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<document id="3332DEAA84D9841FCD95C289CC4197A8" ID-CLB-Dataset="46351" ID-DOI="10.11646/zootaxa.3768.3.4" ID-GBIF-Dataset="ae50092c-8740-40db-89cc-959efad637da" ID-ISSN="1175-5326" ID-Zenodo-Dep="227248" ID-ZooBank="D315EBBE-ACEE-4E23-A6D8-C9BF6CB6ABA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460389200188" checkinUser="plazi" docAuthor="Liu, Yang, Fletcher, Murray J., Dietrich, Christopher H. &amp; Zhang, Ya-Lin" docDate="2014" docId="2547AB19FFC8FF89FF087212FBDCFD4B" docLanguage="en" docName="zt03768p350.pdf" docOrigin="Zootaxa 3768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asymmetrasca helica Liu and Zhang, sp. nov." docType="treatment" docVersion="9" lastPageNumber="333" masterDocId="D97ED361FFCCFF8FFF9F747DFFC0FFE3" masterDocTitle="New species and records of Asymmetrasca (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) from China and name changes in Empoasca (Matsumurasca)" masterLastPageNumber="350" masterPageNumber="327" pageNumber="331" updateTime="1698298032949" updateUser="ExternalLinkService">
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<mods:title id="E1842D22401EF07F66311C6232C33361">New species and records of Asymmetrasca (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) from China and name changes in Empoasca (Matsumurasca)</mods:title>
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<mods:namePart id="B617904FB1C392BCD19966869A6B5C85">Liu, Yang</mods:namePart>
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<mods:namePart id="115BAC17E43F59876D96925BD27C90E8">Fletcher, Murray J.</mods:namePart>
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<mods:namePart id="386ABEBFD092DF1B54A710CFBAB5604F">Dietrich, Christopher H.</mods:namePart>
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<mods:namePart id="77A2B8839373C82AD03D91076117976C">Zhang, Ya-Lin</mods:namePart>
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<mods:number id="36FD00818C33F3FD45CD49A3F569C159">3768</mods:number>
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<treatment id="2547AB19FFC8FF89FF087212FBDCFD4B" ID-DOI="http://doi.org/10.5281/zenodo.6133897" ID-GBIF-Taxon="119416307" ID-Zenodo-Dep="6133897" LSID="urn:lsid:plazi:treatment:2547AB19FFC8FF89FF087212FBDCFD4B" httpUri="http://treatment.plazi.org/id/2547AB19FFC8FF89FF087212FBDCFD4B" lastPageId="6" lastPageNumber="333" pageId="4" pageNumber="331">
<subSubSection id="E5F44984FFC8FF8BFF087212FEBBF948" pageId="4" pageNumber="331" type="nomenclature">
<paragraph id="AD511A0FFFC8FF8BFF087212FD0DF969" blockId="4.[151,717,1647,1707]" box="[151,717,1647,1674]" pageId="4" pageNumber="331">
<heading id="F619AD63FFC8FF8BFF087212FD0DF969" bold="true" box="[151,717,1647,1674]" fontSize="11" level="1" pageId="4" pageNumber="331" reason="1">
<emphasis id="9F9AC61DFFC8FF8BFF087212FD0DF969" bold="true" box="[151,717,1647,1674]" pageId="4" pageNumber="331">
<taxonomicName id="6AEE618CFFC8FF8BFF087212FDA3F969" ID-CoL="J6NB" authority="Liu and Zhang" authorityName="Liu and Zhang" box="[151,611,1647,1674]" class="Insecta" family="Cicadellidae" genus="Asymmetrasca" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="331" phylum="Arthropoda" rank="species" species="helica" status="sp. nov.">
<emphasis id="9F9AC61DFFC8FF8BFF087212FE57F96A" bold="true" box="[151,407,1647,1673]" italics="true" pageId="4" pageNumber="331">Asymmetrasca helica</emphasis>
, Liu and Zhang
</taxonomicName>
,
<taxonomicNameLabel id="84A97B66FFC8FF8BFDEF720DFD0DF969" box="[624,717,1648,1674]" pageId="4" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="AD511A0FFFC8FF8BFF0872EFFEBBF948" blockId="4.[151,717,1647,1707]" box="[151,379,1682,1707]" pageId="4" pageNumber="331">
(
<figureCitation id="35D5068AFFC8FF8BFF0072EFFEE0F948" box="[159,288,1682,1707]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="4" pageNumber="331">Figs 2129</figureCitation>
,
<figureCitation id="35D5068AFFC8FF8BFEB472EEFEB3F948" box="[299,371,1683,1707]" captionStart="FIGURES 94 97" captionStartId="17.[151,264,1823,1846]" captionTargetBox="[171,1435,1111,1780]" captionTargetId="figure@17.[159,1444,1089,1797]" captionTargetPageId="17" captionText="FIGURES 94 97. Asymmetrasca helica, 94, head and thorax, dorsal view; 95, face; 96, whole body, lateral view; 97, whole body, dorsal view." httpUri="https://zenodo.org/record/227266/files/figure.png" pageId="4" pageNumber="331">9497</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E5F44984FFC8FF89FF0872A7FEABFDFB" lastPageId="6" lastPageNumber="333" pageId="4" pageNumber="331" type="materials_examined">
<paragraph id="AD511A0FFFC8FF8BFF0872A7FD8EF8F4" blockId="4.[151,1436,1754,1995]" pageId="4" pageNumber="331">
<emphasis id="9F9AC61DFFC8FF8BFF0872A7FE8FF910" bold="true" box="[151,335,1754,1779]" pageId="4" pageNumber="331">
<typeStatus id="7255A4ADFFC8FF8BFF0872A7FF12F910" box="[151,210,1754,1779]" pageId="4" pageNumber="331">Type</typeStatus>
materials.
</emphasis>
<collectingCountry id="D5F95A9FFFC8FF8BFECA72A7FE5CF910" box="[341,412,1754,1779]" name="China" pageId="4" pageNumber="331">China</collectingCountry>
:
<typeStatus id="7255A4ADFFC8FF8BFE3972A7FDC8F910" box="[422,520,1754,1779]" pageId="4" pageNumber="331" type="holotype">holotype</typeStatus>
♂, Zhejiang Prov.: Mt. Tianmu,
<date id="D9503CCFFFC8FF8BFCEB72A7FC30F910" box="[884,1008,1754,1779]" pageId="4" pageNumber="331" value="2007-08-08">8.viii.2007</date>
, coll. Xinmin Zhang;
<typeStatus id="7255A4ADFFC8FF8BFB7B72A6FA94F910" box="[1252,1364,1755,1779]" pageId="4" pageNumber="331" type="paratype">paratypes</typeStatus>
: 2♂♂, same data as
<typeStatus id="7255A4ADFFC8FF8BFEB47283FE51F8F4" box="[299,401,1790,1815]" pageId="4" pageNumber="331" type="holotype">holotype</typeStatus>
. All in
<collectionCode id="CBFF82CAFFC8FF8BFE7C7282FD8AF8F4" box="[483,586,1791,1815]" pageId="4" pageNumber="331">NWAFU</collectionCode>
.
</paragraph>
<paragraph id="AD511A0FFFC8FF8BFF58735FFDC2F8D9" blockId="4.[151,1436,1754,1995]" box="[199,514,1826,1851]" pageId="4" pageNumber="331">Length: ♂ 3.2 mm3.3 mm.</paragraph>
<paragraph id="AD511A0FFFC8FF8BFF58733BFBABF828" blockId="4.[151,1436,1754,1995]" pageId="4" pageNumber="331">
General color of body yellow-greenish (
<figureCitation id="35D5068AFFC8FF8BFD19733BFD1AF8BC" box="[646,730,1862,1887]" captionStart="FIGURES 94 97" captionStartId="17.[151,264,1823,1846]" captionTargetBox="[171,1435,1111,1780]" captionTargetId="figure@17.[159,1444,1089,1797]" captionTargetPageId="17" captionText="FIGURES 94 97. Asymmetrasca helica, 94, head and thorax, dorsal view; 95, face; 96, whole body, lateral view; 97, whole body, dorsal view." httpUri="https://zenodo.org/record/227266/files/figure.png" pageId="4" pageNumber="331">Fig. 97</figureCitation>
). Crown yellowish, with some obvious green markings on each side of coronal suture (
<figureCitation id="35D5068AFFC8FF8BFE087317FE2BF860" box="[407,491,1898,1923]" captionStart="FIGURES 94 97" captionStartId="17.[151,264,1823,1846]" captionTargetBox="[171,1435,1111,1780]" captionTargetId="figure@17.[159,1444,1089,1797]" captionTargetPageId="17" captionText="FIGURES 94 97. Asymmetrasca helica, 94, head and thorax, dorsal view; 95, face; 96, whole body, lateral view; 97, whole body, dorsal view." httpUri="https://zenodo.org/record/227266/files/figure.png" pageId="4" pageNumber="331">Fig. 94</figureCitation>
). Eyes black. Pronotum broad, with irregular patches of cream-yellowish on anterior margin. Scutoscutellar sulcus beige. Lateral parts of scutellum with cream-yellowish triangular spots. Forewing and hind wing semitransparent. Abdomen yellow. Legs pale yellow to yellow (
<figureCitation id="35D5068AFFC8FF8BFB9573CFFB9EF828" box="[1034,1118,1970,1995]" captionStart="FIGURES 94 97" captionStartId="17.[151,264,1823,1846]" captionTargetBox="[171,1435,1111,1780]" captionTargetId="figure@17.[159,1444,1089,1797]" captionTargetPageId="17" captionText="FIGURES 94 97. Asymmetrasca helica, 94, head and thorax, dorsal view; 95, face; 96, whole body, lateral view; 97, whole body, dorsal view." httpUri="https://zenodo.org/record/227266/files/figure.png" pageId="4" pageNumber="331">Fig. 96</figureCitation>
).
</paragraph>
<caption id="F9914A87FFC9FF8AFF0877B8FF19FBFA" httpUri="https://zenodo.org/record/227251/files/figure.png" pageId="5" pageNumber="332" targetBox="[254,1336,210,933]" targetPageId="5">
<paragraph id="AD511A0FFFC9FF8AFF0877B8FF19FBFA" blockId="5.[151,1436,965,1049]" pageId="5" pageNumber="332">
<emphasis id="9F9AC61DFFC9FF8AFF0877B8FE94FC38" bold="true" box="[151,340,965,988]" pageId="5" pageNumber="332">FIGURES 1320.</emphasis>
<taxonomicName id="6AEE618CFFC9FF8AFEC677BAFD90FC3F" box="[345,592,966,988]" class="Insecta" family="Cicadellidae" genus="Asymmetrasca" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="decedens">
<emphasis id="9F9AC61DFFC9FF8AFEC677BAFD90FC3F" box="[345,592,966,988]" italics="true" pageId="5" pageNumber="332">Asymmetrasca decedens</emphasis>
</taxonomicName>
, 13, male pygofer, lateral view; 14, anal tube appendage; 15, abdominal apodemes; 16, aedeagus, ventral view; 17, same, lateral view; 18, connective; 19, subgenital plate; 20, paramere. (all after Dworakowska, 1970).
</paragraph>
</caption>
<caption id="F9914A87FFC9FF8AFF0873C7FA5BF80C" httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="5" pageNumber="332" targetBox="[232,1355,1115,1941]" targetPageId="5">
<paragraph id="AD511A0FFFC9FF8AFF0873C7FA5BF80C" blockId="5.[151,1435,1978,2031]" pageId="5" pageNumber="332">
<emphasis id="9F9AC61DFFC9FF8AFF0873C7FE94F833" bold="true" box="[151,340,1978,2001]" pageId="5" pageNumber="332">FIGURES 2129.</emphasis>
<taxonomicName id="6AEE618CFFC9FF8AFEC573C6FDF2F832" box="[346,562,1979,2001]" class="Insecta" family="Cicadellidae" genus="Asymmetrasca" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="helica">
<emphasis id="9F9AC61DFFC9FF8AFEC573C6FDF2F832" box="[346,562,1979,2001]" italics="true" pageId="5" pageNumber="332">Asymmetrasca helica</emphasis>
</taxonomicName>
, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate.
</paragraph>
</caption>
<paragraph id="AD511A0FFFCAFF89FF5874EAFEABFDFB" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="333">
Ventral abdominal apodemes subparallel-sided, reaching segment 5 (
<figureCitation id="35D5068AFFCAFF89FC7E74EAFBF9FF53" box="[993,1081,151,176]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 23</figureCitation>
). Male pygofer in lateral view slightly narrowing caudad (
<figureCitation id="35D5068AFFCAFF89FE5274C1FDE2FF37" box="[461,546,188,213]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 21</figureCitation>
); pygofer appendage slightly curved along margin of lobe in lateral view (
<figureCitation id="35D5068AFFCAFF89FAF174C1FF77FF1B" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 22</figureCitation>
). Anal hook well-developed, papillose apically (
<figureCitation id="35D5068AFFCAFF89FD4274A2FCF3FF1B" box="[733,819,223,248]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 26</figureCitation>
). Subgenital plate with ventral margin curved dorsally in nearly half apical part, sinuate, with numerous macrosetae and fine setae somewhat irregularly arranged; setal group A with 3 stout setae; group B with 1723 short microsetae, group C with three rows of macrosetae, group D arranged irregularly, lateral side with some slim setae (
<figureCitation id="35D5068AFFCAFF89FD627531FC92FE87" box="[765,850,332,357]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 29</figureCitation>
). Paramere long, subapically curved, dentifer with 5 teeth, 46 setae present subapically (
<figureCitation id="35D5068AFFCAFF89FDD27512FD66FE6B" box="[589,678,367,392]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 28</figureCitation>
). Connective broad, with posterior margin shallowly emarginate medially (
<figureCitation id="35D5068AFFCAFF89FE9175E9FEA8FE4F" box="[270,360,404,429]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Fig. 27</figureCitation>
). Aedeagus with preatrium nearly as long as shaft, slightly sinuate in lateral view; dorsal apodeme nearly as long as preatrium; shaft nearly straight, without dorsal denticuli; distal process elongate, slender, strongly and evenly curved throughout length, forming semicircle in posterior view; apex slender, without teeth (
<figureCitation id="35D5068AFFCAFF89FF437582FE9FFDFB" box="[220,351,511,536]" captionStart="FIGURES 21 29" captionStartId="5.[151,264,1978,2001]" captionTargetBox="[232,1355,1115,1941]" captionTargetId="figure@5.[224,1363,1099,1957]" captionTargetPageId="5" captionText="FIGURES 21 29. Asymmetrasca helica, 21, male pygofer, lateral view; 22, pygofer appendage; 23, abdominal apodemes; 24, aedeagus, ventral view; 25, aedeagus, lateral view; 26, anal tube appendage; 27, connective; 28, paramere; 29, subgenital plate." httpUri="https://zenodo.org/record/227252/files/figure.png" pageId="6" pageNumber="333">Figs 24, 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E5F44984FFCAFF89FF587659FDF5FD83" pageId="6" pageNumber="333" type="etymology">
<paragraph id="AD511A0FFFCAFF89FF587659FDF5FD83" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="333">
<emphasis id="9F9AC61DFFCAFF89FF587659FE8EFDDE" bold="true" box="[199,334,548,573]" pageId="6" pageNumber="333">Etymology.</emphasis>
The specific epithet is derived from the Greek word &quot;
<emphasis id="9F9AC61DFFCAFF89FC4B7659FBCAFDDF" box="[980,1034,548,572]" italics="true" pageId="6" pageNumber="333">helix</emphasis>
&quot; meaning &quot;curl&quot; and refers to the strongly curved aedeagal appendage.
</paragraph>
</subSubSection>
<subSubSection id="E5F44984FFCAFF89FF587611FBDCFD4B" pageId="6" pageNumber="333" type="diagnosis">
<paragraph id="AD511A0FFFCAFF89FF587611FBDCFD4B" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="333">
<emphasis id="9F9AC61DFFCAFF89FF587611FE81FD66" bold="true" box="[199,321,620,645]" pageId="6" pageNumber="333">Diagnosis.</emphasis>
This species is similar to
<taxonomicName id="6AEE618CFFCAFF89FDF37611FC13FD67" authority="Dworakowska, 1982" authorityName="Dworakowska" authorityYear="1982" box="[620,979,620,645]" class="Insecta" family="Cicadellidae" genus="Asymmetrasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="333" phylum="Arthropoda" rank="species" species="kaicola">
<emphasis id="9F9AC61DFFCAFF89FDF37611FD1EFD67" box="[620,734,620,644]" italics="true" pageId="6" pageNumber="333">A. kaicola</emphasis>
<bibRefCitation id="C97F67FEFFCAFF89FD787611FC13FD67" author="Dworakowska" box="[743,979,620,645]" pageId="6" pageNumber="333" refString="Dworakowska, I. (1982) Empoascini of Japan, Korea and north - east part of China (Homoptera: Auchenorrhyncha: Cicadellidae: Typhlocybinae). Reichenbachia, 20, 33 - 57." type="journal article" year="1982">Dworakowska, 1982</bibRefCitation>
</taxonomicName>
, but differs from the latter in having the aedeagal shaft straight and the apical process evenly curved and tapered distally.
</paragraph>
</subSubSection>
</treatment>
</document>