sea level. Native gardens and occasionally banana and coconut trees bordered the stream.
</paragraph>
</subSubSection>
<subSubSection box="[1400,1901,1309,1342]" pageId="49" pageNumber="807" type="food_feeding">
<paragraph blockId="49.[1398,2609,284,1663]" box="[1400,1901,1309,1342]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1400,1662,1309,1342]" pageId="49" pageNumber="807">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection box="[1400,1774,1348,1381]" pageId="49" pageNumber="807" type="breeding">
<paragraph blockId="49.[1398,2609,284,1663]" box="[1400,1774,1348,1381]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1400,1535,1348,1381]" pageId="49" pageNumber="807">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection box="[1399,1873,1388,1421]" pageId="49" pageNumber="807" type="activity">
<paragraph blockId="49.[1398,2609,284,1663]" box="[1399,1873,1388,1421]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1399,1634,1388,1421]" pageId="49" pageNumber="807">Activity patterns.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="biology_ecology">
<paragraph blockId="49.[1398,2609,284,1663]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1400,2108,1428,1461]" pageId="49" pageNumber="807">Movements, Home range and Social organization.</emphasis>
The Sunda Long-eared Bat apparently forms small colonies.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="807" type="conservation">
<paragraph blockId="49.[1398,2609,284,1663]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1400,1747,1507,1540]" pageId="49" pageNumber="807">Status and Conservation.</emphasis>
Classified as Data Deficient on The IUCN Red List. The Sunda Long-eared Bat is currently known only from the type locality on Lembata Island. It is likely threatened given its apparent rarity and restricted distribution.
</paragraph>
</subSubSection>
<subSubSection box="[1400,2603,1634,1659]" pageId="49" pageNumber="807" type="bibRefCitation_list">
<paragraph blockId="49.[1398,2609,284,1663]" box="[1400,2603,1634,1659]" pageId="49" pageNumber="807">
<emphasis bold="true" box="[1400,1552,1634,1659]" pageId="49" pageNumber="807">Bibliography.</emphasis>
Corbet &amp; Hill (1992), Hutson,
<collectingRegion box="[1876,1967,1634,1659]" country="Jamaica" name="Kingston" pageId="49" pageNumber="807">Kingston</collectingRegion>
et al. (2016), Kitchener et al. (1991), Parnaby (2002a, 2009).
</paragraph>
</subSubSection>
</treatment>
</document>