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<document id="BECE9C238F70400170A65754A21FB942" ID-CLB-Dataset="26066" ID-DOI="10.11646/zootaxa.4652.1.3" ID-GBIF-Dataset="8acfb4ea-c855-4bf4-beba-7fad6e4ffd4a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3363573" ID-ZooBank="9E0BB4A0-B4F5-4ADA-A107-B187AD14465F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565270355150" checkinUser="plazi" docAuthor="Polizei, Thiago T. S. &amp; Barclay, Maxwell V. L." docDate="2019" docId="03D0D1714168FFBDFF5AC4CEFE9A0896" docLanguage="en" docName="zootaxa.4652.1.3.pdf" docOrigin="Zootaxa 4652 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cylloepus segurae Polizei &amp; Barclay 2019, sp. nov." docType="treatment" docUuid="367CFDC0-ED3E-445F-915C-ABD96C8D150E" docUuidSource="ZooBank" docVersion="11" lastPageNumber="99" masterDocId="FFE9A909416CFFBBFFCDC633FFAE0A65" masterDocTitle="The genus Cylloepus Erichson, 1847 (Coleoptera: Elmidae: Elminae) new species and combinations" masterLastPageNumber="100" masterPageNumber="93" pageNumber="97" updateTime="1698749095823" updateUser="ExternalLinkService">
<mods:mods id="4DFBA9862FFBCF17C9F4DA775B3B4E7D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="68E5568A7E7572FA77335911A1E6F9BE">
<mods:title id="DCEFBDAE1D465E0475201C32234837D9">The genus Cylloepus Erichson, 1847 (Coleoptera: Elmidae: Elminae) new species and combinations</mods:title>
</mods:titleInfo>
<mods:name id="1F66CCF4BCF89E435B9B1EE65F6FFD89" type="personal">
<mods:role id="62B8AD098977AD61F3816A846FB43E0F">
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<mods:namePart id="D065371099EA22F9E41DB366CB0D1F20">Polizei, Thiago T. S.</mods:namePart>
</mods:name>
<mods:name id="5F4E178C8F6E08EFF8418133C780C117" type="personal">
<mods:role id="575CBE80E1A2F96B9D4854BEEE355789">
<mods:roleTerm id="07A20FEF0B015D514251672D15284E5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27A345D6F200833D9C225DCE651C6974">Barclay, Maxwell V. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="EDFC4BF32FFFAA66181F91686D84DC97">text</mods:typeOfResource>
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<mods:title id="2C8EEB96DAE6B24E68D54942E4F6AF12">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5F4592A4E9A0F78DAF7BD059FF2F302F">
<mods:date id="D227B9AD51B36E42276BBD38DA71924B">2019</mods:date>
<mods:detail id="18EFD13EADCA5CF23D41342B7D8AC261" type="pubDate">
<mods:number id="A1466BD023C686633EAE2C8D999EC978">2019-08-07</mods:number>
</mods:detail>
<mods:detail id="8C76E0A0C0B6FF38AE118FB56D0746FF" type="volume">
<mods:number id="4FC42F976A497307D0D27D0E03B93620">4652</mods:number>
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<mods:number id="74FF28BE67043E93236DDF654BAB99A8">1</mods:number>
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<mods:start id="A5E6A57F85C349C5FD806A24EA36B57B">93</mods:start>
<mods:end id="418A49B01FF6F9995710D2119719EC4E">100</mods:end>
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<mods:classification id="EDDA5128B8A701F1DB96D358284A1766">journal article</mods:classification>
<mods:identifier id="34C7B48351064647608544D3DA24F5A9" type="CLB-Dataset">26066</mods:identifier>
<mods:identifier id="81677DF4D471CA0CA6F8F19F3497E30C" type="DOI">10.11646/zootaxa.4652.1.3</mods:identifier>
<mods:identifier id="B98A483C3178692FB28AEA187BAF96CD" type="GBIF-Dataset">8acfb4ea-c855-4bf4-beba-7fad6e4ffd4a</mods:identifier>
<mods:identifier id="C7C4DF34D1C097B1DCEDEDBFF94E2981" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0DC965A99D3A4D5A01B57CF569227832" type="Zenodo-Dep">3363573</mods:identifier>
<mods:identifier id="18394CAA184304FC53194DC72717A00D" type="ZooBank">9E0BB4A0-B4F5-4ADA-A107-B187AD14465F</mods:identifier>
</mods:mods>
<treatment id="03D0D1714168FFBDFF5AC4CEFE9A0896" ID-DOI="http://doi.org/10.5281/zenodo.5933300" ID-GBIF-Taxon="159043976" ID-Zenodo-Dep="5933300" LSID="urn:lsid:zoobank.org:act:367CFDC0-ED3E-445F-915C-ABD96C8D150E" httpUri="http://treatment.plazi.org/id/03D0D1714168FFBDFF5AC4CEFE9A0896" lastPageId="6" lastPageNumber="99" pageId="4" pageNumber="97">
<subSubSection id="C36333EC4168FFBFFF5AC4CEFD120972" box="[151,700,765,792]" pageId="4" pageNumber="97" type="nomenclature">
<paragraph id="8BC660674168FFBFFF5AC4CEFD120972" blockId="4.[151,984,765,863]" box="[151,700,765,792]" pageId="4" pageNumber="97">
<heading id="D08ED70B4168FFBFFF5AC4CEFD120972" bold="true" box="[151,700,765,792]" fontSize="11" level="1" pageId="4" pageNumber="97" reason="1">
<emphasis id="B90DBC754168FFBFFF5AC4CEFD120972" bold="true" box="[151,700,765,792]" pageId="4" pageNumber="97">
<taxonomicName id="4C791BE44168FFBFFF5AC4CEFDF90972" authority="Polizei &amp; Barclay" authorityName="Polizei &amp; Barclay" authorityYear="2019" box="[151,599,765,792]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="segurae" status="sp. nov.">
<emphasis id="B90DBC754168FFBFFF5AC4CEFEDA0972" bold="true" box="[151,372,765,792]" italics="true" pageId="4" pageNumber="97">Cylloepus segurae</emphasis>
Polizei &amp; Barclay
</taxonomicName>
<taxonomicNameLabel id="A23E010E4168FFBFFD93C4CEFD120972" box="[606,700,765,791]" pageId="4" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36333EC4168FFBFFF5AC512FC76093A" pageId="4" pageNumber="97" type="description">
<paragraph id="8BC660674168FFBFFF5AC512FEB1095E" blockId="4.[151,984,765,863]" box="[151,287,801,827]" pageId="4" pageNumber="97">
(
<figureCitation id="13427CE24168FFBFFF52C512FF75095E" box="[159,219,801,827]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="4" pageNumber="97">Fig 2</figureCitation>
A–C)
</paragraph>
<paragraph id="8BC660674168FFBFFF5AC577FC76093A" blockId="4.[151,984,765,863]" box="[151,984,836,863]" pageId="4" pageNumber="97">
<uri id="FFE86C654168FFBFFF5AC577FC76093A" box="[151,984,836,863]" pageId="4" pageNumber="97">
urn:lsid:zoobank.org:act:
<uuid id="FFDF5AB24168FFBFFE65C577FC76093A" box="[424,984,836,863]" pageId="4" pageNumber="97">367CFDC0-ED3E-445F-915C-ABD96C8D150E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C36333EC4168FFBDFF5AC5BEFE9A0896" lastPageId="6" lastPageNumber="99" pageId="4" pageNumber="97" type="materials_examined">
<paragraph id="8BC660674168FFBFFF5AC5BEFD8E09C2" blockId="4.[151,1437,908,2015]" box="[151,544,908,935]" pageId="4" pageNumber="97">
<emphasis id="B90DBC754168FFBFFF5AC5BEFE9809C2" bold="true" box="[151,310,908,935]" pageId="4" pageNumber="97">Type locality.</emphasis>
<collectingRegion id="49BDAE854168FFBFFEF1C5BEFE1509C2" box="[316,443,909,935]" country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="97">Santa Cruz</collectingRegion>
,
<collectingCountry id="F36E20F74168FFBFFE0BC5BEFDB209C2" box="[454,540,909,935]" name="Bolivia" pageId="4" pageNumber="97">Bolivia</collectingCountry>
.
</paragraph>
<paragraph id="8BC660674168FFBFFF0AC582FD5C0E77" blockId="4.[151,1437,908,2015]" pageId="4" pageNumber="97">
<emphasis id="B90DBC754168FFBFFF0AC582FEAA09AE" bold="true" box="[199,260,945,971]" pageId="4" pageNumber="97">Male</emphasis>
<materialsCitation id="3B116A3A4168FFBFFEC3C582FD5C0E77" ID-GBIF-Occurrence="2313474995" collectingDate="2004-11-20" collectingDateMax="2004-12-12" collectingDateMin="2004-11-20" collectionCode="MV, BMNH" collectorName="Barclay, M. V. L. &amp; Mendel, H." country="Bolivia" elevation="1000" latitude="-18.1" location="Los Volcanes" longLatPrecision="1276" longitude="-63.6" municipality="Amboro National Park" pageId="4" pageNumber="97" specimenCode="NHMUK010583885" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="54C2DEC54168FFBFFEC3C582FEDB09AE" box="[270,373,945,971]" pageId="4" pageNumber="97" type="holotype">
<emphasis id="B90DBC754168FFBFFEC3C582FEDB09AE" bold="true" box="[270,373,945,971]" pageId="4" pageNumber="97">holotype</emphasis>
</typeStatus>
:
<specimenCount id="9D7FABEE4168FFBFFE4BC582FE3509AE" box="[390,411,945,971]" pageId="4" pageNumber="97" type="male">♂</specimenCount>
|
<typeStatus id="54C2DEC54168FFBFFE78C582FDB309AE" box="[437,541,945,971]" pageId="4" pageNumber="97" type="holotype">Holotype</typeStatus>
|
<collectingCountry id="F36E20F74168FFBFFDFAC582FD0109AE" box="[567,687,945,971]" name="Bolivia" pageId="4" pageNumber="97">BOLIVIA</collectingCountry>
:
<collectingRegion id="49BDAE854168FFBFFD71C582FC9409AE" box="[700,826,945,971]" country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="97">Santa Cruz</collectingRegion>
/
<collectingMunicipality id="6BA2FA1D4168FFBFFC98C582FBF409AE" box="[853,1114,945,971]" pageId="4" pageNumber="97">Amboro National Park</collectingMunicipality>
/
<location id="8EA636BC4168FFBFFBBBC582FABC09AE" LSID="urn:lsid:plazi:treatment:03D0D1714168FFBDFF5AC4CEFE9A0896:8EA636BC4168FFBFFBBBC582FABC09AE" box="[1142,1298,945,971]" country="Bolivia" latitude="-18.1" longLatPrecision="1276" longitude="-63.6" municipality="Amboro National Park" name="Los Volcanes" pageId="4" pageNumber="97" stateProvince="Santa Cruz">Los Volcanes</location>
, c.
<elevation id="005487544168FFBFFAF0C582FA2509AE" box="[1341,1419,945,971]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="97" unit="m" value="1000.0">
<quantity id="4C81CD824168FFBFFAF0C582FA2509AE" box="[1341,1419,945,971]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="97" unit="m" value="1000.0">1000m</quantity>
</elevation>
/
<geoCoordinate id="EE4D06A04168FFBFFF5AC5E6FEAF098A" box="[151,257,981,1007]" degrees="18" direction="south" minutes="06" orientation="latitude" pageId="4" pageNumber="97" precision="925" value="-18.1">S 18º 06’</geoCoordinate>
:
<geoCoordinate id="EE4D06A04168FFBFFED4C5E6FE2A098A" box="[281,388,981,1007]" degrees="63" direction="west" minutes="36" orientation="longitude" pageId="4" pageNumber="97" precision="925" value="-63.6">W63º 36’</geoCoordinate>
/
<collectingDate id="EF83BF4F4168FFBFFE50C5E6FDCA098A" box="[413,612,981,1007]" pageId="4" pageNumber="97" value="2004-11-20" valueMax="2004-12-12" valueMin="2004-11-20">
<date id="FFC746A74168FFBFFE50C5E6FDCA098A" box="[413,612,981,1007]" pageId="4" pageNumber="97" value="2004-11-20" valueMax="2004-12-12" valueMin="2004-11-20">20.xi–12.xii.2004</date>
</collectingDate>
|
<collectionCode id="ED68F8A24168FFBFFDB7C5E6FD09098A" box="[634,679,981,1007]" country="0" name="University of Montana Museum" pageId="4" pageNumber="97">MV</collectionCode>
Light Sheet / on stream beach /
<collectorName id="268C05B14168FFBFFBD2C5E6FB7A098A" box="[1055,1236,981,1007]" pageId="4" pageNumber="97">Barclay, M.V.L.</collectorName>
/ &amp;
<collectorName id="268C05B14168FFBFFAC6C5E6FA22098A" box="[1291,1420,981,1007]" pageId="4" pageNumber="97">Mendel, H.</collectorName>
/
<collectionCode id="ED68F8A24168FFBFFF5AC5CBFF400E77" box="[151,238,1016,1042]" pageId="4" pageNumber="97">BMNH</collectionCode>
(E)2004-280 |
<specimenCode id="DBDFC81C4168FFBFFE43C5C4FDD40E77" box="[398,634,1015,1042]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="97" type="Museum">NHMUK010583885</specimenCode>
. (
<collectionCode id="ED68F8A24168FFBFFD41C5CBFD480E77" box="[652,742,1016,1042]" pageId="4" pageNumber="97">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC660674168FFBFFF0AC229FACF0E51" blockId="4.[151,1437,908,2015]" box="[199,1377,1049,1077]" pageId="4" pageNumber="97">
<materialsCitation id="3B116A3A4168FFBFFF0AC229FC680E51" ID-GBIF-Occurrence="2313475000" box="[199,966,1050,1077]" collectingDate="2004-11-20" collectingDateMax="2004-12-12" collectingDateMin="2004-11-20" collectionCode="BMNH" collectorName="Barclay, M. V. L. &amp; Mendel, H." country="Bolivia" elevation="1000" latitude="-18.1" location="Los Volcanes" longLatPrecision="1276" longitude="-63.6" municipality="Amboro National Park" pageId="4" pageNumber="97" specimenCount="3" stateProvince="Santa Cruz" typeStatus="paratype">
<emphasis id="B90DBC754168FFBFFF0AC229FEE60E51" bold="true" box="[199,328,1050,1076]" pageId="4" pageNumber="97">
<typeStatus id="54C2DEC54168FFBFFF0AC229FEED0E51" box="[199,323,1050,1076]" pageId="4" pageNumber="97" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D7FABEE4168FFBFFE82C229FE770E51" box="[335,473,1050,1076]" pageId="4" pageNumber="97" type="generic">3 specimens</specimenCount>
with same data as the holotype. 1 (
<collectionCode id="ED68F8A24168FFBFFCAEC229FC130E51" box="[867,957,1050,1076]" pageId="4" pageNumber="97">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B116A3A4168FFBFFC1CC229FAF30E51" ID-GBIF-Occurrence="2313474998" box="[977,1373,1049,1076]" collectionCode="MZUSP" country="Bolivia" pageId="4" pageNumber="97" specimenCode="MZSP 35379" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
2 (
<collectionCode id="ED68F8A24168FFBFFC20C229FBE00E51" box="[1005,1102,1050,1076]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="97">MZUSP</collectionCode>
) (
<specimenCode id="DBDFC81C4168FFBFFBA9C229FB510E51" box="[1124,1279,1049,1076]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="97" type="Museum">MZSP 35379</specimenCode>
; 35380)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC660674168FFBFFF0AC20FFEEE0F42" blockId="4.[151,1437,908,2015]" pageId="4" pageNumber="97">
<emphasis id="B90DBC754168FFBFFF0AC20FFEEC0E32" bold="true" box="[199,322,1084,1111]" pageId="4" pageNumber="97">Diagnosis.</emphasis>
Body elongate, slightly convex. Head with granules; antenna with short antennomeres. Pronotum slightly wider than long, covered with long golden setae; deep longitudinal impression on the disc. Elytra with the anterior margin yellow and two yellow patches near the apex. Pair of flat spines at the apex of each tibia. Abdomen with a pair of carinae on the first ventrite. Male genitalia with the phallobase slightly shorter than parameres; parameres slightly compressed dorsoventrally, wider at the base, strongly narrowed towards apex, apex sharply curved towards the inner margin; penis with the basal two thirds broad, almost parallel sided, and then tapering towards the sharp apex.
</paragraph>
<paragraph id="8BC660674168FFBFFF0AC31CFD220FEA" blockId="4.[151,1437,908,2015]" pageId="4" pageNumber="97">
<emphasis id="B90DBC754168FFBFFF0AC31CFEF70F2F" bold="true" box="[199,345,1327,1354]" pageId="4" pageNumber="97">Description.</emphasis>
Male. Maximum length
<quantity id="4C81CD824168FFBFFDA2C303FD600F2F" box="[623,718,1328,1354]" metricMagnitude="-3" metricUnit="m" metricValue="2.84" pageId="4" pageNumber="97" unit="mm" value="2.84">2.84mm</quantity>
, maximum width
<quantity id="4C81CD824168FFBFFC55C31CFC560F2F" box="[920,1016,1327,1354]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="97" unit="mm" value="1.15">1.15mm</quantity>
. Body elongate, slightly convex. Pronotum and mouthparts dark-brown colour, with antenna and legs reddish-brown, and elytra brown with the anterior portion and two patches near to apex yellow.
</paragraph>
<paragraph id="8BC660674168FFBFFF0AC3ABFB8E0C7F" blockId="4.[151,1437,908,2015]" pageId="4" pageNumber="97">
<emphasis id="B90DBC754168FFBFFF0AC3ABFEA30FD7" box="[199,269,1432,1458]" italics="true" pageId="4" pageNumber="97">Head:</emphasis>
Partly retractable, without impressions, surface with granules, clearly separated by their own diameter. Eyes projected laterally. Antenna filiform with 11 short antennomeres, the three apical antennomeres with setae (
<figureCitation id="13427CE24168FFBFFF6DC3EEFF570F92" box="[160,249,1501,1527]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="4" pageNumber="97">Fig. 2A</figureCitation>
). Clypeus slightly concave with granules with the same pattern as the head. Frontoclypeal suture distinctly straight. Labrum with the anterior margin rounded, covered by thin golden setae.
</paragraph>
<paragraph id="8BC660674168FFBEFF0AC011FE520BEE" blockId="4.[151,1437,908,2015]" lastBlockId="5.[151,1436,153,539]" lastPageId="5" lastPageNumber="98" pageId="4" pageNumber="97">
<emphasis id="B90DBC754168FFBFFF0AC011FE8F0C59" box="[199,289,1570,1596]" italics="true" pageId="4" pageNumber="97">Thorax:</emphasis>
Pronotum slightly wider than long (
<figureCitation id="13427CE24168FFBFFD77C011FCBA0C59" box="[698,788,1570,1596]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="4" pageNumber="97">Fig. 2A</figureCitation>
), (length:
<quantity id="4C81CD824168FFBFFC4AC011FC480C59" box="[903,998,1570,1596]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="97" unit="mm" value="0.74">0.74mm</quantity>
, width:
<quantity id="4C81CD824168FFBFFB8CC011FB0C0C59" box="[1089,1186,1570,1596]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="97" unit="mm" value="0.82">0.82mm</quantity>
); anterior margin convex; lateral margin sinuous and granulated, anterior angles produced, acute; posterior margin trisinuate. Pronotum surface with punctures and long golden setae. Sublateral carinae sinuous, reaching the anterior and posterior margin, wider on the middle portion. Deep depression on anterior portion of the longitudinal carinae. Longitudinal impression wide and deep on the disc and with a depression on each side of the posterior portion (
<figureCitation id="13427CE24168FFBFFB57C09EFB5B0CA2" box="[1178,1269,1709,1735]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="4" pageNumber="97">Fig. 2A</figureCitation>
). Elytra longer than wide (length:
<quantity id="4C81CD824168FFBFFEA5C0E3FE660C8F" box="[360,456,1744,1770]" metricMagnitude="-3" metricUnit="m" metricValue="2.09" pageId="4" pageNumber="97" unit="mm" value="2.09">2.09mm</quantity>
, width:
<quantity id="4C81CD824168FFBFFDEDC0FCFD2F0C8F" box="[544,641,1743,1770]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="97" unit="mm" value="1.15">1.15mm</quantity>
), anterior margin sinuous, lateral margin straight, apex rounded. Elytral suture elevated. Elytra with ten intervals, surface with rows of dense granules, separated by two rows of long setae on each interval. Brown colour, with the anterior margin yellow, extending for 1/5, and two yellow patches of triangular shape near to apex (
<figureCitation id="13427CE24168FFBFFE1CC10BFD850D37" box="[465,555,1848,1874]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="4" pageNumber="97">Fig. 2A</figureCitation>
). Carinae on the intervals; 3, 4, 6 and 8, on the third interval extending for 1/5, on fourth interval extending half of the elytra, and the sublateral carinae (sixth and eighth intervals), extending for ¾ of elytral length. Epipleura with the anterior-half covered by tomentum (
<figureCitation id="13427CE24168FFBFFC12C14EFB970DF2" box="[991,1081,1917,1943]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="4" pageNumber="97">Fig. 2A</figureCitation>
). Scutellum dark-brown colour, long and narrow, without punctures, with two rows of three golden setae. Hind wings macropterous. Prosternum covered by tomentum and punctures separated by their own diameter (except on the hypomera). Prosternal process U-shaped (
<figureCitation id="13427CE24169FFBEFEDEC6AAFEC30AD6" box="[275,365,153,179]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="5" pageNumber="98">Fig. 2B</figureCitation>
), reaching the coxae, with the surface with punctures and tomentum. Mesoventrite short, with a shallow depression to receive the prosternal process, surface covered by tomentum. Metaventrite broad, moderately pronounced, with the surface similar to the surface of the prosternal process (
<figureCitation id="13427CE24169FFBEFBFBC6D2FB200A9E" box="[1078,1166,225,251]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="5" pageNumber="98">Fig. 2B</figureCitation>
). Legs (except the tarsi) covered by tomentum. Coxae slightly elongate, hind coxae with two rounded depressions; femora with microsetae; tibiae long and thin, with fringe of tomentum on distal ¾, and a pair of long, flat spines at the apex, tarsi long, 5- 5-5, with a few setae at the apex of each tarsomere, and the last tarsomere slightly smaller than the previous four combined. Claws long and thin.
</paragraph>
<paragraph id="8BC660674169FFBEFF0AC7A6FF45087E" blockId="5.[151,1436,153,539]" pageId="5" pageNumber="98">
<emphasis id="B90DBC754169FFBEFF0AC7A6FE940BCA" box="[199,314,405,431]" italics="true" pageId="5" pageNumber="98">Abdomen:</emphasis>
Five ventrites (
<figureCitation id="13427CE24169FFBEFE23C7A6FDE90BCA" box="[494,583,405,431]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="5" pageNumber="98">Fig. 2B</figureCitation>
), (length:
<quantity id="4C81CD824169FFBEFD70C7A6FCB30BCA" box="[701,797,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="98" unit="mm" value="1.24">1.24mm</quantity>
, width:
<quantity id="4C81CD824169FFBEFCB4C7A7FC750BCA" box="[889,987,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="98" unit="mm" value="1.05">1.05mm</quantity>
). Surface covered by tomentum, except the disc of the first and second ventrite, and with punctures scattered randomly up to twice their own diameter. First ventrite carinate on the disc. Fifth with a few setae on the apex, and with lateral margins slightly produced posterolaterad.
</paragraph>
<caption id="DF0630EF4169FFBEFF5AC195FD4C0D86" httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="5" pageNumber="98" startId="5.[151,250,1958,1983]" targetBox="[151,1436,580,1912]" targetPageId="5">
<paragraph id="8BC660674169FFBEFF5AC195FD4C0D86" blockId="5.[151,1436,1958,2019]" pageId="5" pageNumber="98">
<emphasis id="B90DBC754169FFBEFF5AC195FEBC0DDA" bold="true" box="[151,274,1958,1983]" pageId="5" pageNumber="98">FIGURE 2.</emphasis>
<taxonomicName id="4C791BE44169FFBEFED5C194FE7D0DDA" authority="Polizei &amp; Barclay, 2019" authorityName="Polizei &amp; Barclay" authorityYear="2019" box="[280,467,1959,1983]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="segurae" status="sp. nov.">
<emphasis id="B90DBC754169FFBEFED5C194FE7D0DDA" box="[280,467,1959,1983]" italics="true" pageId="5" pageNumber="98">Cylloepus segurae</emphasis>
</taxonomicName>
<emphasis id="B90DBC754169FFBEFE14C194FD870DDA" bold="true" box="[473,553,1959,1983]" pageId="5" pageNumber="98">
<taxonomicNameLabel id="A23E010E4169FFBEFE14C194FD870DDA" box="[473,553,1959,1983]" pageId="5" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm.
</paragraph>
</caption>
<paragraph id="8BC66067416AFFBDFF0AC6AAFC5F0B26" blockId="6.[151,1437,152,755]" pageId="6" pageNumber="99">
<emphasis id="B90DBC75416AFFBDFF0AC6AAFE940AD6" box="[199,314,153,179]" italics="true" pageId="6" pageNumber="99">Genitalia:</emphasis>
(
<figureCitation id="13427CE2416AFFBDFE87C6AAFE0A0AD6" box="[330,420,153,179]" captionStart="FIGURE 2" captionStartId="5.[151,250,1958,1983]" captionTargetBox="[151,1436,580,1912]" captionTargetId="figure@5.[151,1436,580,1912]" captionTargetPageId="5" captionText="FIGURE 2. Cylloepus segurae sp. nov. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." httpUri="https://zenodo.org/record/3363577/files/figure.png" pageId="6" pageNumber="99">Fig. 2C</figureCitation>
); (length:
<quantity id="4C81CD82416AFFBDFDD5C6ABFDD90AD6" box="[536,631,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="6" pageNumber="99" unit="mm" value="0.57">0.57mm</quantity>
, width:
<quantity id="4C81CD82416AFFBDFD1FC6AAFC9A0AD6" box="[722,820,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="6" pageNumber="99" unit="mm" value="0.18">0.18mm</quantity>
): Phallobase twice as long as wide and slightly shorter than parameres. Parameres slightly shorter than penis and slightly compressed dorsoventrally, wider at the basal, strongly narrowed towards apex, apex sharply curved towards the inner margin. Penis with the basal two thirds broad, almost parallel sided, and then tapering towards the sharp apex. Fibula elongate, approximately half the length of penis. Corona present, arrangement of tiny rod-like sclerotizations.
</paragraph>
<paragraph id="8BC66067416AFFBDFF0AC77EFDA40B02" blockId="6.[151,1437,152,755]" box="[199,522,333,359]" pageId="6" pageNumber="99">
<emphasis id="B90DBC75416AFFBDFF0AC77EFE8B0B02" bold="true" box="[199,293,333,359]" pageId="6" pageNumber="99">Female.</emphasis>
Similar to the male.
</paragraph>
<paragraph id="8BC66067416AFFBDFF0AC742FDA60BCA" blockId="6.[151,1437,152,755]" pageId="6" pageNumber="99">
<emphasis id="B90DBC75416AFFBDFF0AC742FEE00BEE" bold="true" box="[199,334,369,395]" pageId="6" pageNumber="99">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C791BE4416AFFBDFDF3C741FD390BEE" authorityName="Polizei &amp; Barclay" authorityYear="2019" box="[574,663,370,395]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="segurae">
<emphasis id="B90DBC75416AFFBDFDF3C741FD390BEE" box="[574,663,370,395]" italics="true" pageId="6" pageNumber="99">segurae</emphasis>
</taxonomicName>
is in honour of Dr. Melissa Ottoboni Segura, one of the pioneers in the study of the elmids of
<collectingCountry id="F36E20F7416AFFBDFE71C7A6FDAA0BCA" box="[444,516,405,431]" name="Brazil" pageId="6" pageNumber="99">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BC66067416AFFBDFF0AC78BFE9A0896" blockId="6.[151,1437,152,755]" pageId="6" pageNumber="99">
<emphasis id="B90DBC75416AFFBDFF0AC78BFE1F0BB6" bold="true" box="[199,433,440,467]" pageId="6" pageNumber="99">Comparative notes.</emphasis>
<taxonomicName id="4C791BE4416AFFBDFE74C78AFD240BB6" authority="Polizei &amp; Barclay, 2019" authorityName="Polizei &amp; Barclay" authorityYear="2019" box="[441,650,441,467]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="segurae" status="sp. nov.">
<emphasis id="B90DBC75416AFFBDFE74C78AFD240BB6" box="[441,650,441,467]" italics="true" pageId="6" pageNumber="99">Cylloepus segurae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E010E416AFFBDFD5CC78AFD450BB6" box="[657,747,441,467]" pageId="6" pageNumber="99" rank="species">
<emphasis id="B90DBC75416AFFBDFD5CC78AFD450BB6" bold="true" box="[657,747,441,467]" pageId="6" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C791BE4416AFFBDFC4DC78AFB580BB6" authority="Hinton, 1940" authorityName="Hinton" authorityYear="1940" box="[896,1270,441,467]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="B90DBC75416AFFBDFC4DC78AFBF40BB6" box="[896,1114,441,467]" italics="true" pageId="6" pageNumber="99">Cylloepus ventralis</emphasis>
Hinton,
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</taxonomicName>
in the antenna with short antennomeres; pronotum with lateral margin sinuous, very deep longitudinal impression on the disc, and tibiae with fringe of tomentum on distal ¾. However it can easily be differentiated by the dark brown colour of the elytra of
<emphasis id="B90DBC75416AFFBDFE89C416FE6B085A" box="[324,453,549,575]" italics="true" pageId="6" pageNumber="99">
<taxonomicName id="4C791BE4416AFFBDFE89C416FE6E085A" authorityName="Polizei &amp; Barclay" authorityYear="2019" box="[324,448,549,575]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="segurae">C. segurae</taxonomicName>
,
</emphasis>
with the yellow anterior margin extending for 1/5 of the elytra, and two yellow patches of triangular shape near to the apex; sublateral carinae on the pronotum reaching the anterior and posterior margin; epipleura with the anterior-half covered by tomentum and disc of the first and second ventrite not covered by tomentum, while
<taxonomicName id="4C791BE4416AFFBDFED5C4A2FE3208CE" authorityName="Hinton" authorityYear="1940" box="[280,412,657,683]" class="Insecta" family="Elmidae" genus="Cylloepus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="B90DBC75416AFFBDFED5C4A2FE3208CE" box="[280,412,657,683]" italics="true" pageId="6" pageNumber="99">C. ventralis</emphasis>
</taxonomicName>
has the elytra uniformly brown; sublateral carinae on pronotum only reaching the anterior margin; epipleura are entirely covered by dense tomentum, and the disc of the first to fourth ventrites is not covered by tomentum.
</paragraph>
</subSubSection>
</treatment>
</document>
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