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<document ID-CLB-Dataset="122795" ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-GBIF-Dataset="5f938945-980c-433c-942a-f05d39529a25" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="F64697E7AAAF5C5A845C7FC2C4942AEC" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho vargasllosai Benedetti &amp; Pinto-da-Rocha 2022, sp. nov." docType="treatment" docUuid="C3D23CAE-1661-4036-AD00-E4D4F304070F" docUuidSource="ZooBank" docVersion="5" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1732651490011" updateUser="ExternalLinkService">
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<mods:title id="8FF597D76ADD6D12157E7CA9D4161F9C">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:name id="5367C1C9D3B6EF98D423E646D841E1B2" type="personal">
<mods:role id="43BC781F187B0BC41C960954B491828F">
<mods:roleTerm id="AADEFFD428DE16C24E9AFBED4FB308A0">Author</mods:roleTerm>
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<mods:namePart id="C3327213D0E5EF37C8C17493F56A31E8">Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation id="A4186C191249BF27E76D8B8E83E9E436">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier id="0CEA5078D859F544E62E0D4CD8139E52" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0DD8446AF42265113D44A830B8AE2ADE">Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier id="AAF9C61C22E9316BF91801A5F1902C3E" type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation id="4864FBA829BD1C2735FE0E966E8D6BE8">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="9546AD5C898E8454B6C85141AA13B75D">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="91BFE662DE1E1634C52A84050D2F845A">2022</mods:date>
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<mods:number id="F1497FF393649D6A13F3B483A19C6909">2022-08-16</mods:number>
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<treatment id="F64697E7AAAF5C5A845C7FC2C4942AEC" ID-GBIF-Taxon="199468718" LSID="urn:lsid:zoobank.org:act:C3D23CAE-1661-4036-AD00-E4D4F304070F" httpUri="http://treatment.plazi.org/id/F64697E7AAAF5C5A845C7FC2C4942AEC" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
<subSubSection id="5AEDE10A0581F884BFB6DFE8D957D565" pageId="0" pageNumber="309" type="nomenclature">
<paragraph id="9910A11D47CCBE587AE1C4CECD03A353" pageId="0" pageNumber="309">
3.26.
<taxonomicName id="C7C8928429D4FFCD0ED27E7313A3CCF2" ID-CoL="9X66B" LSID="http://zoobank.org/C3D23CAE-1661-4036-AD00-E4D4F304070F" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho vargasllosai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vargasllosai" status="sp. nov.">Ayacucho vargasllosai</taxonomicName>
<taxonomicNameLabel id="CB3B72FF5EF687F767D4877CAC4D76A9" pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="895637B105820AEB505A0AE3341EF996" pageId="0" pageNumber="309" type="description">
<paragraph id="8D6001E2FFD1FB971422E5C4263108ED" pageId="0" pageNumber="309">
<figureCitation id="CA0CA3D86BEC9C29042128FD63403015" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">Figs 3G</figureCitation>
<figureCitation id="6C24692933E44DC513AB145AFFAF4248" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">, 8C, D</figureCitation>
<figureCitation id="BE6C72090514A82EA1192A7F919B8768" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury &amp; Maury, 1998; G - H Incasarcus dianae Kury &amp; Maury, 1998; I - J: Incasarcus pictus Kury &amp; Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure13" httpUri="https://binary.pensoft.net/fig/729887" pageId="0" pageNumber="309">, 13A, B</figureCitation>
<figureCitation id="0130568525991A8ADC1AB5042CE4B0AB" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">, 23F</figureCitation>
<figureCitation id="D869C0B626A4B7C5D3684DA7A2A2EAB5" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1983EA17ED8A9BE8688F268A4FC5344" pageId="0" pageNumber="309" type="description">
<paragraph id="29C1EFE0B75CCE2BFFC08B45A6DC8EA9" pageId="0" pageNumber="309">Description.</paragraph>
<paragraph id="311735ACC3C9031A838F9F2D284C3F3F" pageId="0" pageNumber="309">
<emphasis id="832D0C1E3855C33C90063A24AF849EF3" bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis id="EBD7FBA554F3D4407BD792D2E5EE818A" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="79FFA2204745F4010208D1E12FB4944A" italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) DSW: 3.3; DSL: 4.6; CL: 1.7. FIVL: 4.7. ChL: 2.5.
<emphasis id="131CAB189895A354EC7A79E79B9DC610" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation id="75BDD92875FB1E83E87D67C7B2D48619" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">23F</figureCitation>
) Chelicerae, pedipalpus, carapace and lateral margin of dorsal scutum and legs orange. Scutal areas, posterior margin of dorsal scutum and free tergites brownish.
<emphasis id="D02465F74BB14991D9999BB2A34265EF" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation id="1C4BC5C64660DA8BCD1936EC84695E35" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3G</figureCitation>
) Alpha-type DSS, with shallower constrictions. Anterior margin of carapace with median elevation totally covered with granules. Ocularium with tiny acuminate tubercles. Areas I-IV with one pair of small median tubercles (larger than the pair of the ocularium). Posterior margin of dorsal scutum with a transverse row of four tubercles. Free tergites I--III with a row of 3-6 tubercles, larger than tubercles of areas I-IV.
<emphasis id="CA1F4B82AA0B891CF0B7E4718958F0DF" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation id="737624D5C9B7D0DE28F444DD38E2AA57" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3G</figureCitation>
) Swollen in male. Segment I densely covered with granules throughout its length. Segment II predominantly smooth, with some hairs on the front surface; finger with four teeth. Segment III with two teeth.
<taxonomicName id="61EAC93A1ABC0099A8A0C035FE1C69D7" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis id="6C759DFFB8779EB3DA347D2A8226005C" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: With very small granules sparsely distributed on the dorsal surface of the femur-tibia. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of five small ventromedian setiferous tubercles. Tibia: prolateral IIII, IiI; retrolateral iII(II), iIII. Tarsus: prolateral IIi; retrolateral Iii, Ii.
<emphasis id="0DDCE3D463551D183E657085406C7C67" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxae I-IV and stigmatic area with sparse small granules; coxa I with a longitudinal row of small setiferous tubercles. Genital area smooth. Free sternites I-IV and anal operculum with sparse small granules.
<emphasis id="FE2076DEFEB76EB40A80AE7F33298162" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation id="1F7E416941B39780B47B4EA456514859" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3G</figureCitation>
,
<figureCitation id="6F48346B6783256EB14ED7AD79DB76F4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">8C, D</figureCitation>
) Coxae I-III each one with a retrolateral and a prolateral apophysis. Coxa IV densely granulate, unarmed. Trochanters I-IV densely granulate, unarmed. Femora I-IV densely granulate throughout their dorsal, retrolateral and prolateral faces. Femora I-III with two rows of granules in the ventral face, larger than other granules of these segments. Femur IV with two ventral rows of granules that increase in size apically, greater than other granules of femur IV. Patellae I-IV densely granulate. Tibiae I-IV granulate (larger granules on ventral face). Tarsal segmentation: (
<emphasis id="319AB0A9F7452A9235EFE3905CFF3C87" italics="true" pageId="0" pageNumber="309">n</emphasis>
= 1) 7, 11, 7, 7.
<emphasis id="64BAE715BC22C535ADA50BE69EA5B9F5" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation id="E37BE059E53B0B8526C3BB5C926D5263" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury &amp; Maury, 1998; G - H Incasarcus dianae Kury &amp; Maury, 1998; I - J: Incasarcus pictus Kury &amp; Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure13" httpUri="https://binary.pensoft.net/fig/729887" pageId="0" pageNumber="309">13A, B</figureCitation>
) VP subrectangular in dorsal view, with distal half larger than basal half; distal margin straight. MS C1-C7 (C8) apical, long and curved; MS A1 median placed, long and straight (smaller than MS C); MS B1 sub basal, long and straight (smaller than MS A). Lateral sacs long, apically tapered; with long T3-like microsetae. Stylus apically inflated, with a conspicuous ventral projection. Dorsal process absent. Promontory straight. -
<emphasis id="AC8AC212CEDB1F44C2092ACB9DFDD9F3" bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis id="DBD4C5DB1C228D29A38F399ABC967481" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(n=3) DSW: 3.0-3.1; DSL: 4.4-4.5; CL: 1.7. FIVL: 4.5-4.7. ChL: 2.2-2.3. Areas I-IV with one pair of small median tubercles, smaller than in males.
<taxonomicName id="5A9BB36DB426E08585E47B0B749E13FA" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">Pedipalpus</taxonomicName>
with ventral tubercles slightly smaller than in males. Tibia: prolateral IiI; retrolateral iIIIi, iIII. Tarsus: retrolateral iIIi, iIii. Genital area with sparse granules. Femur IV with ventral tubercles smaller than in males. Tarsal segmentation (
<emphasis id="43FB38895D587BEF57B29A0F97B68529" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3): 7,11,7,8.
</paragraph>
<caption id="6890D760ED04D1E71D493F7DEB632E2E" doi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309" start="Figure 19" startId="F19">
<paragraph id="EDE05F30B66780EB14E75EA496671B42" pageId="0" pageNumber="309">
<emphasis id="7123EB00E53759066AEB835EEC0F748F" bold="true" pageId="0" pageNumber="309">Figure 19.</emphasis>
Penis of
<taxonomicName id="961CA0C57D02BE41CAC58B2E82255DB5" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="C3677E0A842DCDB2C2CA0369BFC3EC82" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
</taxonomicName>
, dorsal, lateral and ventral views, respectively.
<emphasis id="4F0B4E1066BA5543F2ADFBC57B79EF96" bold="true" pageId="0" pageNumber="309">A</emphasis>
-
<emphasis id="935624AA8786B284286C27C9E184002F" bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName id="88B727DC11EE50E6FA309A8B06B45E07" class="Arachnida" kingdom="Animalia" lsidName="M. bergmani" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bergmani">
<emphasis id="72C16DF784943F6BCDD2881A06234257" italics="true" pageId="0" pageNumber="309">M. bergmani</emphasis>
</taxonomicName>
<emphasis id="D9DD5C9A15B68145AEBF4EF05703CCEA" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="D76D1B5612EC6B2518DEB3567821AA03" bold="true" pageId="0" pageNumber="309">D</emphasis>
-
<emphasis id="AC7633A5C6CF527F500B8991A3984E2C" bold="true" pageId="0" pageNumber="309">F</emphasis>
<taxonomicName id="632B42D3BB64E26C93C362D1A2C1EDB3" class="Arachnida" kingdom="Animalia" lsidName="M. clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
<emphasis id="92B642A4C42261CA7E36C6E53262CF90" italics="true" pageId="0" pageNumber="309">M. clavifemur</emphasis>
</taxonomicName>
(Roewer, 1929);
<emphasis id="25DDFCDBABE5646264BCC424AB56A9F1" bold="true" pageId="0" pageNumber="309">G</emphasis>
-
<emphasis id="66F115E3CE663B5B8B128A69A8BE39BD" bold="true" pageId="0" pageNumber="309">I</emphasis>
<taxonomicName id="92C7DD0E49BC075453E1605CF77D461D" class="Arachnida" kingdom="Animalia" lsidName="M. fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
<emphasis id="846A2B914CB98C62667E090FD1DBB341" italics="true" pageId="0" pageNumber="309">M. fellinii</emphasis>
</taxonomicName>
<emphasis id="C2111AF30397C1C3AFC7A8B4F9C214D0" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
; Legend bars = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DF301C4F02BCAE84005DC55A83306DE4" pageId="0" pageNumber="309" type="diagnosis">
<paragraph id="505D6B8302478A665598CAB33B55F85D" pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph id="ADB963195403B7B2432E83D66325B037" pageId="0" pageNumber="309">
Similar to
<taxonomicName id="D9C5C65CEFBA461FB8F9754A33B8A6D9" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho glauberrochai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="glauberrochai">
<emphasis id="B68EA394D536F59FD1DC79DB513EFE4D" italics="true" pageId="0" pageNumber="309">Ayacucho glauberrochai</emphasis>
</taxonomicName>
<emphasis id="5171195D7ABB254B51E4CDE82FC80E49" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
<taxonomicName id="A79D670EB050C24E1D80F6F9E12BE85C" lsidName="A. pomacocha" pageId="0" pageNumber="309" rank="species" species="pomacocha">
<emphasis id="33CEBC8A475E712E84931702A15152B8" italics="true" pageId="0" pageNumber="309">A. pomacocha</emphasis>
</taxonomicName>
<emphasis id="59D05C5DF45A2295230B1203E2A9148D" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
,
<taxonomicName id="C6C83DB326EB0BE2047C30E89AC5405B" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="06EB71841E754C20C0DF7FE8281D1398" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="BAE48107F4D78FBF5A30E57ECE2BD335" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, and
<taxonomicName id="D2B91587AAA88823D3EED22DB2F50C14" lsidName="A. titschacki" pageId="0" pageNumber="309" rank="species" species="titschacki">
<emphasis id="A024F260C14B9B91D759E2861FE961F1" italics="true" pageId="0" pageNumber="309">A. titschacki</emphasis>
</taxonomicName>
in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig.
<figureCitation id="064F3F0001A6102629B60F378F241F88" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3G</figureCitation>
); femur IV of males without strong armature (except in ventral surface of femur IV in
<taxonomicName id="1DD91D6E230E38CC3E0A0B49E9E54BE2" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="F806214810EF242FCA0EA921EA7FD3E0" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="4592B990C6B31BA022DA8D68CB076F55" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
; Fig.
<figureCitation id="CD0E52FA0017A3A03D1A65920EB540CC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">8C, D</figureCitation>
). It differs from the previously mentioned species in the following combination of characteristics: ocularium with a pair of small tubercles; areas I-IV with a pair of tubercles (larger than those in ocularium; Fig.
<figureCitation id="B89C681A99D47B93B8A735062F7134C7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3G</figureCitation>
); male femur IV unarmed (unlike
<taxonomicName id="5F10AFEBAB1852A30628CE2745E4ED4D" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="6D0940A38A2F113BB95F38192E3ED4FF" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="6A6D0A7F3C8680A847924881D8317483" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
; Fig.
<figureCitation id="75B97E119A1804070E7637A466BC95F5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male trochanter-tibia IV of leg IV of Ayacucho, dorsal and ventral view, respectively. A - B A. uniseriatus (Roewer, 1959) comb. nov.; C - D A. vargasllosai sp. nov.; E - F A. weyrauchi (Roewer, 1952) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure8" httpUri="https://binary.pensoft.net/fig/729882" pageId="0" pageNumber="309">8C, D</figureCitation>
); and penis VP subrectangular, with 7-8 apical MS C (Fig.
<figureCitation id="E93FAD363E629CA34A6A468C70D979AA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury &amp; Maury, 1998; G - H Incasarcus dianae Kury &amp; Maury, 1998; I - J: Incasarcus pictus Kury &amp; Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure13" httpUri="https://binary.pensoft.net/fig/729887" pageId="0" pageNumber="309">13A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4BB60DD9FA065AB876EC679DD7C67C6F" pageId="0" pageNumber="309" type="etymology">
<paragraph id="84FC78A54101F819C505E54D61766CD3" pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph id="6F6F47FF7A274EA587A4E47A51D4AC52" pageId="0" pageNumber="309">The specific epithet of masculine gender, in the genitive form, dedicated to the Peruvian writer, politician, journalist, essayist, filmmaker, college professor and Nobel Prize winner Jorge Mario Pedro Vargas Llosa (born 1936), more commonly known as Mario Vargas Llosa.</paragraph>
</subSubSection>
<subSubSection id="EBA2C211BF4EB1A9A167C4D5BFB1D5E6" pageId="0" pageNumber="309" type="distribution">
<paragraph id="3FD6651E280F6EB7B8CDD9E9340396CB" pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph id="9DCE85C74C4DECECDCFD136EA92A7C6A" pageId="0" pageNumber="309">
(Fig.
<figureCitation id="A2D14F0466E31BBF92AC8FD02F5618BF" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
) PERU.
<normalizedToken id="1D16437209C07B2F4992BF44668A3EBA" originalValue="Junín">Junin</normalizedToken>
. Cruce Mina.
</paragraph>
</subSubSection>
<subSubSection id="86FD9BE5642B24B0F5D6C020524904A6" pageId="0" pageNumber="309" type="materials_examined">
<paragraph id="7A40537B33459D8DBA3BB8EEDC0AF5E3" pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph id="5E95E760E445CB1E59F087A9B74D54E4" pageId="0" pageNumber="309">
<emphasis id="A85855D378B8DE61AE63E882839C7C70" bold="true" pageId="0" pageNumber="309">
<emphasis id="AED29852CF2378BD85A08509E48913DD" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation id="A47104F2A4DE972D02F555A7EE1C6A07" ID-GBIF-Occurrence="3889641310" collectingDate="2011-04-22" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, D. Silva" country="PERU" latitude="-11.379278" location="Cemento Andino" longLatPrecision="1" longitude="-75.87875" municipality="Cruce Mina" specimenCount="♂" stateProvince="Junin" typeStatus="Holotype">
<typeStatus id="7022E370F1132077B7C676BE202E3A40">Holotype</typeStatus>
<specimenCount id="C3422B654104940CB7BAE16DAD0E5CFD">♂</specimenCount>
, '
<collectingCountry id="360587EA76F4D51405BB8E1B63CC0BEE" name="Peru">PERU</collectingCountry>
,
<normalizedToken id="049D244C51C7AB994D226856159B7AE1" originalValue="Junín">
<collectingRegion id="0713CCE13F0AA890FC0566AC705F4A3D" country="Peru" name="Junin">Junin</collectingRegion>
</normalizedToken>
,
<collectingMunicipality id="95B78B404FE19EFFEA036CA01B7A2AB6">Cruce Mina</collectingMunicipality>
,
<location id="718E5277DA4454725DF7A5DBB1328100" LSID="urn:lsid:plazi:treatment:F64697E7AAAF5C5A845C7FC2C4942AEC:718E5277DA4454725DF7A5DBB1328100" country="PERU" latitude="-11.379278" longLatPrecision="1" longitude="-75.87875" municipality="Cruce Mina" name="Cemento Andino" stateProvince="Junin">Cemento Andino</location>
,
<geoCoordinate id="2905F1E114DD8E414226D25E69647DBA" degrees="11" direction="south" minutes="22" orientation="latitude" precision="1" seconds="45.4" value="-11.379278">11°22′45.4″S</geoCoordinate>
<geoCoordinate id="92A8697E27800B1791D837F4F8176BAF" degrees="75" direction="west" minutes="52" orientation="longitude" precision="1" seconds="43.5" value="-75.87875">75°52′43.5″W</geoCoordinate>
,|
<collectingDate id="1D5795D3E218A643AAA49E6AD69592EF" value="2011-04-22">22/IV/2011</collectingDate>
,
<collectorName id="D8357E88399A48BC07B5359B6525988E">R. Pinto-da-Rocha</collectorName>
,
<collectorName id="A3F2D017E9CA91D2E742500E0D57C7BC">A. Benedetti</collectorName>
,
<collectorName id="869B0954CF0C55C4E10DD6B320A2D7B0">J. Ochoa</collectorName>
&amp;
<collectorName id="42B9604A9DEDB8355AA004810D4D8117">D. Silva</collectorName>
leg. (MUBI)
</materialsCitation>
</emphasis>
-
<emphasis id="9E78DA88415F2B63BDFE2E890AD3D140" bold="true" pageId="0" pageNumber="309">
<typeStatus id="9C9C4D9DC7C592364A2054C77DD6E243">Paratype</typeStatus>
</emphasis>
<specimenCount id="ECA476AF6211AE037D32612A4FDEBF11" type="female">1 ♀</specimenCount>
,
<normalizedToken id="4FBB6998E55E90A494BBC41C7A2495FD" originalValue="‘ditto’">'ditto'</normalizedToken>
(MUBI);
<emphasis id="C0717598A991ABC3252B655B747C2DF5" bold="true" pageId="0" pageNumber="309">
<typeStatus id="371858A32368B40781D611142AA0DD06">Paratype</typeStatus>
</emphasis>
<specimenCount id="EC79992E09ABACEB27B825B86F180C1F" type="female">1 ♀</specimenCount>
,
<normalizedToken id="6E5A51352B86B58B3395C94AFAA416C3" originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 36975);
<emphasis id="DA648959BC8E56019478E84AD52FA965" bold="true" pageId="0" pageNumber="309">
<typeStatus id="E5BE22B0130AEF551C9E7EB38CF21FEE">Paratype</typeStatus>
</emphasis>
<specimenCount id="04294C6476EBE69FC44F909E042DD301" type="male">1 ♂</specimenCount>
,
<specimenCount id="6DF34CEC7E4DDF328657635B5EE07503" type="female">1 ♀</specimenCount>
,
<normalizedToken id="E50DEBA77392FE3CF6D7B9DE1EA819FA" originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 73007).
</paragraph>
</subSubSection>
</treatment>
</document>
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