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<document ID-DOI="http://doi.org/10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" approvalRequired="37" approvalRequired_for_taxonomicNames="8" approvalRequired_for_treatments="29" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FF716ACFFA519FCD1DFEBFB9" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Kerivoula titania Bates 2007" docType="treatment" docVersion="11" lastPageNumber="899" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="898" updateTime="1658433834459" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Vespertilionidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>716</mods:start>
<mods:end>981</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6397752</mods:identifier>
<mods:identifier type="GBIF-Dataset">45351c32-25dd-422c-bdb2-00e73deb4943</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">6397752</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6580638" ID-GBIF-Taxon="195628329" ID-Zenodo-Dep="6580638" LSID="urn:lsid:plazi:treatment:4C3D87E8FF716ACFFA519FCD1DFEBFB9" httpUri="http://treatment.plazi.org/id/4C3D87E8FF716ACFFA519FCD1DFEBFB9" lastPageId="139" lastPageNumber="899" pageId="138" pageNumber="898">
<subSubSection box="[1453,1535,2452,2498]" pageId="138" pageNumber="898" type="multiple">
<paragraph blockId="138.[1451,2341,2452,2538]" box="[1453,1535,2452,2498]" pageId="138" pageNumber="898">
<heading box="[1453,1535,2452,2498]" pageId="138" pageNumber="898">
<figureCitation box="[1453,1535,2452,2498]" captionStart="Plate 67: Vespertilionidae" captionStartId="135.[114,144,3260,3285]" captionTargetBox="[11,2751,19,3659]" captionTargetPageId="134" captionText="293. Groove-toothed Trumpet-eared Bat (Phoniscus atrox), 294. Peterss Trumpet-eared Bat (Phoniscus jagorii), 295. Dubious Trumpet-eared Bat (Phoniscus aerosus), 296. Golden-tipped Bat (Phonuscus papuensis), 297. Hardwickes Woolly Bat (Kerivoula hardwickii), 298. Flatskulled Woolly Bat (Kerivoula depressa), 299. Sri Lankan Woolly Bat (Kerivoula malpasi), 300. Cryptic Woolly Bat (Kerivoula crypta), 301. Dark Woolly Bat (Kerivoula furva), 302. Kachin Woolly Bat (Kerivoula kachinensis), 303. Indochinese Woolly Bat (Kerivoula dongduongana), 304. Krau Woolly Bat (Kerwoula krauensis), 305. Titanias Woolly Bat (Kerwoula titania), 306. Least Woolly Bat (Kerivoula minuta), 307. Small Woolly Bat (Kerivoula intermedia), 308. Papillose Woolly Bat (Kerivoula papillosa), 309. Lenis Woolly Bat (Kerivoula lenis), 310. Clear-winged Woolly Bat (Kerivoula pellucida), 311. Painted Woolly Bat (Kerwoula picta), 312. Whiteheads Woolly Bat (Kerivoula whiteheadi), 313. Flores Woolly Bat (Kerivoula flora), 314. Fly River Woolly Bat (Kerivoula muscina), 315. Saint Aignans Woolly Bat (Kerivoula agnella), 316. Bismarck Woolly Bat (Kerwoula myrella), 317. Lesser Woolly Bat (Kerivoula lanosa), 318. Ethiopian Woolly Bat (Kerwoula eriophora), 319. Tanzanian Woolly Bat (Kerwwoula africana), 320. Smiths Woolly Bat (Kerivoula smithii), 321. Coppery Woolly Bat (Kerivoula cuprosa), 322. Spurrells Woolly Bat (Kerivoula phalaena), 323. Damara Woolly Bat (Kerwoula argentata)" figureDoi="http://doi.org/10.5281/zenodo.6398545" httpUri="https://zenodo.org/record/6398545/files/figure.png" pageId="138" pageNumber="898">305.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1552,1965,2452,2498]" pageId="138" pageNumber="898" type="vernacular_names">
<paragraph blockId="138.[1451,2341,2452,2538]" box="[1552,1965,2452,2498]" pageId="138" pageNumber="898">
<heading box="[1552,1965,2452,2498]" pageId="138" pageNumber="898">
<vernacularName box="[1552,1965,2452,2498]" pageId="138" pageNumber="898">Titanias Woolly Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[2036,2341,2452,2498]" pageId="138" pageNumber="898" type="nomenclature">
<paragraph blockId="138.[1451,2341,2452,2538]" box="[2036,2341,2452,2498]" pageId="138" pageNumber="898">
<heading box="[2036,2341,2452,2498]" pageId="138" pageNumber="898">
<taxonomicName authority="Bates et al., 2007" authorityName="Bates" authorityYear="2007" box="[2036,2341,2452,2498]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="titania">
<emphasis box="[2036,2341,2452,2498]" italics="true" pageId="138" pageNumber="898">Kerivoula titania</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1453,2336,2516,2537]" pageId="138" pageNumber="898" type="vernacular_names">
<paragraph blockId="138.[1451,2341,2452,2538]" box="[1453,2336,2516,2537]" pageId="138" pageNumber="898">
<heading box="[1453,2336,2516,2537]" pageId="138" pageNumber="898">
<emphasis bold="true" box="[1453,1528,2516,2537]" pageId="138" pageNumber="898">French:</emphasis>
<vernacularName box="[1538,1695,2516,2537]" pageId="138" pageNumber="898">Kérivoule titania</vernacularName>
/
<emphasis bold="true" box="[1716,1807,2516,2537]" pageId="138" pageNumber="898">German:</emphasis>
<vernacularName box="[1812,2041,2516,2537]" pageId="138" pageNumber="898">Titania-Wollfledermaus</vernacularName>
/
<emphasis bold="true" box="[2062,2153,2516,2537]" pageId="138" pageNumber="898">Spanish:</emphasis>
<vernacularName box="[2163,2269,2516,2537]" pageId="138" pageNumber="898">Querivoula</vernacularName>
titania
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="898" type="reference_group">
<paragraph blockId="138.[2064,2659,2585,3009]" pageId="138" pageNumber="898">
<emphasis bold="true" box="[2064,2216,2585,2614]" pageId="138" pageNumber="898">Taxonomy.</emphasis>
<taxonomicName authority="Bates et al., 2007" authorityName="Bates" authorityYear="2007" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="titania">Kerivoula titania Bates et al., 2007</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="898" type="materials_examined">
<paragraph blockId="138.[2064,2659,2585,3009]" pageId="138" pageNumber="898">
<materialsCitation ID-GBIF-Occurrence="3783319635" latitude="10.49" longitude="106.97583" pageId="138" pageNumber="898">
“Seima Biodiversity Conservation Area,
<collectingRegion box="[2158,2479,2669,2694]" country="Cambodia" name="Mondolkiri" pageId="138" pageNumber="898">Mondul Kiri Province</collectingRegion>
,
<collectingCountry box="[2506,2649,2669,2694]" name="Cambodia" pageId="138" pageNumber="898">Cambodia</collectingCountry>
,
<geoCoordinate box="[2068,2219,2708,2733]" degrees="12" direction="north" minutes="10.49" orientation="latitude" pageId="138" pageNumber="898" precision="555" value="10.49">12°10.49°N</geoCoordinate>
,
<geoCoordinate box="[2238,2401,2708,2733]" degrees="106" direction="east" minutes="58.55" orientation="longitude" pageId="138" pageNumber="898" precision="9" value="106.97583">106°58.55'E</geoCoordinate>
, 290. m asl.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="898" type="discussion">
<paragraph blockId="138.[2064,2659,2585,3009]" pageId="138" pageNumber="898">
<taxonomicName box="[2066,2292,2747,2772]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="titania">Kerivoula titania</taxonomicName>
appears to be sister to
<taxonomicName authorityName="G. S. Miller" authorityYear="1898" box="[2066,2197,2782,2811]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="minuta">K. minuta</taxonomicName>
, and this clade is sister to the rest of the
<taxonomicName authorityName="Horsfield" authorityYear="1824" box="[2170,2299,2826,2851]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="hardwickii">hardwickii</taxonomicName>
species group, although it might also be related to
<taxonomicName authorityName="Hill &amp; Francis" authorityYear="1984" box="[2474,2651,2858,2891]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="intermedia">K. intermedia</taxonomicName>
. Specimens previously attributed to K. fitania from Hainan and
<collectingCountry box="[2384,2480,2936,2969]" name="Taiwan" pageId="138" pageNumber="898">Taiwan</collectingCountry>
islands now represent
<taxonomicName box="[2207,2314,2980,3009]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="138" pageNumber="898" phylum="Chordata" rank="species" species="furva">K. furva</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection box="[1453,2456,3025,3050]" pageId="138" pageNumber="898" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6398458" ID-Zenodo-Dep="6398458" box="[1453,2456,3025,3050]" httpUri="https://zenodo.org/record/6398458/files/figure.png" inLine="true" pageId="138" pageNumber="898" targetBox="[1450,2041,2590,3004]" targetPageId="138">
<paragraph blockId="138.[1453,2659,3025,3484]" box="[1453,2456,3025,3050]" pageId="138" pageNumber="898">
<emphasis bold="true" box="[1453,1629,3025,3050]" pageId="138" pageNumber="898">Distribution.</emphasis>
E
<collectingCountry box="[1670,1801,3025,3050]" name="Myanmar" pageId="138" pageNumber="898">Myanmar</collectingCountry>
, N
<collectingCountry box="[1850,1979,3025,3050]" name="Thailand" pageId="138" pageNumber="898">Thailand</collectingCountry>
,
<collectingCountry box="[1997,2062,3025,3050]" name="Laos" pageId="138" pageNumber="898">Laos</collectingCountry>
,
<collectingCountry box="[2077,2199,3025,3050]" name="Vietnam" pageId="138" pageNumber="898">Vietnam</collectingCountry>
, and E
<collectingCountry box="[2306,2452,3025,3050]" name="Cambodia" pageId="138" pageNumber="898">Cambodia</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="139" lastPageNumber="899" pageId="138" pageNumber="898" type="description">
<paragraph blockId="138.[1453,2659,3025,3484]" lastBlockId="139.[135,1345,302,1165]" lastPageId="139" lastPageNumber="899" pageId="138" pageNumber="898">
<emphasis bold="true" box="[1454,1698,3055,3088]" pageId="138" pageNumber="898">Descriptive notes.</emphasis>
Head—body 38-
<quantity box="[1915,2054,3055,3088]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="4.1" metricValueMin="0.4" pageId="138" pageNumber="898" unit="mm" value="22.5" valueMax="41.0" valueMin="4.0">4—41 mm</quantity>
, tail 45-8-53-
<quantity box="[2240,2324,3055,3088]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="138" pageNumber="898" unit="mm" value="5.0">5 mm</quantity>
, ear 12-
<quantity box="[2441,2568,3055,3088]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="138" pageNumber="898" unit="mm" value="10.0" valueMax="15.0" valueMin="5.0">5-15 mm</quantity>
, hindfoot 7-1-8-
<quantity box="[1600,1683,3095,3128]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="138" pageNumber="898" unit="mm" value="3.0">3 mm</quantity>
, forearm 32-4-35-
<quantity box="[1934,2016,3095,3128]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="138" pageNumber="898" unit="mm" value="9.0">9 mm</quantity>
; weight 4-7-
<quantity box="[2185,2229,3095,3128]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="138" pageNumber="898" unit="g" value="9.0">9 g</quantity>
. Dorsal pelage is brownish gray (hairs with black bases, light gray middles, and dark gray tips); venteris a little lighter (hairs black with whitish tips tinged with gray-brown). Face and lips are hairy, except nostrils. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragusis narrow and tall, with virtually straight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are semi-translucent and gray, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes. Fleshy lobe at base of thumb is oval and smooth in middle. Skull has broad, flattened braincase; rostrum has large narial pit and well developed rostral sulcus; and basioccipital pits are shallow. I? is unicuspid, with cingulum on its internal posterior border; I is equal in crown area to I?, with well-defined internally positioned cusp; canines are smooth, with deep longitudinal groove on outer sides; upper premolars are compressed in tooth row; M? lacks metastyle; I and I, are tricuspid, and I, is smaller with only well-defined central cusp and no lateral cusps; lower premolars are compressed in tooth row; and talonid of M,is reduced to ¢.70% of trigonid size.
</paragraph>
</subSubSection>
<subSubSection pageId="139" pageNumber="899" type="biology_ecology">
<paragraph blockId="139.[135,1345,302,1165]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[139,250,539,572]" pageId="139" pageNumber="899">Habitat.</emphasis>
Lowland and montane primary and secondary evergreen and deciduous forests at elevations of
<quantity box="[417,585,581,610]" metricMagnitude="2" metricUnit="m" metricValue="9.3" metricValueMax="16.0" metricValueMin="2.6" pageId="139" pageNumber="899" unit="m" value="930.0" valueMax="1600.0" valueMin="260.0">260-1600 m</quantity>
(
<collectingCountry box="[608,735,581,610]" name="Vietnam" pageId="139" pageNumber="899">Vietnam</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection box="[139,639,616,649]" pageId="139" pageNumber="899" type="food_feeding">
<paragraph blockId="139.[135,1345,302,1165]" box="[139,639,616,649]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[139,400,616,649]" pageId="139" pageNumber="899">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="139" pageNumber="899" type="breeding">
<paragraph blockId="139.[135,1345,302,1165]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[138,272,657,690]" pageId="139" pageNumber="899">Breeding.</emphasis>
Female Titanias Woolly Bats in late pregnancy were caughtin early May in Ba Be National Park, northern
<collectingCountry box="[575,697,696,729]" name="Vietnam" pageId="139" pageNumber="899">Vietnam</collectingCountry>
, and late April in
<collectingCountry box="[950,1017,696,729]" name="Laos" pageId="139" pageNumber="899">Laos</collectingCountry>
. Two pregnant females were captured in mid-May and two lactating females in early June in
<collectingCountry box="[1112,1178,736,769]" name="Laos" pageId="139" pageNumber="899">Laos</collectingCountry>
, suggesting births occur in late May and possibly earlyJune.
</paragraph>
</subSubSection>
<subSubSection pageId="139" pageNumber="899" type="activity">
<paragraph blockId="139.[135,1345,302,1165]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[136,373,819,848]" pageId="139" pageNumber="899">Activity patterns.</emphasis>
In
<collectingCountry box="[428,557,819,848]" name="Thailand" pageId="139" pageNumber="899">Thailand</collectingCountry>
, call shape is steep FM sweep, with start frequencies of 217-243 kHz (mean 227 kHz), end frequencies of 74-8-90 kHz (82-6 kHz), and peak frequencies of 137-7-168-7 kHz (157 kHz).
</paragraph>
</subSubSection>
<subSubSection box="[137,1076,932,965]" pageId="139" pageNumber="899" type="biology_ecology">
<paragraph blockId="139.[135,1345,302,1165]" box="[137,1076,932,965]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[137,837,932,965]" pageId="139" pageNumber="899">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="139" pageNumber="899" type="conservation">
<paragraph blockId="139.[135,1345,302,1165]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[137,490,976,1005]" pageId="139" pageNumber="899">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Titania's Woolly Bat does not seem to currently face any major threats, but becauseit is forest dependent, logging and agricultural development might be future threats.
</paragraph>
</subSubSection>
<subSubSection pageId="139" pageNumber="899" type="bibRefCitation_list">
<paragraph blockId="139.[135,1345,302,1165]" pageId="139" pageNumber="899">
<emphasis bold="true" box="[137,290,1099,1124]" pageId="139" pageNumber="899">Bibliography.</emphasis>
Bates, Struebig et al. (2007), Chanson (2008), Douangboubpha et al. (2016), Kruskop (2013a), Kuo Haochih et al. (2017), Tu Vuong Tan et al. (2018), Wu Yi, Li Yuchun et al. (2012).
</paragraph>
</subSubSection>
</treatment>
</document>