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<document ID-DOI="http://doi.org/10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" approvalRequired="5" approvalRequired_for_matCits="1" approvalRequired_for_treatments="4" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E2753527FE96E183296D73628F1B" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Maxomys hellwaldii" docType="treatment" docVersion="5" lastPageNumber="862" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="862" updateTime="1658538965639" updateUser="ExternalLinkService">
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<mods:title>Muridae</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Thomas E. Lacher, Jr</mods:namePart>
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<mods:dateIssued>2017</mods:dateIssued>
<mods:dateOther type="pubDate">2017-11-30</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
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<mods:placeTerm>Barcelona</mods:placeTerm>
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<mods:title>Handbook of the Mammals of the World Volume 7 Rodents II</mods:title>
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<mods:start>536</mods:start>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6887260</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6833357" ID-GBIF-Taxon="197825624" ID-Zenodo-Dep="6833357" LSID="urn:lsid:plazi:treatment:1E30E2753527FE96E183296D73628F1B" httpUri="http://treatment.plazi.org/id/1E30E2753527FE96E183296D73628F1B" lastPageNumber="862" pageId="257" pageNumber="862">
<heading pageId="257" pageNumber="862">
<subSubSection box="[180,262,1380,1426]" pageId="257" pageNumber="862" type="multiple">
<paragraph blockId="257.[178,1317,1380,1504]" box="[180,262,1380,1426]" pageId="257" pageNumber="862">
<figureCitation box="[180,262,1380,1426]" captionStart="Plate 56: Muridae" captionStartId="248.[116,147,3133,3158]" captionTargetBox="[14,2745,12,3649]" captionTargetPageId="247" captionText="724. Burmese Tree Rat (Chiromyscus chiropus), 725. Langbian Tree Rat (Chiromyscus langbianis), 726. Thomas's Tree Rat (Chiromyscus thomasi), 727. Millets Long-tailed Giant Rat (Leopoldamys milleti), 728. Edwardss Long-tailed Giant Rat (Leopoldamys edwardsi), 729. Indomalayan Long-tailed Giant Rat (Leopoldamys sabanus), 730. Herbert's Long-tailed Giant Rat (Leopoldamys herberti), 731. Neills Long-tailed Giant Rat (Leopoldamys neilli), 733. Diwangkaras Long-tailed Giant Rat (Leopoldamys diwangkarai), 734. Mentawai Archipelago Long-tailed Giant Rat (Leopoldamys siporanus), 735. Paulinas Limestone Rat (Saxatilomys paulinae), 736. Daovantiens Limestone Rat (Tonkinomys daovantieni), 737. Javan Bamboo Rat (Kadarsanomys sodyi), 739. Indomalayan Spiny Rat (Maxomys surifer), 740. Indochinese Mountain Spiny Rat (Maxomys moi), 741. Bornean Mountain Spiny Rat (Maxomys alticola), 742. Small Bornean Spiny Rat (Maxomys baeodon), 743. Bartelss Javan Spiny Rat (Maxomys bartelsii), 744. Sumatran Mountain Spiny Rat (Maxomys hylomyoides), 745. Malayan Mountain Spiny Rat (Maxomys inas), 746. Broad-nosed Sumatran Spiny Rat (Maxomys inflatus), 747. Ochraceous-bellied Bornean Spiny Rat (Maxomys ochracerventer), 748. Mentawai Archipelago Spiny Rat (Maxomys pagensis), 749. Palawan Spiny Rat (Maxomys panglima), 750. Rajah Sundaic Spiny Rat (Maxomys rajah), 751. Whiteheads Sundaic Spiny Rat (Maxomys whiteheadi), 752. Dollmans Sulawesi Spiny Rat (Maxomys dollmani), 753. Hellwalds Sulawesi Spiny Rat (Maxomys hellwaldii), 755. Tajuddins Spiny Rat (Maxomys tajuddinii)" figureDoi="http://doi.org/10.5281/zenodo.6888489" httpUri="https://zenodo.org/record/6888489/files/figure.png" pageId="257" pageNumber="862">753.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[279,892,1380,1426]" pageId="257" pageNumber="862" type="vernacular_names">
<paragraph blockId="257.[178,1317,1380,1504]" box="[279,892,1380,1426]" pageId="257" pageNumber="862">
<vernacularName box="[279,892,1380,1426]" pageId="257" pageNumber="862">Hellwalds Sulawesi Spiny Rat</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[942,1317,1380,1426]" pageId="257" pageNumber="862" type="nomenclature">
<paragraph blockId="257.[178,1317,1380,1504]" box="[942,1317,1380,1426]" pageId="257" pageNumber="862">
<taxonomicName baseAuthorityName="Jentink" baseAuthorityYear="1878" box="[942,1317,1380,1426]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="species" species="hellwaldii">
<emphasis box="[942,1317,1380,1426]" italics="true" pageId="257" pageNumber="862">Maxomys hellwaldii</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="vernacular_names">
<paragraph blockId="257.[178,1317,1380,1504]" box="[178,1317,1444,1465]" pageId="257" pageNumber="862">
<emphasis bold="true" box="[178,254,1444,1465]" pageId="257" pageNumber="862">French:</emphasis>
<vernacularName box="[264,484,1444,1465]" pageId="257" pageNumber="862">Maxomys de Hellwald</vernacularName>
/
<emphasis bold="true" box="[505,596,1444,1465]" pageId="257" pageNumber="862">German:</emphasis>
<vernacularName box="[606,811,1444,1465]" pageId="257" pageNumber="862">Hellwald-Rajah-Ratte</vernacularName>
/
<emphasis bold="true" box="[832,923,1444,1465]" pageId="257" pageNumber="862">Spanish:</emphasis>
<vernacularName box="[934,1317,1444,1465]" pageId="257" pageNumber="862">Rata espinosa de Célebes de Hellwald</vernacularName>
</paragraph>
<paragraph blockId="257.[178,1317,1380,1504]" box="[178,954,1483,1504]" pageId="257" pageNumber="862">
<emphasis bold="true" box="[178,424,1483,1504]" pageId="257" pageNumber="862">Other common names:</emphasis>
<vernacularName box="[435,638,1483,1504]" pageId="257" pageNumber="862">Hellwald's Spiny Rat</vernacularName>
,
<vernacularName box="[652,954,1483,1504]" pageId="257" pageNumber="862">Hellwald's Sulawesi Maxomys</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection box="[791,1380,1550,1584]" pageId="257" pageNumber="862" type="reference_group">
<paragraph blockId="257.[790,1383,1550,1978]" box="[791,1380,1550,1584]" pageId="257" pageNumber="862">
Taxonomy.
<taxonomicName authority="Jentink, 1879" authorityName="Jentink" authorityYear="1879" box="[967,1375,1550,1584]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="species" species="hellwaldii">Mus hellwaldii Jentink, 1879</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="materials_examined">
<paragraph blockId="257.[790,1383,1550,1978]" pageId="257" pageNumber="862">
<materialsCitation ID-GBIF-Occurrence="3864424717" pageId="257" pageNumber="862">“Celebes [= Sulawesi, Indonesia], Me-nado.”</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="discussion">
<paragraph blockId="257.[790,1383,1550,1978]" pageId="257" pageNumber="862">
<taxonomicName baseAuthorityName="Jentink" baseAuthorityYear="1878" box="[792,1050,1663,1701]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="species" species="hellwaldii">Maxomys hellwaldii</taxonomicName>
is most closely related to
<taxonomicName baseAuthorityName="Ellerman" baseAuthorityYear="1941" box="[833,997,1708,1742]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="species" species="dollmani">M. dollmani</taxonomicName>
, forming a clade together with at least three undescribed species, which is sister to
<taxonomicName box="[1050,1177,1791,1820]" class="Mammalia" family="Muridae" genus="Crunomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="genus">Crunomys</taxonomicName>
.
<taxonomicName box="[1198,1318,1791,1820]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="genus">Maxomys</taxonomicName>
hell- waldn might also represent multiple species because there is a large amount of ge- ographical variation between populations. Taxonomy requires further investigation.
</paragraph>
</subSubSection>
<subSubSection box="[176,486,1985,2014]" pageId="257" pageNumber="862" type="distribution">
<paragraph blockId="257.[171,1381,1985,3080]" box="[176,486,1985,2014]" pageId="257" pageNumber="862">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6888525" ID-Zenodo-Dep="6888525" box="[176,482,1985,2014]" httpUri="https://zenodo.org/record/6888525/files/figure.png" inLine="true" pageId="257" pageNumber="862" startId="257.[176,348,1985,2014]" targetBox="[176,766,1559,1972]" targetPageId="257">Distribution. Sulawesi</caption>
.
</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="description">
<paragraph blockId="257.[171,1381,1985,3080]" pageId="257" pageNumber="862">
Descriptive notes. Head-body 180-220 mm, tail 160-200 mm, ear 2630 mm, hindfoot 43-49-9 mm; weight 150-270 g. Hellwalds Sulawesi Spiny Rat is large, with short, dense, and soft pelage and long bristle-like guard hairs that are gray basally and brown distally. Dorsum is very dark brownish orange along back and top of head, becoming lighter orange brown on sides with whitish gray cheeks. Venteris white, sometimes suf- fused with gray on thorax and neck. Feet are pale yellow and sparsely haired. Ears are rounded; vibrissae are long and black. Tail is 87-89% of head-body length, sparsely haired, and grayish brown above and unpigmented below and for distal one-third. Skull is larger, more strongly built, and more angular than in Dollmans Sulawesi Spiny Rat (
<taxonomicName baseAuthorityName="Ellerman" baseAuthorityYear="1941" box="[330,498,2385,2410]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="species" species="dollmani">M. dollmani</taxonomicName>
). The mite Laelaps, fleas Nestivalius sulawesiensis and Musserella, and louse Hoplopleura sembeli have been recorded from Hellwalds Sulawesi Spiny Rat. There are four pairs of mammae: one pectoral, one post-axillary, one abdominal, and one inguinal. Chromosomal complement is 2n = 34, FN = 61 (males) and 62 (females).
</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="biology_ecology">
<paragraph blockId="257.[171,1381,1985,3080]" pageId="257" pageNumber="862">Habitat. Tropical lowland evergreen rain forests from sea level up to elevations of 1006 m.</paragraph>
<paragraph blockId="257.[171,1381,1985,3080]" pageId="257" pageNumber="862">Food and Feeding. Hellwalds Sulawesi Spiny Rat probably eats fruits, arthropods, snails, earthworms, and small vertebrates.</paragraph>
<paragraph blockId="257.[171,1381,1985,3080]" box="[173,1112,2691,2725]" pageId="257" pageNumber="862">Breeding. Litters of Hellwalds Sulawesi Spiny Rats have 2—4 young.</paragraph>
<paragraph blockId="257.[171,1381,1985,3080]" box="[171,1221,2731,2765]" pageId="257" pageNumber="862">Activity patterns. Hellwalds Sulawesi Spiny Rat is nocturnal and terrestrial.</paragraph>
<paragraph blockId="257.[171,1381,1985,3080]" box="[172,1112,2774,2803]" pageId="257" pageNumber="862">Movements, Home range and Social organization. No information.</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="conservation">
<paragraph blockId="257.[171,1381,1985,3080]" pageId="257" pageNumber="862">
Status and Conservation. Classified as Least Concern on The IUCN Red List. Hellwalds Sulawesi Spiny Rat has a relatively large distribution and occurs in Lore Lindu National Park and no doubt other protected areas. Habitat destruction and possibly hunting in the north-western peninsula are its only
<taxonomicName authorityName="Hatt" authorityYear="1934" box="[854,936,2933,2962]" class="Mammalia" family="Muridae" genus="Thamnomys" kingdom="Animalia" order="Rodentia" pageId="257" pageNumber="862" phylum="Chordata" rank="species" species="major">major</taxonomicName>
threats. Additional research is needed to fully understand its natural history, taxonomy, and threats.
</paragraph>
</subSubSection>
<subSubSection pageId="257" pageNumber="862" type="bibRefCitation_list">
<paragraph blockId="257.[171,1381,1985,3080]" pageId="257" pageNumber="862">Bibliography. Achmadi et al. (2013), Musser (1969c, 1991, 2014), Musser &amp; Carleton (2005), Musser &amp; Holden (1991), Musser &amp; Ruedas (2016b), Musser et al. (1979), Steppan &amp; Schenk (2017).</paragraph>
</subSubSection>
</treatment>
</document>