treatments-xml/data/3E/50/87/3E5087BBFFF2372CFCFEFA85FB06E4CC.xml

<document id="0236535EBF939023245D2AA6AE502D6B" ID-DOI="10.3853/j.2201-4349.66.2014.1596" ID-ISSN="2201-4349" ID-Zenodo-Dep="4684561" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618331494905" checkinUser="carolina" docAuthor="Harris, Vernon A." docDate="2014" docId="3E5087BBFFF2372CFCFEFA85FB06E4CC" docLanguage="en" docName="RecAustMus.66.2.167-196.pdf" docOrigin="Records of the Australian Museum 66 (2)" docSource="http://dx.doi.org/10.3853/j.2201-4349.66.2014.1596" docStyle="DocumentStyle:A2A8AC5C71CC92C4EA50773B9B9E6D6A.1:RecAustMus.2008-2018.journal_article.1cover" docStyleId="A2A8AC5C71CC92C4EA50773B9B9E6D6A" docStyleName="RecAustMus.2008-2018.journal_article.1cover" docStyleVersion="1" docTitle="Cereudorsum verrucosum Harris 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="177" masterDocId="C269FFC3FFF03727FF81FFADFFD7E574" masterDocTitle="Porcellidiidae of Australia (Harpacticoida, Copepoda). III. Synopsis of Genera and Species" masterLastPageNumber="196" masterPageNumber="167" pageNumber="168" updateTime="1698904935843" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5DEEDE8B273084192749EA0B7D5087CB">Porcellidiidae of Australia (Harpacticoida, Copepoda). III. Synopsis of Genera and Species</mods:title>
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<paragraph id="B64636ADFFF23725FCFEFA85FAFFE036" blockId="2.[895,1320,1320,1347]" box="[895,1320,1320,1347]" pageId="2" pageNumber="168">
<heading id="ED0E81C1FFF23725FCFEFA85FAFFE036" box="[895,1320,1320,1347]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="168" reason="2">
<emphasis id="848DEABFFFF23725FCFEFA85FAFFE036" bold="true" box="[895,1320,1320,1347]" pageId="2" pageNumber="168">
<taxonomicName id="71F94D2EFFF23725FCFEFA85FB17E036" authority="Harris, 2014" authorityName="Harris" authorityYear="2014" box="[895,1216,1320,1347]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="168" phylum="Arthropoda" rank="species" species="verrucosum" status="sp. nov.">
<emphasis id="848DEABFFFF23725FCFEFA85FB17E036" bold="true" box="[895,1216,1320,1347]" italics="true" pageId="2" pageNumber="168">Cereudorsum verrucosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FBE57C4FFF23725FB49FA85FAFFE036" box="[1224,1320,1320,1346]" pageId="2" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFF23725FC58FAF0FB19E00C" box="[985,1230,1373,1400]" pageId="2" pageNumber="168" type="description">
<paragraph id="B64636ADFFF23725FC58FAF0FB19E00C" blockId="2.[985,1230,1373,1400]" box="[985,1230,1373,1400]" pageId="2" pageNumber="168">
<figureCitation id="2EC22A28FFF23725FC58FAF0FB93E00C" box="[985,1092,1373,1400]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[166,229,1867,1889]" captionStartId-1="4.[121,184,1862,1884]" captionStartId-2="5.[166,229,1862,1884]" captionStartId-3="8.[121,184,1834,1856]" captionStartId-4="9.[166,229,1872,1894]" captionTargetBox-0="[243,1386,160,1829]" captionTargetBox-1="[207,1351,176,1821]" captionTargetBox-2="[228,1404,153,1804]" captionTargetBox-3="[197,1336,172,1783]" captionTargetBox-4="[239,1403,166,1825]" captionTargetId-0="figure-8@3.[166,1466,151,1833]" captionTargetId-1="figure-10@4.[121,1421,151,1830]" captionTargetId-2="figure-8@5.[166,1466,151,1819]" captionTargetId-3="figure-10@8.[121,1421,151,1786]" captionTargetId-4="figure-8@9.[166,1466,151,1833]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." captionText-1="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." captionText-2="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." captionText-3="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." captionText-4="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4684563" figureDoi-1="http://doi.org/10.5281/zenodo.4684565" figureDoi-2="http://doi.org/10.5281/zenodo.4684567" figureDoi-3="http://doi.org/10.5281/zenodo.4684573" figureDoi-4="http://doi.org/10.5281/zenodo.4684577" httpUri-0="https://zenodo.org/record/4684563/files/figure.png" httpUri-1="https://zenodo.org/record/4684565/files/figure.png" httpUri-2="https://zenodo.org/record/4684567/files/figure.png" httpUri-3="https://zenodo.org/record/4684573/files/figure.png" httpUri-4="https://zenodo.org/record/4684577/files/figure.png" pageId="2" pageNumber="168">Figs 1–5</figureCitation>
; Plates 1, 2
</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFF23725FC9BFA20FB69E321" pageId="2" pageNumber="168" type="materials_examined">
<paragraph id="B64636ADFFF23725FC9BFA20FB69E321" blockId="2.[793,1422,1420,1621]" pageId="2" pageNumber="168">
<emphasis id="848DEABFFFF23725FC9BFA20FC68E0D0" bold="true" box="[794,959,1420,1445]" pageId="2" pageNumber="168">Type material</emphasis>
. 
<materialsCitation id="06913CF0FFF23725FC51FA20FCB8E0B6" accessionNumber="P81216" pageId="2" pageNumber="168" specimenCount="1" typeStatus="holotype">
<typeStatus id="6942880FFFF23725FC51FA20FB95E0D1" box="[976,1090,1421,1445]" pageId="2" pageNumber="168" type="holotype">HOLOTYPE</typeStatus>
adult male, length 
<quantity id="71019B48FFF23725FAA2FA20FA5EE0D0" box="[1315,1417,1421,1445]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="168" unit="mm" value="0.73">0.73 mm</quantity>
, P81216
</materialsCitation>
; 
<materialsCitation id="06913CF0FFF23725FCFBFA07FA5EE0B6" accessionNumber="P81217" box="[890,1417,1450,1474]" pageId="2" pageNumber="168" specimenCount="1" typeStatus="allotype">
<typeStatus id="6942880FFFF23725FCFBFA07FC34E0B6" box="[890,995,1450,1474]" pageId="2" pageNumber="168" type="allotype">ALLOTYPE</typeStatus>
adult female, length 
<quantity id="71019B48FFF23725FB46FA07FAFEE0B5" box="[1223,1321,1450,1474]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="168" unit="mm" value="0.92">0.92 mm</quantity>
, P81217
</materialsCitation>
; 
<materialsCitation id="06913CF0FFF23725FC9BFA6AFB6DE321" accessionNumber="P81218" collectionCode="AM, V" collectorName="A. Harris" county="Gunnamatta Bay" latitude="-151.13333" location="Sydney" municipality="Port Hacking" pageId="2" pageNumber="168" specimenCount="16" specimenCount-female="10" specimenCount-male="6" stateProvince="New South Wales" typeStatus="paratype">
<typeStatus id="6942880FFFF23725FC9BFA6AFCADE0AB" box="[794,890,1479,1503]" pageId="2" pageNumber="168" type="paratype">PARATYPE</typeStatus>
specimens 
<specimenCount id="A0FFFD24FFF23725FC6EFA6AFBE1E0AB" box="[1007,1078,1479,1503]" pageId="2" pageNumber="168" type="female">10 ♀♀</specimenCount>
, 
<specimenCount id="A0FFFD24FFF23725FBC1FA6AFBABE0AB" box="[1088,1148,1479,1503]" pageId="2" pageNumber="168" type="male">6 ♂♂</specimenCount>
, P81218, deposited at 
<collectionCode id="D0E8AE68FFF23725FADDFA6AFA5EE0AB" box="[1372,1417,1479,1503]" country="Australia" name="Australian Museum" pageId="2" pageNumber="168" type="Museum">AM</collectionCode>
, Sydney. Additional 
<typeStatus id="6942880FFFF23725FC6EFA48FB82E089" box="[1007,1109,1509,1533]" pageId="2" pageNumber="168" type="paratype">paratypes</typeStatus>
deposited at NHM, London. All collected from rotting 
<taxonomicName id="71F94D2EFFF23725FBADF9AFFB08E36E" box="[1068,1247,1538,1562]" class="Phaeophyceae" family="Lessoniaceae" genus="Ecklonia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Laminariales" pageId="2" pageNumber="168" phylum="Ochrophyta" rank="species" species="radiata">
<emphasis id="848DEABFFFF23725FBADF9AFFB08E36E" box="[1068,1247,1538,1562]" italics="true" pageId="2" pageNumber="168">Ecklonia radiata</emphasis>
</taxonomicName>
near entrance to 
<collectingCounty id="5F274E21FFF23725FC98F9B2FC1AE34C" box="[793,973,1567,1592]" pageId="2" pageNumber="168">Gunnamatta Bay</collectingCounty>
, 
<collectingMunicipality id="5622ACD7FFF23725FC59F98DFBBFE343" box="[984,1128,1567,1591]" pageId="2" pageNumber="168">Port Hacking</collectingMunicipality>
, 
<location id="B3266076FFF23725FBF2F9B2FB13E343" LSID="urn:lsid:plazi:treatment:3E5087BBFFF2372CFCFEFA85FB06E4CC:B3266076FFF23725FBF2F9B2FB13E343" box="[1139,1220,1567,1591]" county="Gunnamatta Bay" latitude="-151.13333" municipality="Port Hacking" name="Sydney" pageId="2" pageNumber="168" stateProvince="New South Wales">Sydney</location>
, 
<collectingRegion id="743DF84FFFF23725FB4FF98DFA5BE343" box="[1230,1420,1567,1591]" country="Australia" name="New South Wales" pageId="2" pageNumber="168">New South Wales</collectingRegion>
(34°05' 
<geoCoordinate id="D3CD506AFFF23725FCE9F991FC06E321" box="[872,977,1596,1621]" degrees="151" direction="south" minutes="08" orientation="latitude" pageId="2" pageNumber="168" precision="925" value="-151.13333">S 151°08'</geoCoordinate>
E), 
<collectionCode id="D0E8AE68FFF23725FC74F990FBDFE320" box="[1013,1032,1597,1620]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="168" type="Museum">V</collectionCode>
. 
<collectorName id="1B0C537BFFF23725FB90F990FBA1E320" box="[1041,1142,1596,1621]" pageId="2" pageNumber="168">A. Harris</collectorName>
, 1977
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFF2372DFC9BF9C7FDBCE636" lastPageId="10" lastPageNumber="176" pageId="2" pageNumber="168" type="diagnosis">
<paragraph id="B64636ADFFF23725FC9BF9C7FBE5E2D3" blockId="2.[792,1423,1642,1959]" pageId="2" pageNumber="168">
<emphasis id="848DEABFFFF23725FC9BF9C7FC5FE3F6" bold="true" box="[794,904,1642,1666]" pageId="2" pageNumber="168">Diagnosis</emphasis>
. Female with blocks of microtubules near edge on anterior half of cephalosome, microtubules absent from male cephalosome; both male and female have two dorsal organs with honeycomb-like cuticle on cephalosome and one each on metasome segment 3 and genital double-somite; numerous dorsal sensilla (&gt; 100) on cephalosome, metasome and genital double-somite; female caudal rami with two longitudinal ridges; internal seta absent from segment 1 of male and female P4 endopod; first (lateral) seta on male P5 with five strong ventral setules, each terminal seta with row of three setules at its base.
</paragraph>
<caption id="E2866625FFF33724FF27F8E6FAF7E20F" ID-DOI="http://doi.org/10.5281/zenodo.4684563" ID-Zenodo-Dep="4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="3" pageNumber="169" startId="3.[166,229,1867,1889]" targetBox="[243,1386,160,1829]" targetPageId="3">
<paragraph id="B64636ADFFF33724FF27F8E6FAF7E20F" blockId="3.[166,1467,1867,1916]" pageId="3" pageNumber="169">
Figure 1. 
<taxonomicName id="71F94D2EFFF33724FE84F8E6FE28E215" authority="Harris, 2014" authorityName="Harris" authorityYear="2014" box="[261,511,1867,1889]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="169" phylum="Arthropoda" rank="species" species="verrucosum" status="sp. nov.">
<emphasis id="848DEABFFFF33724FE84F8E6FE28E215" box="[261,511,1867,1889]" italics="true" pageId="3" pageNumber="169">Cereudorsum verrucosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FBE57C4FFF33724FD87F8E6FD85E214" box="[518,594,1867,1888]" pageId="3" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
Female: 
<emphasis id="848DEABFFFF33724FD2DF8E6FD1DE214" box="[684,714,1867,1888]" italics="true" pageId="3" pageNumber="169">(A)</emphasis>
adult; 
<emphasis id="848DEABFFFF33724FC8EF8E6FC9CE214" box="[783,843,1867,1888]" italics="true" pageId="3" pageNumber="169">(C, E)</emphasis>
genital double-somite (
<emphasis id="848DEABFFFF33724FBB0F8E1FB97E215" box="[1073,1088,1868,1889]" italics="true" pageId="3" pageNumber="169">E</emphasis>
detail of posterior lobe); 
<emphasis id="848DEABFFFF33724FAB8F8E6FAA1E214" box="[1337,1398,1867,1888]" italics="true" pageId="3" pageNumber="169">(D, F)</emphasis>
caudal rami (
<emphasis id="848DEABFFFF33724FF61F8CBFF26E20F" box="[224,241,1894,1915]" italics="true" pageId="3" pageNumber="169">D</emphasis>
detail of terminal setae). Male: 
<emphasis id="848DEABFFFF33724FDA4F8CBFD94E20F" box="[549,579,1894,1915]" italics="true" pageId="3" pageNumber="169">(B)</emphasis>
adult. Scale bar: A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm.
</paragraph>
</caption>
<caption id="E2866625FFF43723FFF8F8EBFDDBE2E4" ID-DOI="http://doi.org/10.5281/zenodo.4684565" ID-Zenodo-Dep="4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="4" pageNumber="170" startId="4.[121,184,1862,1884]" targetBox="[207,1351,176,1821]" targetPageId="4">
<paragraph id="B64636ADFFF43723FFF8F8EBFDDBE2E4" blockId="4.[120,1422,1862,1936]" pageId="4" pageNumber="170">
Figure 2. 
<taxonomicName id="71F94D2EFFF43723FF55F8EBFE1BE22F" authority="Harris, 2014" authorityName="Harris" authorityYear="2014" box="[212,460,1862,1883]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="170" phylum="Arthropoda" rank="species" species="verrucosum" status="sp. nov.">
<emphasis id="848DEABFFFF43723FF55F8EBFE1BE22F" box="[212,460,1862,1883]" italics="true" pageId="4" pageNumber="170">Cereudorsum verrucosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FBE57C4FFF43723FE53F8EBFDCBE22F" box="[466,540,1862,1883]" pageId="4" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
Female: 
<emphasis id="848DEABFFFF43723FDF2F8EBFD46E22F" box="[627,657,1862,1883]" italics="true" pageId="4" pageNumber="170">(A)</emphasis>
antenna; 
<emphasis id="848DEABFFFF43723FD6CF8EBFCDBE22F" box="[749,780,1862,1883]" italics="true" pageId="4" pageNumber="170">(B)</emphasis>
maxillule; 
<emphasis id="848DEABFFFF43723FCF9F8EBFC40E22F" box="[888,919,1862,1883]" italics="true" pageId="4" pageNumber="170">(C)</emphasis>
maxillipeds (note, coxae do not touch); 
<emphasis id="848DEABFFFF43723FA96F8EBFAEFE22F" box="[1303,1336,1862,1883]" italics="true" pageId="4" pageNumber="170">(D)</emphasis>
maxilla; 
<emphasis id="848DEABFFFF43723FFF9F8CCFF40E202" box="[120,151,1889,1910]" italics="true" pageId="4" pageNumber="170">(E)</emphasis>
P5 and detail of border setules; 
<emphasis id="848DEABFFFF43723FE4FF8CCFE3AE202" box="[462,493,1889,1910]" italics="true" pageId="4" pageNumber="170">(F)</emphasis>
P3; 
<emphasis id="848DEABFFFF43723FD98F8CCFDEDE202" box="[537,570,1889,1910]" italics="true" pageId="4" pageNumber="170">(G)</emphasis>
P4; 
<emphasis id="848DEABFFFF43723FDE6F8CCFD5FE202" box="[615,648,1889,1910]" italics="true" pageId="4" pageNumber="170">(H)</emphasis>
maxilliped; 
<emphasis id="848DEABFFFF43723FC83F8CCFCCDE202" box="[770,794,1889,1910]" italics="true" pageId="4" pageNumber="170">(I)</emphasis>
genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm.
</paragraph>
</caption>
<caption id="E2866625FFF53722FF27F8EBFE88E2E4" ID-DOI="http://doi.org/10.5281/zenodo.4684567" ID-Zenodo-Dep="4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="5" pageNumber="171" startId="5.[166,229,1862,1884]" targetBox="[228,1404,153,1804]" targetPageId="5">
<paragraph id="B64636ADFFF53722FF27F8EBFE88E2E4" blockId="5.[166,1467,1861,1936]" pageId="5" pageNumber="171">
Figure 3. 
<taxonomicName id="71F94D2EFFF53722FE83F8EBFE2CE22F" authority="Harris, 2014" authorityName="Harris" authorityYear="2014" box="[258,507,1862,1883]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="171" phylum="Arthropoda" rank="species" species="verrucosum" status="sp. nov.">
<emphasis id="848DEABFFFF53722FE83F8EBFE2CE22F" box="[258,507,1862,1883]" italics="true" pageId="5" pageNumber="171">Cereudorsum verrucosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FBE57C4FFF53722FD80F8EBFD9BE22F" box="[513,588,1862,1883]" pageId="5" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
Male: 
<emphasis id="848DEABFFFF53722FD0FF8EBFD7AE22F" box="[654,685,1862,1883]" italics="true" pageId="5" pageNumber="171">(A)</emphasis>
caudal ramus; 
<emphasis id="848DEABFFFF53722FCBEF8EBFCADE22F" box="[831,890,1862,1883]" italics="true" pageId="5" pageNumber="171">(B, D)</emphasis>
P5 ventral view (pits on dorsal side); 
<emphasis id="848DEABFFFF53722FB67F8EBFAD1E22F" box="[1254,1286,1862,1883]" italics="true" pageId="5" pageNumber="171">(C)</emphasis>
P2; 
<emphasis id="848DEABFFFF53722FAB3F8EBFA87E22F" box="[1330,1360,1862,1883]" italics="true" pageId="5" pageNumber="171">(E)</emphasis>
antennule; 
<emphasis id="848DEABFFFF53722FF27F8CCFF08E202" box="[166,223,1889,1910]" italics="true" pageId="5" pageNumber="171">(F, G)</emphasis>
details of coupling denticles and dactylus. Female: 
<emphasis id="848DEABFFFF53722FD50F8CCFD24E202" box="[721,755,1889,1910]" italics="true" pageId="5" pageNumber="171">(H)</emphasis>
rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm.
</paragraph>
</caption>
<paragraph id="B64636ADFFF63721FFF8F932FB62E3BB" blockId="6.[121,1422,1695,1744]" pageId="6" pageNumber="172">
Plate 1. Scanning electron micrographs of 
<taxonomicName id="71F94D2EFFF63721FD92F932FCDBE3C1" authorityName="Harris" authorityYear="2014" box="[531,780,1695,1717]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="172" phylum="Arthropoda" rank="species" species="verrucosum">
<emphasis id="848DEABFFFF63721FD92F932FCDBE3C1" box="[531,780,1695,1717]" italics="true" pageId="6" pageNumber="172">Cereudorsum verrucosum</emphasis>
</taxonomicName>
. 
<emphasis id="848DEABFFFF63721FC99F932FCE0E3C0" box="[792,823,1695,1716]" italics="true" pageId="6" pageNumber="172">(A)</emphasis>
adult female (dorsal, note caudal rami and P5 limbs missing); 
<emphasis id="848DEABFFFF63721FFF8F917FF63E3BB" box="[121,180,1722,1743]" italics="true" pageId="6" pageNumber="172">(B, C)</emphasis>
honeycomb-like cuticle; 
<emphasis id="848DEABFFFF63721FE29F917FE1EE3BB" box="[424,457,1722,1743]" italics="true" pageId="6" pageNumber="172">(D)</emphasis>
dorsal pits. Scale bar: A = 
<quantity id="71019B48FFF63721FD4DF917FCF3E3BB" box="[716,804,1722,1743]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="172" unit="mm" value="0.13">0.13 mm</quantity>
. B = 
<quantity id="71019B48FFF63721FCD6F917FC6CE3BB" box="[855,955,1722,1744]" metricMagnitude="-5" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="172" unit="mm" value="0.016">0.016 mm</quantity>
. C = 
<quantity id="71019B48FFF63721FC6FF914FBE2E3BB" box="[1006,1077,1721,1743]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="172" unit="mm" value="8.5">8.5 µm</quantity>
. D = 
<quantity id="71019B48FFF63721FBEBF917FB66E3BB" box="[1130,1201,1722,1743]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="172" unit="mm" value="3.2">3.2 µm</quantity>
.
</paragraph>
<paragraph id="B64636ADFFF73720FF27F905FED1E387" blockId="7.[166,1466,1704,1779]" pageId="7" pageNumber="173">
Plate 2. Scanning electron micrographs of 
<taxonomicName id="71F94D2EFFF73720FDC0F905FCEDE3CA" authorityName="Harris" authorityYear="2014" box="[577,826,1704,1726]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="173" phylum="Arthropoda" rank="species" species="verrucosum">
<emphasis id="848DEABFFFF73720FDC0F905FCEDE3CA" box="[577,826,1704,1726]" italics="true" pageId="7" pageNumber="173">Cereudorsum verrucosum</emphasis>
</taxonomicName>
. 
<emphasis id="848DEABFFFF73720FCC7F905FCB2E3C9" box="[838,869,1704,1725]" italics="true" pageId="7" pageNumber="173">(A)</emphasis>
right “shoulder” region of female cephalosome; 
<emphasis id="848DEABFFFF73720FABCF905FA8CE3C9" box="[1341,1371,1704,1725]" italics="true" pageId="7" pageNumber="173">(B)</emphasis>
border of cephalosome showing blocks of micro-tubules; 
<emphasis id="848DEABFFFF73720FDF5F96EFD65E3AC" box="[628,690,1731,1752]" italics="true" pageId="7" pageNumber="173">(C, D)</emphasis>
detail of microtubules. Scale bar: A = 
<quantity id="71019B48FFF73720FBA6F96FFB5CE3AC" box="[1063,1163,1730,1752]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="173" unit="mm" value="0.045">0.045 mm</quantity>
. B = 
<quantity id="71019B48FFF73720FB3EF96EFAF3E3AC" box="[1215,1316,1731,1752]" metricMagnitude="-5" metricUnit="m" metricValue="2.1999999999999997" pageId="7" pageNumber="173" unit="mm" value="0.022">0.022 mm</quantity>
. C = 
<quantity id="71019B48FFF73720FAD9F96EFA48E3AC" box="[1368,1439,1731,1753]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="7" pageNumber="173" unit="mm" value="6.4">6.4 µm</quantity>
. D = 
<quantity id="71019B48FFF73720FF3BF970FED6E387" box="[186,257,1757,1779]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="173" unit="mm" value="3.2">3.2 µm</quantity>
.
</paragraph>
<caption id="E2866625FFF8372FFFF8F887FC14E2B1" ID-DOI="http://doi.org/10.5281/zenodo.4684573" ID-Zenodo-Dep="4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="8" pageNumber="174" startId="8.[121,184,1834,1856]" targetBox="[197,1336,172,1783]" targetPageId="8">
<paragraph id="B64636ADFFF8372FFFF8F887FC14E2B1" blockId="8.[121,1422,1834,1989]" pageId="8" pageNumber="174">
Figure 4. 
<taxonomicName id="71F94D2EFFF8372FFF56F887FE06E234" authority="Harris, 2014" authorityName="Harris" authorityYear="2014" box="[215,465,1834,1856]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="174" phylum="Arthropoda" rank="species" species="verrucosum" status="sp. nov.">
<emphasis id="848DEABFFFF8372FFF56F887FE06E234" box="[215,465,1834,1856]" italics="true" pageId="8" pageNumber="174">Cereudorsum verrucosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FBE57C4FFF8372FFE56F887FDF5E24B" box="[471,546,1834,1855]" pageId="8" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="848DEABFFFF8372FFDA9F887FD90E24B" box="[552,583,1834,1855]" italics="true" pageId="8" pageNumber="174">(A)</emphasis>
P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; 
<emphasis id="848DEABFFFF8372FFE55F8E8FE24E22E" box="[468,499,1861,1882]" italics="true" pageId="8" pageNumber="174">(B)</emphasis>
stage V female copepodid; 
<emphasis id="848DEABFFFF8372FFD7CF8E8FCCBE22E" box="[765,796,1861,1882]" italics="true" pageId="8" pageNumber="174">(C)</emphasis>
lateral view of adult male; 
<emphasis id="848DEABFFFF8372FFBA3F8E8FB94E22E" box="[1058,1091,1861,1882]" italics="true" pageId="8" pageNumber="174">(D)</emphasis>
border of male cephalosome, note absence of micro-tubules (
<emphasis id="848DEABFFFF8372FFEF6F8F2FE7CE200" box="[375,427,1887,1908]" italics="true" pageId="8" pageNumber="174">hy.m.</emphasis>
hyaline membrane; 
<emphasis id="848DEABFFFF8372FFDECF8F2FD73E200" box="[621,676,1887,1908]" italics="true" pageId="8" pageNumber="174">hy.so.</emphasis>
hyaline sense organ; 
<emphasis id="848DEABFFFF8372FFCF0F8F2FC70E200" box="[881,935,1887,1908]" italics="true" pageId="8" pageNumber="174">mg.d.</emphasis>
duct of marginal gland; 
<emphasis id="848DEABFFFF8372FFB11F8CDFB10E201" box="[1168,1223,1888,1909]" italics="true" pageId="8" pageNumber="174">mg.o.</emphasis>
opening of marginal duct); 
<emphasis id="848DEABFFFF8372FFF38F8D7FF0FE2FB" box="[185,216,1914,1935]" italics="true" pageId="8" pageNumber="174">(E)</emphasis>
border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for 
<emphasis id="848DEABFFFF8372FFE35F838FE12E2DE" box="[436,453,1941,1962]" italics="true" pageId="8" pageNumber="174">D</emphasis>
; 
<emphasis id="848DEABFFFF8372FFE53F838FDD1E2DE" box="[466,518,1941,1962]" italics="true" pageId="8" pageNumber="174">d.pits</emphasis>
, dorsal pits); 
<emphasis id="848DEABFFFF8372FFD0CF838FD7CE2DE" box="[653,683,1941,1962]" italics="true" pageId="8" pageNumber="174">(F)</emphasis>
dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar: A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm.
</paragraph>
</caption>
<caption id="E2866625FFF9372EFF27F8FDFE8CE2EF" ID-DOI="http://doi.org/10.5281/zenodo.4684577" ID-Zenodo-Dep="4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="9" pageNumber="175" startId="9.[166,229,1872,1894]" targetBox="[239,1403,166,1825]" targetPageId="9">
<paragraph id="B64636ADFFF9372EFF27F8FDFE8CE2EF" blockId="9.[166,1468,1872,1947]" pageId="9" pageNumber="175">
Figure 5. 
<taxonomicName id="71F94D2EFFF9372EFE86F8FDFDD5E212" authority="Harris, 2014" authorityName="Harris" authorityYear="2014" box="[263,514,1872,1894]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="175" phylum="Arthropoda" rank="species" species="verrucosum" status="sp. nov.">
<emphasis id="848DEABFFFF9372EFE86F8FDFDD5E212" box="[263,514,1872,1894]" italics="true" pageId="9" pageNumber="175">Cereudorsum verrucosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FBE57C4FFF9372EFD8AF8FCFD8FE212" box="[523,600,1873,1894]" pageId="9" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
Male: 
<emphasis id="848DEABFFFF9372EFD1EF8FCFD6AE212" box="[671,701,1873,1894]" italics="true" pageId="9" pageNumber="175">(A)</emphasis>
rostrum and “shoulder” (ventral); 
<emphasis id="848DEABFFFF9372EFB94F8FCFBE3E212" box="[1045,1076,1873,1894]" italics="true" pageId="9" pageNumber="175">(F)</emphasis>
genital double-somite and caudal rami. Female: 
<emphasis id="848DEABFFFF9372EFF7BF8C6FECFE2F4" box="[250,280,1899,1920]" italics="true" pageId="9" pageNumber="175">(B)</emphasis>
P2; 
<emphasis id="848DEABFFFF9372EFEC4F8C6FEB2E2F4" box="[325,357,1899,1920]" italics="true" pageId="9" pageNumber="175">(C)</emphasis>
mandible and molar process; 
<emphasis id="848DEABFFFF9372EFD06F8C6FD7FE2F4" box="[647,680,1899,1920]" italics="true" pageId="9" pageNumber="175">(D)</emphasis>
labrum; 
<emphasis id="848DEABFFFF9372EFD7EF8C6FCCAE2F4" box="[767,797,1899,1920]" italics="true" pageId="9" pageNumber="175">(E)</emphasis>
P1. Scale bar: A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm.
</paragraph>
</caption>
<paragraph id="B64636ADFFFA372DFFF8FF3AFE57E42B" blockId="10.[119,751,151,498]" pageId="10" pageNumber="176">
<emphasis id="848DEABFFFFA372DFFF8FF3AFEF5E5C4" bold="true" box="[121,290,151,176]" pageId="10" pageNumber="176">Biometric data</emphasis>
. 
<emphasis id="848DEABFFFFA372DFEAEFF35FE5EE5C4" box="[303,393,152,176]" italics="true" pageId="10" pageNumber="176">Females</emphasis>
(N = 6): maximum length (
<subScript id="2A7D34E8FFFA372DFD31FF35FD34E5C5" attach="left" box="[688,739,152,177]" fontSize="8" pageId="10" pageNumber="176">Lmax</subScript>
) mean 
<quantity id="71019B48FFFA372DFF3CFF18FEF5E5B8" box="[189,290,181,205]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="10" pageNumber="176" unit="mm" value="0.92">0.92 mm</quantity>
, range 
<quantity id="71019B48FFFA372DFEF7FF18FDC0E5B8" box="[374,535,181,205]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.8" metricValueMin="9.0" pageId="10" pageNumber="176" unit="mm" value="0.94" valueMax="0.98" valueMin="0.9">0.90–0.98 mm</quantity>
, body length (
<subScript id="2A7D34E8FFFA372DFD3BFF18FD33E5BB" attach="left" box="[698,740,181,207]" fontSize="8" pageId="10" pageNumber="176">Lurs</subScript>
) mean 
<quantity id="71019B48FFFA372DFF3AFF7FFEC8E59E" box="[187,287,210,234]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="176" unit="mm" value="0.85">0.85 mm</quantity>
, range 
<quantity id="71019B48FFFA372DFEEEFF7FFDD8E59E" box="[367,527,210,234]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" metricValueMax="8.6" metricValueMin="8.2" pageId="10" pageNumber="176" unit="mm" value="0.84" valueMax="0.86" valueMin="0.82">0.82–0.86 mm</quantity>
; cephalosome width (W) 
<quantity id="71019B48FFFA372DFF2BFF42FEDFE473" box="[170,264,239,263]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="176" unit="mm" value="0.65">0.65 mm</quantity>
, range 
<quantity id="71019B48FFFA372DFECEFF42FE3FE473" box="[335,488,239,263]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="6.6" metricValueMin="6.3" pageId="10" pageNumber="176" unit="mm" value="0.645" valueMax="0.66" valueMin="0.63">0.63–0.66 mm</quantity>
, length 
<quantity id="71019B48FFFA372DFDB7FF5DFD43E473" box="[566,660,240,264]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="10" pageNumber="176" unit="mm" value="0.47">0.47 mm</quantity>
; rostrum width (R) 
<quantity id="71019B48FFFA372DFF64FEA0FE9FE450" box="[229,328,269,293]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="176" unit="mm" value="0.13">0.13 mm</quantity>
; genital double-somite width 
<quantity id="71019B48FFFA372DFD04FEA0FD31E450" box="[645,742,269,293]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="176" unit="mm" value="0.35">0.35 mm</quantity>
, length 
<quantity id="71019B48FFFA372DFF47FE87FEFBE436" box="[198,300,298,322]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="176" unit="mm" value="0.2">0.20 mm</quantity>
; caudal ramus width 
<quantity id="71019B48FFFA372DFD9CFE87FD57E436" box="[541,640,298,322]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="176" unit="mm" value="0.08">0.08 mm</quantity>
(3/4 down ramus), length 
<quantity id="71019B48FFFA372DFE9BFEEAFEABE42B" box="[282,380,327,351]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="176" unit="mm" value="0.16">0.16 mm</quantity>
.
</paragraph>
<paragraph id="B64636ADFFFA372DFF16FEC8FDF6E486" blockId="10.[119,751,151,498]" pageId="10" pageNumber="176">
<materialsCitation id="06913CF0FFFA372DFF16FEC8FDD0E486" collectionCode="R" collectorName="Lurs, W" longitude="-2.0" pageId="10" pageNumber="176" specimenCount="1">
Ratios: 
<subScript id="2A7D34E8FFFA372DFF70FEC8FECCE40B" attach="left" box="[241,283,357,383]" fontSize="8" pageId="10" pageNumber="176">Lurs</subScript>
/ 
<geoCoordinate id="D3CD506AFFFA372DFEA4FEC8FEB5E409" box="[293,354,357,381]" degrees="1.3" direction="west" orientation="longitude" pageId="10" pageNumber="176" precision="5555" value="-1.3">W1.3</geoCoordinate>
, 
<subScript id="2A7D34E8FFFA372DFEF3FEC8FE71E40B" attach="right" box="[370,422,357,383]" fontSize="8" pageId="10" pageNumber="176">Lmax</subScript>
/ 
<geoCoordinate id="D3CD506AFFFA372DFE2EFEC8FE39E409" box="[431,494,357,381]" degrees="1.4" direction="west" orientation="longitude" pageId="10" pageNumber="176" precision="5555" value="-1.4">W1.4</geoCoordinate>
, cephalosome 55% of 
<collectorName id="1B0C537BFFFA372DFFF8FE2EFF10E4EE" box="[121,199,386,412]" pageId="10" pageNumber="176">
<subScript id="2A7D34E8FFFA372DFFF8FE2EFF76E4E8" attach="left" box="[121,161,387,412]" fontSize="8" pageId="10" pageNumber="176">Lurs</subScript>
, W
</collectorName>
/ 
<collectionCode id="D0E8AE68FFFA372DFF51FE2FFF35E4EE" box="[208,226,386,410]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="176">R</collectionCode>
5.0; genital double-somite w/l 1.75, width 54% of cephalosome, anterior lobe 77% of lateral border, posterior lobe 23% of lateral border; caudal ramus 19% of 
<subScript id="2A7D34E8FFFA372DFD12FE10FD6CE4A3" attach="left" box="[659,699,445,471]" fontSize="8" pageId="10" pageNumber="176">Lurs</subScript>
, l/ 
<geoCoordinate id="D3CD506AFFFA372DFD58FE10FF49E486" degrees="2.0" direction="west" orientation="longitude" pageId="10" pageNumber="176" precision="5555" value="-2.0">w 2.0</geoCoordinate>
, Hicks’ index for α 86%, for β 60
</materialsCitation>
%.
</paragraph>
<paragraph id="B64636ADFFFA372DFFF8FDA8FF08E7DB" blockId="10.[119,749,517,836]" pageId="10" pageNumber="176">
<emphasis id="848DEABFFFFA372DFFF8FDA8FF6CE769" box="[121,187,517,541]" italics="true" pageId="10" pageNumber="176">Males</emphasis>
(N =8): maximum length (
<subScript id="2A7D34E8FFFA372DFE72FDA8FDFFE76B" attach="left" box="[499,552,517,543]" fontSize="8" pageId="10" pageNumber="176">Lmax</subScript>
) 
<quantity id="71019B48FFFA372DFDBAFDA8FD75E768" box="[571,674,517,541]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="10" pageNumber="176" unit="mm" value="0.72">0.72 mm</quantity>
, range 
<quantity id="71019B48FFFA372DFFF8FD8FFECBE74E" box="[121,284,546,570]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.6" metricValueMin="7.0" pageId="10" pageNumber="176" unit="mm" value="0.73" valueMax="0.76" valueMin="0.7">0.70–0.76 mm</quantity>
; cephalosome width (W) 
<quantity id="71019B48FFFA372DFDBCFD8FFD75E74E" box="[573,674,546,570]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="176" unit="mm" value="0.55">0.55 mm</quantity>
, range 
<quantity id="71019B48FFFA372DFFF8FD92FEC2E723" box="[121,277,575,599]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.699999999999999" metricValueMin="4.9" pageId="10" pageNumber="176" unit="mm" value="0.53" valueMax="0.57" valueMin="0.49">0.49–0.57 mm</quantity>
, length 
<quantity id="71019B48FFFA372DFEE4FDEDFE11E723" box="[357,454,576,600]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="176" unit="mm" value="0.32">0.32 mm</quantity>
; rostrum (ventral) 
<quantity id="71019B48FFFA372DFD09FDEDFD3FE723" box="[648,744,576,600]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="10" pageNumber="176" unit="mm" value="0.07">0.07 mm</quantity>
; caudal ramus width 
<quantity id="71019B48FFFA372DFECAFDF0FE6EE700" box="[331,441,605,629]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="176" unit="mm" value="0.055">0.055 mm</quantity>
, length 
<quantity id="71019B48FFFA372DFD8AFDF0FDADE700" box="[523,634,605,629]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="176" unit="mm" value="0.065">0.065 mm</quantity>
; antennule length (fully extended) 
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; spermatophore 0.18 × 
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.
</paragraph>
<paragraph id="B64636ADFFFA372DFF16FD18FDBCE636" blockId="10.[119,749,517,836]" pageId="10" pageNumber="176">
Ratios: 
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/ 
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, cephalosome 43% of 
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, W/R 7.8; caudal ramus l/ 
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, Hicks’ index for α 68%, for β 60%; antennule 27% of bodY length, antennule segment 2 34%, segments 3+4 34%, dactYlus 24%, aesthetasc 78% of antennule length; spermatophore 25% of 
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.
</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFFA372DFFF8FCF8FAE6E2E4" pageId="10" pageNumber="176" type="description">
<paragraph id="B64636ADFFFA372DFFF8FCF8FC09E149" blockId="10.[119,751,853,1933]" lastBlockId="10.[792,1424,151,1936]" pageId="10" pageNumber="176">
<emphasis id="848DEABFFFFA372DFFF8FCF8FF2AE619" bold="true" box="[121,253,853,877]" pageId="10" pageNumber="176">Description</emphasis>
. 
<emphasis id="848DEABFFFFA372DFE8FFCF8FE73E619" box="[270,420,853,877]" italics="true" pageId="10" pageNumber="176">Adult females</emphasis>
(
<figureCitation id="2EC22A28FFFA372DFE37FCF8FDD9E619" box="[438,526,853,877]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1A</figureCitation>
): colourless or very pale amber, heavily sclerotized regions brown. Body outline elliptical, truncated anteriorly with convex bulge above rostrum, rostrum (
<figureCitation id="2EC22A28FFFA372DFEB5FC00FE53E6B1" box="[308,388,941,965]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Fig. 3H</figureCitation>
) partly obscured by anterior bulge. Hyaline border clear, 
<quantity id="71019B48FFFA372DFED8FC67FE12E697" box="[345,453,970,995]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="10" pageNumber="176" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 µm</quantity>
wide. Rows of conspicuous pits (
<quantity id="71019B48FFFA372DFF2DFC45FF34E174" box="[172,227,1000,1024]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="176" unit="mm" value="3.0">3 µm</quantity>
) surrounded by rim of thickened cuticle ornament dorsal surface of cephalosome, metasome, genital doublesomite and P5s (Plate 1D, p. 172), conspicuous cuticular ridge runs parallel to edge of cephalosome and medial to the blocks of microtubules (
<figureCitation id="2EC22A28FFFA372DFEA2FBF0FEA4E101" box="[291,371,1117,1141]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1A</figureCitation>
; Plate 2B, p. 173). Numerous (&gt; 100) sensilla form a regular pattern on the dorsal surface of the cephalosome, metasome segments and genital double-somite (Plate 1A, p. 172; 
<figureCitation id="2EC22A28FFFA372DFEBAFB18FE5AE1B9" box="[315,397,1205,1229]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1C</figureCitation>
). Some resemble setae, but most are short tubular structures with a basal collar that project through holes in the cuticle (Plate 2B,C, p. 173). Two massive cuticular outgrowths resembling honeycomb 
<quantity id="71019B48FFFA372DFDCDFAA0FD6EE051" box="[588,697,1293,1317]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="10" pageNumber="176" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 µm</quantity>
high (
<figureCitation id="2EC22A28FFFA372DFFFEFA87FF02E036" box="[127,213,1322,1346]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1A</figureCitation>
, 
<figureCitation id="2EC22A28FFFA372DFF62FA87FED2E036" box="[227,261,1322,1346]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="10" pageNumber="176">4C</figureCitation>
; Plate 1A,B,C, p. 172) are present on dorsal midline of cephalosome. Individual cells range from 
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in diameter and have very thin walls (&lt;
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). Similar, but smaller, cuticular outgrowths are present on metasome 3 and genital double-somite. On either side of anterior half of cephalosome there are eleven blocks of microtubules close to hyaline border (
<figureCitation id="2EC22A28FFFA372DFEFDFA77FE03E086" box="[380,468,1498,1522]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Figs 1A</figureCitation>
, 
<figureCitation id="2EC22A28FFFA372DFE61FA77FE29E086" box="[480,510,1498,1522]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="10" pageNumber="176">4E</figureCitation>
left side; Plate 2A–D, p. 173). Each block contains 80–100 tubules and each tubule is 
<quantity id="71019B48FFFA372DFF10F9B8FF3AE359" box="[145,237,1557,1581]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="176" unit="mm" value="0.75">0.75 µm</quantity>
in diameter, 
<quantity id="71019B48FFFA372DFEFAF9B8FE1FE359" box="[379,456,1557,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="176" unit="mm" value="2.5">2.5 µm</quantity>
high and has a lumen 
<quantity id="71019B48FFFA372DFD3FF9B8FF4AE33F" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="176" unit="mm" value="0.15">0.15 µm</quantity>
in diameter (Plate 2D, p. 173). Labrum without ridge plates, posterior edge with three or four setules at lateral corner (
<figureCitation id="2EC22A28FFFA372DFF4AF9C0FEC8E3F1" box="[203,287,1645,1669]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">Fig. 5D</figureCitation>
). Genital double-somite about half width of cephalosome (
<figureCitation id="2EC22A28FFFA372DFE8EF927FE89E3D6" box="[271,350,1674,1698]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1C</figureCitation>
, note that width is distorted in drawing by pressure of coverglass), a distinct notch marks boundary between anterior and posterior lobes, one sensillum mid-way along edge of anterior lobe, posterior lobe about 1/4 length of genital double-somite, (in normal view posterior lobe appears pointed, but when laid flat it is rounded, 
<figureCitation id="2EC22A28FFFA372DFD9FF8B0FDB9E241" box="[542,622,1821,1845]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1C</figureCitation>
), setules on posterior lobe smaller than those on anterior lobe (
<figureCitation id="2EC22A28FFFA372DFD0FF897FD08E226" box="[654,735,1850,1874]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1E</figureCitation>
). Dorsal surface pitted, many seta-like sensilla present plus area of honeycomb cuticle in midline. Posterior arch deep, surrounds about 1/2 of caudal furca. Genital opening straight (
<figureCitation id="2EC22A28FFFA372DFCA3FF18FCA7E5B9" box="[802,880,181,205]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2I</figureCitation>
). Metasome segment 4 with fimbriate setules on posterior edge of sternum. Caudal rami broad, elongate (
<figureCitation id="2EC22A28FFFA372DFAE2FF7FFCE2E47C" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1F</figureCitation>
), medial edge straight with setules down length, lateral edge slightly convex with few setules distally. Seta T1 small pinnate, deeply recessed, posterior border slightly convex with three very large pinnate setae (T2, T3 and T4) evenly spaced (
<figureCitation id="2EC22A28FFFA372DFCEEFEC8FC17E409" box="[879,960,357,381]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1D</figureCitation>
). Dorsal surface with two longitudinal ridges and some net-like markings, α and β setae verY long (about 1/2 length of ramus).Seta on first segment of antennule pinnate. Structure and setation of mouthparts and ambulatory limbs typical of family. Basis of antenna with row of triangular setules (
<figureCitation id="2EC22A28FFFA372DFCEEFE5AFC14E764" box="[879,963,503,528]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2A</figureCitation>
), exopod with six plumulose setae, segment 2 of endopod with three lateral setae, geniculate setae plain, claw comb-like. Mandible with small group of setules on anterior lobe (
<figureCitation id="2EC22A28FFFA372DFC33FDE2FBD1E713" box="[946,1030,591,615]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">Fig. 5C</figureCitation>
). Maxillule with single bulbous seta on exopod (
<figureCitation id="2EC22A28FFFA372DFC15FDC0FC32E7F1" box="[916,997,621,645]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2B</figureCitation>
). Maxilla as in 
<figureCitation id="2EC22A28FFFA372DFB01FDC0FB06E7F1" box="[1152,1233,621,645]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2D</figureCitation>
. Maxilliped coxae do not meet in midline (
<figureCitation id="2EC22A28FFFA372DFB8FFD27FB88E7D6" box="[1038,1119,650,674]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2C</figureCitation>
), coxa with fimbriate border, fimbriate process greatlY reduced in size and represented bY a bunch of fine fimbriate setules (
<figureCitation id="2EC22A28FFFA372DFBDDFD68FB78E7A9" box="[1116,1199,709,733]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2H</figureCitation>
). Endopod of P1 with small area of denticulate setules at lateral end of fimbriate crescent (
<figureCitation id="2EC22A28FFFA372DFCFFFD52FC18E663" box="[894,975,767,791]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">Fig. 5E</figureCitation>
). P2 endopod with strong proximal setules on segment 1, serrulate spinous seta on segment 3 2/3 length of endopod (
<figureCitation id="2EC22A28FFFA372DFC1BFC97FC3CE626" box="[922,1003,826,850]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">Fig. 5B</figureCitation>
). Serrate spinous seta on segment 2 of P3 endopod (
<figureCitation id="2EC22A28FFFA372DFC01FCFAFC1BE604" box="[896,972,855,880]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2F</figureCitation>
) shorter than endopod (0.7:1), large serrate spinous seta on segment 3 very much longer than endopod (1.7:1). P4 endopod segment 1 without internal seta (
<figureCitation id="2EC22A28FFFA372DFAE5FC3FFCEAE6BC" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2G</figureCitation>
), seta on segment 2 and first internal seta of segment 3 thin, straight spinous. Exopod of P5 lanceolate (
<figureCitation id="2EC22A28FFFA372DFAADFC60FA57E691" box="[1324,1408,973,997]" captionStart="Figure 2" captionStartId="4.[121,184,1862,1884]" captionTargetBox="[207,1351,176,1821]" captionTargetId="figure-10@4.[121,1421,151,1830]" captionTargetPageId="4" captionText="Figure 2. Cereudorsum verrucosum sp. nov. Female: (A) antenna; (B) maxillule; (C) maxillipeds (note, coxae do not touch); (D) maxilla; (E) P5 and detail of border setules; (F) P3; (G) P4; (H) maxilliped; (I) genital opening. Scale bar: A, C = 0.14 mm. B = 0.08 mm. D, H, I = 0.1 mm. E = 0.3 mm. F, G = 0.23 mm." figureDoi="http://doi.org/10.5281/zenodo.4684565" httpUri="https://zenodo.org/record/4684565/files/figure.png" pageId="10" pageNumber="176">Fig. 2E</figureCitation>
), dorsal surface with rows of pits, one sub-terminal seta and two apical setae, border setules long (
<quantity id="71019B48FFFA372DFB20FBAAFB3EE154" box="[1185,1257,1031,1056]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="176" unit="mm" value="25.0">25 µm</quantity>
). Females carry 
<specimenCount id="A0FFFD24FFFA372DFC98FB88FCBDE149" box="[793,874,1061,1085]" pageId="10" pageNumber="176" type="egg">10 eggs</specimenCount>
per brood.
</paragraph>
<paragraph id="B64636ADFFFA372DFCB9FBEFFAE6E2E4" blockId="10.[792,1424,151,1936]" pageId="10" pageNumber="176">
<emphasis id="848DEABFFFFA372DFCB9FBEFFC61E12E" box="[824,950,1090,1114]" italics="true" pageId="10" pageNumber="176">Adult males</emphasis>
(
<figureCitation id="2EC22A28FFFA372DFC45FBEFFBC1E12E" box="[964,1046,1090,1114]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="10" pageNumber="176">Fig. 1B</figureCitation>
). Outline of cephalosome a sharply truncated hemi-ellipse, convex in midline above rostrum, rostrum rounded (
<figureCitation id="2EC22A28FFFA372DFC52FBD0FBF2E1E1" box="[979,1061,1149,1173]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">Fig. 5A</figureCitation>
), lateral angle of antennule sockets project forward, shoulder with epaulet and several cuticular serrations (in ventral view, 
<figureCitation id="2EC22A28FFFA372DFBC2FB1AFB4EE1BB" box="[1091,1177,1207,1231]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">Fig. 5A</figureCitation>
), hyaline border starts at epaulet. No microtubules on border of cephalosome (
<figureCitation id="2EC22A28FFFA372DFAE5FB78FCEBE07E" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="10" pageNumber="176">Fig. 4D</figureCitation>
), ducts from marginal glands open individually dorsal to hyaline border. Colour, pits, ridges, sensilla and massive dorsal cuticular honeycomb as described for female. Caudal rami rectangular (
<figureCitation id="2EC22A28FFFA372DFC5AFAE7FBE2E016" box="[987,1077,1354,1378]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Figs 3A</figureCitation>
, 
<figureCitation id="2EC22A28FFFA372DFBC2FAE7FBB5E016" box="[1091,1122,1354,1378]" captionStart="Figure 5" captionStartId="9.[166,229,1872,1894]" captionTargetBox="[239,1403,166,1825]" captionTargetId="figure-8@9.[166,1466,151,1833]" captionTargetPageId="9" captionText="Figure 5. Cereudorsum verrucosum sp. nov. Male: (A) rostrum and “shoulder” (ventral); (F) genital double-somite and caudal rami. Female: (B) P2; (C) mandible and molar process; (D) labrum; (E) P1. Scale bar:A = 0.22 mm. B, C = 0.18 mm. D = 0.1 m.m. E = 0.15 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684577" httpUri="https://zenodo.org/record/4684577/files/figure.png" pageId="10" pageNumber="176">5F</figureCitation>
), lateral edge convex, with setules at posterior end, posterior border slightly convex with fine setules, setules down medial edge, dorsal surface with longitudinal ridge and net-like markings, α and β setae very long (almost length of ramus), inserted very close together about 1/3 waY down ramus. Terminal seta T1 small, pinnate, recessed at lateral corner, T2, T3 and T4 large pinnate setae evenly spaced. Antennule (
<figureCitation id="2EC22A28FFFA372DFB59F9BAFAFDE344" box="[1240,1322,1559,1584]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Fig. 3E</figureCitation>
) without denticle on segment 3, four or five closelY grouped finger-like processes (not cuticular denticles) on segment 4 (
<figureCitation id="2EC22A28FFFA372DFAAFF9FFFA57E31E" box="[1326,1408,1618,1642]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Fig. 3F</figureCitation>
), dactylus shorter than segment 3+4, with small hook at end of segment 5 (Figs, 3E, G), aesthetasc long (about 3/4 length of antennule), divided into three sections by two constrictions. Terminal segment of P2 endopod with two plumose setae and one serrulate spinous seta (
<figureCitation id="2EC22A28FFFA372DFBF1F948FB12E389" box="[1136,1221,1765,1789]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Fig. 3C</figureCitation>
). Segment 1 of P4 endopod without internal seta, setae on segments 2 and 3 plumose (not spinous). P5 exopod acutely trapezoid (apical angle 50°) with rows of pits on dorsal surface (
<figureCitation id="2EC22A28FFFA372DFB7EF890FA86E221" box="[1279,1361,1853,1877]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Fig. 3B</figureCitation>
), first (lateral) seta with five strong ventral setules, each terminal seta with two or three setules at its base (
<figureCitation id="2EC22A28FFFA372DFB4FF8DAFAF3E2E4" box="[1230,1316,1911,1936]" captionStart="Figure 3" captionStartId="5.[166,229,1862,1884]" captionTargetBox="[228,1404,153,1804]" captionTargetId="figure-8@5.[166,1466,151,1819]" captionTargetPageId="5" captionText="Figure 3. Cereudorsum verrucosum sp. nov. Male: (A) caudal ramus; (B, D) P5 ventral view (pits on dorsal side); (C) P2; (E) antennule; (F, G) details of coupling denticles and dactylus. Female: (H) rostrum (ventral). Scale bar: A = 0.1 mm. B, C, E, H = 0.14 mm. F = 0.06 mm. G = 0.08 mm." figureDoi="http://doi.org/10.5281/zenodo.4684567" httpUri="https://zenodo.org/record/4684567/files/figure.png" pageId="10" pageNumber="176">Fig. 3D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFFB372CFF27FF35FE6AE0E9" pageId="11" pageNumber="177" type="discussion">
<paragraph id="B64636ADFFFB372CFF27FF35FDF0E78E" blockId="11.[165,796,151,1437]" pageId="11" pageNumber="177">
<emphasis id="848DEABFFFFB372CFF27FF35FEC6E5C4" bold="true" box="[166,273,152,176]" pageId="11" pageNumber="177">Remarks</emphasis>
. This species is remarkable for a number of unusual features. Nothing corresponding to the dorsal organs and their cuticular outgrowths has been found on any other member of the 
<taxonomicName id="71F94D2EFFFB372CFE6BFF42FD52E473" authorityName="Boeck" authorityYear="1865" box="[490,645,239,263]" class="Maxillopoda" family="Porcellidiidae" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="177" phylum="Arthropoda" rank="family">Porcellidiidae</taxonomicName>
. They do not appear to be derived from any ancestral or pre-existing structure. TheY first appear on stage III copepodids, but after metamorphosis to adult the trough is completely obscured by the massive honeycomb cuticle. These structures occur on both sexes and first appear on stage III copepodids in a simple form. They are best interpreted from juvenile stages because the mass of cuticular honeycomb is shorter and does not obscure the dorsal organs (
<figureCitation id="2EC22A28FFFB372CFDB9FE77FD58E486" box="[568,655,474,498]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">Fig. 4B</figureCitation>
). The dorsal organ on stage V copepodids appears as a longitudinal trough with thickened rim (
<figureCitation id="2EC22A28FFFB372CFE5CFDB8FDE4E759" box="[477,563,533,557]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">Fig. 4F</figureCitation>
). This is surrounded by a clear oval area which is bounded by a ridge and pits, similar to the ridge parallel to the edge of the cephalosome but greatly extended in height to form the folds of the cuticular honeycomb. In adults the honeycomb is massive and completely obscures the dorsal trough (
<figureCitation id="2EC22A28FFFB372CFD0CFD0AFD32E7B4" box="[653,741,679,704]" captionStart="Figure 1" captionStartId="3.[166,229,1867,1889]" captionTargetBox="[243,1386,160,1829]" captionTargetId="figure-8@3.[166,1466,151,1833]" captionTargetPageId="3" captionText="Figure 1. Cereudorsum verrucosum sp. nov. Female: (A) adult; (C, E) genital double-somite (E detail of posterior lobe); (D, F) caudal rami (D detail of terminal setae). Male: (B) adult. Scale bar:A, B = 0.5 mm. C = 0.03 mm. D = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684563" httpUri="https://zenodo.org/record/4684563/files/figure.png" pageId="11" pageNumber="177">Fig. 1A</figureCitation>
, 
<figureCitation id="2EC22A28FFFB372CFD73FD05FCC2E7B4" box="[754,789,680,704]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">4C</figureCitation>
; Plate 1A, p. 172). The function of the dorsal organ and cuticular honeycomb is not known.
</paragraph>
<paragraph id="B64636ADFFFB372CFF44FCADFCCCE149" blockId="11.[165,796,151,1437]" pageId="11" pageNumber="177">
Ducts from marginal glands of male animals open individually close to the hyaline border as in other porcellidiid species (
<figureCitation id="2EC22A28FFFB372CFE10FC97FE3BE626" box="[401,492,826,850]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">Fig. 4D</figureCitation>
), but similar ducts are not found in female animals in the region of microtubule blocks (
<figureCitation id="2EC22A28FFFB372CFF2CFCD8FF29E6F9" box="[173,254,885,909]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">Fig. 4E</figureCitation>
, left hand side). From the body cavity passages run through the thickened cuticular border of the cephalosome and end in fine branches under the blocks of microtubules (
<figureCitation id="2EC22A28FFFB372CFF2FFC60FF28E691" box="[174,255,973,997]" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">Fig. 4E</figureCitation>
, right hand side). It is assumed that secretions from the marginal glands exit through the microtubules. Blocks of microtubules are not found on juvenile animals, but first appear at the moult from stage V copepodid to adult female.
</paragraph>
<paragraph id="B64636ADFFFB372CFF44FBEFFD2EE07E" blockId="11.[165,796,151,1437]" pageId="11" pageNumber="177">
The caudal rami of copepodid stages bear typical thin pinnate setae. The large pinnate setae with hollow shaft do not appear until the final moult to the adult stage (
<figureCitation id="2EC22A28FFFB372CFD6FFBD0FF11E1C6" captionStart="Figure 4" captionStartId="8.[121,184,1834,1856]" captionTargetBox="[197,1336,172,1783]" captionTargetId="figure-10@8.[121,1421,151,1786]" captionTargetPageId="8" captionText="Figure 4. Cereudorsum verrucosum sp. nov. (A) P5 and caudal ramus of pharate stage V female copepodid, showing difference in type of setae between adult and juvenile; (B) stage V female copepodid; (C) lateral view of adult male; (D) border of male cephalosome, note absence of micro-tubules (hy.m. hyaline membrane; hy.so. hyaline sense organ; mg.d. duct of marginal gland; mg.o. opening of marginal duct); (E) border of female cephalosome (left superficial focus showing micro-tubules, right deep focus showing branched ducts of marginal glands, lettering as for D; d.pits, dorsal pits); (F) dorsal organs and cuticular honeycomb of stage V copepodid (dorsal view). Scale bar:A, F = 0.14 mm. B = 0.55 mm. C = 0.35 mm. D, E = 0.08 mm. F = 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.4684573" httpUri="https://zenodo.org/record/4684573/files/figure.png" pageId="11" pageNumber="177">Fig. 4A</figureCitation>
shows a pharate stage V female copepodid with both juvenile and adult terminal setae). Unlike the clavate setae of 
<taxonomicName id="71F94D2EFFFB372CFF46FB78FE9DE199" authorityName="Harris &amp; Iwasaki" authorityYear="1996" box="[199,330,1237,1261]" class="Hexanauplia" family="Porcellidiidae" genus="Clavigofera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="848DEABFFFFB372CFF46FB78FE9DE199" box="[199,330,1237,1261]" italics="true" pageId="11" pageNumber="177">Clavigofera</emphasis>
</taxonomicName>
, the pinnae do not originate from a thin lateral expansion of the shaft, but from the shaft itself.
</paragraph>
<paragraph id="B64636ADFFFB372CFF44FAA2FE6AE0E9" blockId="11.[165,796,151,1437]" pageId="11" pageNumber="177">The male antennules are also unique for theY have fingerlike structures on segment 4 that appear to take the place of coupling denticles found on all other species. They may perform the same function of increasing friction during mate guarding behaviour.</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFFB372CFF27FA1FFD31E09C" pageId="11" pageNumber="177" type="etymology">
<paragraph id="B64636ADFFFB372CFF27FA1FFD31E09C" blockId="11.[166,793,1458,1512]" pageId="11" pageNumber="177">
<emphasis id="848DEABFFFFB372CFF27FA1FFEF0E0BE" bold="true" box="[166,295,1458,1482]" pageId="11" pageNumber="177">Etymology</emphasis>
. The specific name refers to the wart-like appearance of the dorsal organs (L. 
<emphasis id="848DEABFFFFB372CFDA4FA7DFDAFE093" box="[549,632,1488,1511]" italics="true" pageId="11" pageNumber="177">verruca</emphasis>
= a wart).
</paragraph>
</subSubSection>
<subSubSection id="FEE36526FFFB372CFCC6FF3AFB06E4CC" pageId="11" pageNumber="177" type="distribution">
<paragraph id="B64636ADFFFB372CFCC6FF3AFB06E4CC" blockId="11.[837,1467,151,440]" pageId="11" pageNumber="177">
<emphasis id="848DEABFFFFB372CFCC6FF3AFC18E5DB" bold="true" box="[839,975,151,175]" pageId="11" pageNumber="177">Distribution</emphasis>
. 
<taxonomicName id="71F94D2EFFFB372CFC5BFF3AFB3FE5DB" authorityName="Harris" authorityYear="2014" box="[986,1256,151,175]" class="Hexanauplia" family="Porcellidiidae" genus="Cereudorsum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="177" phylum="Arthropoda" rank="species" species="verrucosum">
<emphasis id="848DEABFFFFB372CFC5BFF3AFB3FE5DB" box="[986,1256,151,175]" italics="true" pageId="11" pageNumber="177">Cereudorsum verrucosum</emphasis>
</taxonomicName>
has only been found on partly decomposed 
<taxonomicName id="71F94D2EFFFB372CFBAAFF18FB0EE5B9" box="[1067,1241,181,205]" class="Phaeophyceae" family="Lessoniaceae" genus="Ecklonia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Laminariales" pageId="11" pageNumber="177" phylum="Ochrophyta" rank="species" species="radiata">
<emphasis id="848DEABFFFFB372CFBAAFF18FB0EE5B9" box="[1067,1241,181,205]" italics="true" pageId="11" pageNumber="177">Ecklonia radiata</emphasis>
</taxonomicName>
in muddy water at the entrance to Gunnamatta Bay, Port Hacking. This is unusual for it is verY rare to find porcellidiids on rotting algae and only one or two species can tolerate muddy water. Nearly all the animals in the sample are covered with diatoms, suctorian protozoa, moulds and sand particles, which make critical observation of detail difficult. The 
<typeStatus id="6942880FFFFB372CFB41FEC8FB39E408" box="[1216,1262,357,380]" pageId="11" pageNumber="177">tYpe</typeStatus>
series, Cr34. 2/77, contained 
<specimenCount id="A0FFFD24FFFB372CFC34FE2FFC29E4EE" box="[949,1022,386,410]" pageId="11" pageNumber="177" type="female">82 ♀♀</specimenCount>
, 
<specimenCount id="A0FFFD24FFFB372CFB8BFE2FFB85E4EE" box="[1034,1106,386,410]" pageId="11" pageNumber="177" type="male">54 ♂♂</specimenCount>
(8 coupled to female copepodids) and 8 copepodids. V. A. Harris 1977.
</paragraph>
</subSubSection>
</treatment>
</document>