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<document ID-DOI="http://doi.org/10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" approvalRequired="5" approvalRequired_for_matCits="1" approvalRequired_for_treatments="4" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E27534CEFF7FE4512AFE7E8D8F94" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Rattus nikenii" docType="treatment" docVersion="8" lastPageNumber="837" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="837" updateTime="1658538965639" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Muridae</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Don E. Wilson</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Russell A. Mittermeier</mods:namePart>
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<mods:name type="personal">
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<mods:namePart>Thomas E. Lacher, Jr</mods:namePart>
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<mods:dateIssued>2017</mods:dateIssued>
<mods:dateOther type="pubDate">2017-11-30</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
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<mods:placeTerm>Barcelona</mods:placeTerm>
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<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World – Volume 7 Rodents II</mods:title>
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[MISSING], 624. [MISSING], 625. [MISSING], 626. [MISSING], 627. [MISSING], 628. [MISSING], 629. [MISSING], 630. [MISSING], 631. [MISSING], 632. [MISSING], 633. [MISSING], 634. [MISSING], 635. [MISSING], 636. [MISSING], 637. [MISSING], 638. [MISSING], 639. [MISSING], 640. [MISSING], 641. [MISSING], 642. [MISSING], 643. [MISSING], 644. [MISSING], 645. [MISSING], 646. [MISSING], 647. [MISSING], 648. [MISSING], 649. [MISSING], 650. [MISSING], 651. [MISSING], 652. [MISSING], 653. [MISSING], 654. [MISSING], 655. [MISSING], 656. [MISSING], 657. [MISSING], 659. Brown Rat (Rattus norvegicus), 660. Roof Rat (Rattus rattus), 661. [MISSING], 662. [MISSING], 664. Himalayan Rat (Rattus pyctoris), 665. Ranjini’s Rat (Rattus ranjiniae), 666. Sahyadris Forest Rat (Rattus satarae), 667. Andaman Archipelago Rat (Rattus stoicus), 669. Pacific Rat (Rattus exulans), 670. Indochinese Forest Rat (Rattus andamanensis), 672. Losea Rat (Rattus losea), 673. White-footed Indochinese Rat (Rattus nitidus), 674. Osgood’s Vietnamese Rat (Rattus osgoodi), 675. Little Indochinese Field Rat (Rattus sakeratensis), 676. Burnished Enggano Rat (Rattus adustus), 677. Kinabalu Rat (Rattus baluensis), 678. Aceh Rat (Rattus blangorum), 679. Enggano Island Rat (Rattus enganus), 680. Hoogerwert’s Sumatran Rat (Rattus hoogerwerfi), 682. Mentawai Archipelago Rat (Rattus lugens), 683. Gag Island Rat (Rattus nikenu), 684. Simalur Archipelago Rat (Rattus simalurensis), 685. Malaysian Field Rat (Rattus tiomanicus), 686. South-western Xanthurus Rat (Rattus bontanus), 687. Hoffmann’s Sulawesi Rat (Rattus hoffmanni), 688. Koopman' ’ s Peleng Island Rat (Rattus koopmani), 689. Marmoset Xanthurus Rat (Rattus marmosurus), 690. Lampobatang Sulawesi Rat (Rattus mollicomulus)" figureDoi="http://doi.org/10.5281/zenodo.6888389" httpUri="https://zenodo.org/record/6888389/files/figure.png" pageId="232" pageNumber="837">683.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[1482,1786,1783,1829]" pageId="232" pageNumber="837" type="vernacular_names">
<paragraph blockId="232.[1381,2108,1783,1908]" box="[1482,1786,1783,1829]" pageId="232" pageNumber="837">
<vernacularName box="[1482,1786,1783,1829]" pageId="232" pageNumber="837">Gag Island Rat</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[1837,2106,1783,1829]" pageId="232" pageNumber="837" type="nomenclature">
<paragraph blockId="232.[1381,2108,1783,1908]" box="[1837,2106,1783,1829]" pageId="232" pageNumber="837">
<taxonomicName baseAuthorityName="Maryanto et al." baseAuthorityYear="2010" box="[1837,2106,1783,1829]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="species" species="nikenii">
<emphasis box="[1837,2106,1783,1829]" italics="true" pageId="232" pageNumber="837">Rattus nikenii</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="vernacular_names">
<paragraph blockId="232.[1381,2108,1783,1908]" box="[1381,2108,1847,1868]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,1456,1847,1868]" pageId="232" pageNumber="837">French:</emphasis>
<vernacularName box="[1466,1592,1847,1868]" pageId="232" pageNumber="837">Rat de Niken</vernacularName>
/
<emphasis bold="true" box="[1614,1704,1847,1868]" pageId="232" pageNumber="837">German:</emphasis>
<vernacularName box="[1715,1865,1847,1868]" pageId="232" pageNumber="837">Insel-Gag-Ratte</vernacularName>
/
<emphasis bold="true" box="[1887,1977,1847,1868]" pageId="232" pageNumber="837">Spanish:</emphasis>
<vernacularName box="[1988,2108,1847,1868]" pageId="232" pageNumber="837">Rata de Gag</vernacularName>
</paragraph>
<paragraph blockId="232.[1381,2108,1783,1908]" box="[1381,1747,1887,1908]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,1627,1887,1908]" pageId="232" pageNumber="837">Other common names:</emphasis>
<vernacularName box="[1638,1747,1887,1908]" pageId="232" pageNumber="837">Niken's Rat</vernacularName>
</paragraph>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="reference_group">
<paragraph blockId="232.[1992,2585,1956,2380]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1992,2144,1956,1985]" pageId="232" pageNumber="837">Taxonomy.</emphasis>
<taxonomicName authorityName="Maryanto et al." authorityYear="2010" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="species" species="nikenii">Rattus nikenii Maryanto et al., 2010</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection box="[2075,2581,1992,2026]" pageId="232" pageNumber="837" type="materials_examined">
<materialsCitation ID-GBIF-Occurrence="3864424513" box="[2075,2581,1992,2026]" pageId="232" pageNumber="837">
<paragraph blockId="232.[1992,2585,1956,2380]" box="[2075,2241,1992,2026]" pageId="232" pageNumber="837">“Gag Island,</paragraph>
<paragraph blockId="232.[1992,2585,1956,2380]" box="[2250,2581,1992,2026]" pageId="232" pageNumber="837">West Papua, Indonesia.”</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="discussion">
<paragraph blockId="232.[1992,2585,1956,2380]" pageId="232" pageNumber="837">
<taxonomicName box="[1994,2078,2035,2064]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="genus">Rattus</taxonomicName>
nikenai is in the Recent New Guinea clade within
<taxonomicName box="[2177,2261,2070,2104]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="genus">Rattus</taxonomicName>
. Populations of
<taxonomicName class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="species" species="nikenii">R. nikenii</taxonomicName>
were previously allocated to
<taxonomicName baseAuthorityName="Thomas" baseAuthorityYear="1888" box="[2463,2578,2110,2144]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="species" species="praetor">R.praetor</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6888373" ID-Zenodo-Dep="6888373" httpUri="https://zenodo.org/record/6888373/files/figure.png" inLine="true" pageId="232" pageNumber="837" startId="232.[1992,2164,2189,2223]" targetBox="[1378,1968,1963,2376]" targetPageId="232">
<paragraph blockId="232.[1992,2585,1956,2380]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1992,2168,2189,2223]" pageId="232" pageNumber="837">Distribution.</emphasis>
Gag I, Indonesia; and perhaps other islands in the region.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="description">
<paragraph blockId="232.[1992,2585,1956,2380]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1993,2272,2267,2301]" pageId="232" pageNumber="837">Descriptive notes.</emphasis>
Head-body 135-300 mm, tail 138-165 mm, hindfoot 31-36-5 mm; weight 113-175 g. The Gag
</paragraph>
<paragraph blockId="232.[1379,2585,2390,3207]" pageId="232" pageNumber="837">
Island Rat is mediumto large-sized and similar to the Large New Guinea Spiny Rat (
<taxonomicName baseAuthorityName="Thomas" baseAuthorityYear="1888" box="[1394,1532,2425,2459]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="232" pageNumber="837" phylum="Chordata" rank="species" species="praetor">R. praetor</taxonomicName>
), although it is considerably smaller on average, with shorter snout and hindfeet proportionately. Pelage is harsh and spiny. Dorsum is ashy gray, with light brown sides sharply demarcated from venter. Venter is white on chin, rufous merging to tawny on throat, brown to patchy light brown on chest, creamy on abdomen, and pale brown mixed with cream in scrotal area. Feet are grayish white dorsally. Ears are small, rounded, and similarly in color to dorsum; vibrissae are long. Tail is less than 100% of head-body length and unicolored blackish brown. Skull has oval profiled cranium and is altogether similar to Stein’s New Guinea Rat (R. stein). There are four pairs of mammae: two axillary and two inguinal.
</paragraph>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="biology_ecology">
<paragraph blockId="232.[1379,2585,2390,3207]" box="[1381,2120,2784,2813]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,1491,2784,2813]" pageId="232" pageNumber="837">Habitat.</emphasis>
Probably ultrabasic, evergreen valley forests.
</paragraph>
<paragraph blockId="232.[1379,2585,2390,3207]" box="[1381,1880,2819,2853]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,1642,2819,2853]" pageId="232" pageNumber="837">Food and Feeding.</emphasis>
No information.
</paragraph>
<paragraph blockId="232.[1379,2585,2390,3207]" box="[1381,1753,2862,2891]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,1515,2862,2891]" pageId="232" pageNumber="837">Breeding.</emphasis>
No information.
</paragraph>
<paragraph blockId="232.[1379,2585,2390,3207]" box="[1379,1853,2902,2931]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1379,1614,2902,2931]" pageId="232" pageNumber="837">Activity patterns.</emphasis>
No information.
</paragraph>
<paragraph blockId="232.[1379,2585,2390,3207]" box="[1381,2319,2941,2970]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,2080,2941,2970]" pageId="232" pageNumber="837">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="232" pageNumber="837" type="conservation">
<paragraph blockId="232.[1379,2585,2390,3207]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1381,1734,2980,3009]" pageId="232" pageNumber="837">Status and Conservation.</emphasis>
Classified as Vulnerable on The IUCN Red List. The GagIsland Rat was only recently recognized as a distinct species and has a very limited distribution (56 km?) that is completely in a protected area, although it is potentially threatened bynickel mining. More research is needed to fully understand its natural history, taxonomy, and conservation threats.
</paragraph>
</subSubSection>
<subSubSection box="[1382,2500,3186,3207]" pageId="232" pageNumber="837" type="bibRefCitation_list">
<paragraph blockId="232.[1379,2585,2390,3207]" box="[1382,2500,3186,3207]" pageId="232" pageNumber="837">
<emphasis bold="true" box="[1382,1534,3186,3207]" pageId="232" pageNumber="837">Bibliography.</emphasis>
Engelbrektsson &amp; Kennerley (2016a), Maryanto et al. (2010), Timm, Weijola et al. (2016).
</paragraph>
</subSubSection>
</treatment>
</document>
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