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<mods:title id="77CE67701EDD50EAAF7FF3A0EC9F604B">Taxonomic revision of the genus Deltepilissus Pereira, 1949 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="766B756A87E698F070A6757BE8B5ED7E">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="DB1C8C609FD3842463F8461D618CB5A7">Universidade Federal Rural de Pernambuco, Unidade Acadêmica de Serra Talhada. Avenida Gregório Ferraz Nogueira, s / n. Serra Talhada, PE, 56909 - 535, Brazil. &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01, Guamá. Belém, PA, 66075 - 110, Brazil.</mods:affiliation>
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<mods:namePart id="A7B218F7254393C56CC99E1FB762F852">Ferreira, Ana B. M.</mods:namePart>
<mods:affiliation id="1185BCDD8E161D2B9F6C5D1EE7F802EF">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01, Guamá. Belém, PA, 66075 - 110, Brazil.</mods:affiliation>
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<mods:namePart id="2642804775E73E2B663CD69276BC249A">Génier, François</mods:namePart>
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<heading id="D0C42C0CFFDAFFAD09C9E2E9FDE3FC01" bold="true" box="[151,618,978,1005]" fontSize="11" level="1" pageId="6" pageNumber="89" reason="1">
<taxonomicName id="4C33E0E3FFDAFFAD09C9E2E9FDE3FC01" ID-CoL="34K63" authority="(Harold, 1880)" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[151,618,978,1005]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="infernalis">
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<emphasis id="B9474772FFDAFFAD09C9E2E9FE24FC00" bold="true" box="[151,429,978,1005]" italics="true" pageId="6" pageNumber="89">Deltepilissus infernalis</emphasis>
(
<bibRefCitation id="EFA2E691FFDAFFAD08E2E2E9FDEBFC01" author="Harold, E." box="[444,610,978,1004]" pageId="6" pageNumber="89" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
)
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<paragraph id="8B8C9B60FFDAFFAD09C9E2CCFD4DFBFC" blockId="6.[151,708,978,1041]" box="[151,708,1014,1041]" pageId="6" pageNumber="89">
(
<figureCitation id="130887E5FFDAFFAD09C1E2CCFE95FBFC" box="[159,284,1015,1041]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Figs 2AL</figureCitation>
,
<figureCitation id="130887E5FFDAFFAD0879E2CCFD92FBFC" box="[295,539,1015,1041]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="6" pageNumber="89">3DF, 3H, 3J, 3L, 3N</figureCitation>
,
<figureCitation id="130887E5FFDAFFAD0B79E2CCFDE4FBFC" box="[551,621,1015,1041]" captionStart="FIGURE 4" captionStartId="10.[151,250,1910,1934]" captionTargetBox="[278,1321,181,1886]" captionTargetId="figure-30@10.[267,1321,181,1886]" captionTargetPageId="10" captionText="FIGURE 4. Type specimens in Deltepilissus and respective label data. AB, Lectotype of Canthon diabolicus Harold, 1880: A, body, dorsal view; B, labels. CD, Lectotype of Canthon infernalis Harold, 1880: C, body, dorsal view; D, labels. EH, holotype of Deltepilissus travassosi Pereira, 1949: E, body, dorsal view; F, head; G, left protibia, dorsal view; H, protibial and tarsal claws. Photographs AD provided by Fernando Z. Vaz-de-Mello." figureDoi="http://doi.org/10.5281/zenodo.6388917" httpUri="https://zenodo.org/record/6388917/files/figure.png" pageId="6" pageNumber="89">4CH</figureCitation>
,
<figureCitation id="130887E5FFDAFFAD0B26E2CDFD35FBFC" box="[632,700,1014,1041]" captionStart="FIGURE 5" captionStartId="11.[151,250,1103,1127]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-19@11.[151,1436,181,1080]" captionTargetPageId="11" captionText="FIGURE 5. Known distribution of species of Deltepilissus. A, Records plotted on terrestrial biomes layer; B, records plotted on elevational range layer." figureDoi="http://doi.org/10.5281/zenodo.6388921" httpUri="https://zenodo.org/record/6388921/files/figure.png" pageId="6" pageNumber="89">5AB</figureCitation>
)
</paragraph>
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<treatmentCitation id="0A92BD71FFDAFFAD09C9E500FD87FBBE" author="Harold, E." box="[151,526,1083,1107]" page="150" pageId="6" pageNumber="89" year="1880">
<taxonomicName id="4C33E0E3FFDAFFAD09C9E500FD87FBBE" ID-CoL="QPCX" authority="Harold, 1880: 150" authorityName="Harold" authorityPageNumber="150" authorityYear="1880" box="[151,526,1083,1107]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDAFFAD09C9E500FEDDFBBE" box="[151,340,1083,1107]" italics="true" pageId="6" pageNumber="89">Canthon infernalis</emphasis>
<bibRefCitation id="EFA2E691FFDAFFAD0807E500FD87FBBE" author="Harold, E." box="[345,526,1083,1107]" pageId="6" pageNumber="89" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold, 1880: 150</bibRefCitation>
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<bibRefCitation id="EFA2E691FFDAFFAD0BADE500FC01FBBE" author="Gillet, J. J. E." box="[755,904,1083,1107]" pageId="6" pageNumber="89" refId="ref8874" refString="Gillet, J. J. E. (1911) Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin, Germany, 100 pp." type="book" year="1911">Gillet 1911: 30</bibRefCitation>
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<bibRefCitation id="EFA2E691FFDAFFAD0AADE500FB2FFBBE" author="Schmidt, A." box="[1011,1190,1083,1107]" pageId="6" pageNumber="89" pagination="61 - 103" refId="ref9438" refString="Schmidt, A. (1922) 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A, 88 (3), 61 - 103." type="journal article" year="1922">Schmidt 1922: 62</bibRefCitation>
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, 73 (taxonomic remarks, distribution);
<treatmentCitation id="0A92BD71FFDAFFAD080CE561FD97FB9F" author="Balthasar, V." box="[338,542,1114,1138]" page="181" pageId="6" pageNumber="89" year="1939">
<bibRefCitation id="EFA2E691FFDAFFAD080CE561FD97FB9F" author="Balthasar, V." box="[338,542,1114,1138]" pageId="6" pageNumber="89" pagination="179 - 238" refId="ref8462" refString="Balthasar, V. (1939) Eine Vorstudie Zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica, 9, 179 - 238." type="journal article" year="1939">Balthasar 1939: 181</bibRefCitation>
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(identification key);
<treatmentCitation id="0A92BD71FFDAFFAD0BAAE561FC68FB9F" author="Blackwelder, R." box="[756,993,1114,1138]" page="199" pageId="6" pageNumber="89" year="1944">
<bibRefCitation id="EFA2E691FFDAFFAD0BAAE561FC68FB9F" author="Blackwelder, R." box="[756,993,1114,1138]" pageId="6" pageNumber="89" pagination="189 - 341" refId="ref8489" refString="Blackwelder, R. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 199</bibRefCitation>
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(checklist).
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<treatmentCitation id="0A92BD71FFDAFFAD09C9E543FDDDFB7C" author="Pereira, F." box="[151,596,1144,1169]" page="231" pageId="6" pageNumber="89" year="1949">
<taxonomicName id="4C33E0E3FFDAFFAD09C9E543FDDDFB7C" ID-CoL="34K64" authority="Pereira, 1949: 231" authorityName="Pereira" authorityPageNumber="231" authorityYear="1949" box="[151,596,1144,1169]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDAFFAD09C9E543FE03FB7D" box="[151,394,1144,1168]" italics="true" pageId="6" pageNumber="89">Deltepilissus travassosi</emphasis>
<bibRefCitation id="EFA2E691FFDAFFAD08CCE543FDDDFB7C" author="Pereira, F." box="[402,596,1144,1169]" pageId="6" pageNumber="89" pagination="231 - 246" refId="ref9277" refString="Pereira, F. (1949) Escarabeideos americanos II (Coleopt. - Scarabaeidae). Arquivos do Museu Paranaense, 7, 231 - 246." type="journal article" year="1949">Pereira, 1949: 231</bibRefCitation>
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, 232, fig. 1 (original description);
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<bibRefCitation id="EFA2E691FFDAFFAD0AE1E543FBF1FB7C" author="Pereira, F." box="[959,1144,1144,1169]" pageId="6" pageNumber="89" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">Pereira 1953: 393</bibRefCitation>
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(synonymized
<taxonomicName id="4C33E0E3FFDAFFAD0C47E543FEA7FB42" ID-CoL="34K64" authority="Pereira, 1949: 231" authorityName="Pereira" authorityPageNumber="231" authorityYear="1949" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDAFFAD0C47E543FEA7FB42" italics="true" pageId="6" pageNumber="89">Deltepilissus travassosi</emphasis>
</taxonomicName>
with
<taxonomicName id="4C33E0E3FFDAFFAD0836E5ACFDAFFB42" ID-CoL="QPCX" authority="Harold, 1880: 150" authorityName="Harold" authorityPageNumber="150" authorityYear="1880" box="[360,550,1175,1199]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDAFFAD0836E5ACFDAFFB42" box="[360,550,1175,1199]" italics="true" pageId="6" pageNumber="89">Canthon infernalis</emphasis>
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).
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<paragraph id="8B8C9B60FFDAFFAD09C9E58DFCC0FAC7" blockId="6.[151,1437,1083,1322]" pageId="6" pageNumber="89">
<treatmentCitationGroup id="AB23BC4EFFDAFFAD09C9E58DFCC0FAC7" pageId="6" pageNumber="89">
<treatmentCitation id="0A92BD71FFDAFFAD09C9E58DFDB6FB23" author="Pereira, F." box="[151,575,1206,1230]" page="393" pageId="6" pageNumber="89" year="1953">
<taxonomicName id="4C33E0E3FFDAFFAD09C9E58DFDB6FB23" ID-CoL="34K63" authority=": Pereira 1953: 393" authorityName=": Pereira" authorityPageNumber="393" authorityYear="1953" box="[151,575,1206,1230]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDAFFAD09C9E58DFE09FB23" box="[151,384,1206,1230]" italics="true" pageId="6" pageNumber="89">Deltepilissus infernalis</emphasis>
:
<bibRefCitation id="EFA2E691FFDAFFAD08D3E58DFDB6FB23" author="Pereira, F." box="[397,575,1206,1230]" pageId="6" pageNumber="89" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">Pereira 1953: 393</bibRefCitation>
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(distribution; transferred
<taxonomicName id="4C33E0E3FFDAFFAD0A1CE58DFC36FB23" ID-CoL="34K63" authorityName=": Pereira" authorityYear="1953" box="[834,959,1206,1230]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDAFFAD0A1CE58DFC36FB23" box="[834,959,1206,1230]" italics="true" pageId="6" pageNumber="89">C. infernalis</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4C33E0E3FFDAFFAD0D1DE58DFB4FFB23" ID-CoL="42SD" authorityName="Pereira" authorityYear="1949" box="[1091,1222,1206,1230]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDAFFAD0D1DE58DFB4FFB23" box="[1091,1222,1206,1230]" italics="true" pageId="6" pageNumber="89">Deltepilissus</emphasis>
</taxonomicName>
);
<treatmentCitation id="0A92BD71FFDAFFAD0D85E58DFEA4FB01" author="Pereira, F. &amp; Martinez, A." page="125" pageId="6" pageNumber="89" year="1956">
<bibRefCitation id="EFA2E691FFDAFFAD0D85E58DFEA4FB01" author="Pereira, F. &amp; Martinez, A." pageId="6" pageNumber="89" pagination="91 - 192" refId="ref9328" refString="Pereira, F. &amp; Martinez, A. (1956) Os generos de Canthonini americanos. Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956: 125</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="0A92BD71FFDAFFAD08C7E5EFFD36FB01" author="Vulcano, M. A. &amp; Pereira, F." box="[409,703,1236,1261]" page="660" pageId="6" pageNumber="89" year="1964">
<bibRefCitation id="EFA2E691FFDAFFAD08C7E5EFFD36FB01" author="Vulcano, M. A. &amp; Pereira, F." box="[409,703,1236,1261]" pageId="6" pageNumber="89" pagination="570 - 685" refId="ref10315" refString="Vulcano, M. A. &amp; Pereira, F. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey, 15, 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 660</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="0A92BD71FFDAFFAD0A72E5EFFBDDFB01" author="Halffter, G. &amp; Martinez, A." box="[812,1108,1236,1261]" page="66" pageId="6" pageNumber="89" year="1977">
<bibRefCitation id="EFA2E691FFDAFFAD0A72E5EFFBDDFB01" author="Halffter, G. &amp; Martinez, A." box="[812,1108,1236,1261]" pageId="6" pageNumber="89" pagination="29 - 107" refId="ref8982" refString="Halffter, G. &amp; Martinez, A. (1977) Revision monografica de los Canthonina Americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana, 38, 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 66</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0A92BD71FFDAFFAD0D8FE5EFFF64FAE6" author="Vaz-de-Mello, F." page="192" pageId="6" pageNumber="89" year="2000">
<bibRefCitation id="EFA2E691FFDAFFAD0D8FE5EFFF64FAE6" author="Vaz-de-Mello, F." pageId="6" pageNumber="89" pagination="183 - 195" refId="ref9891" refString="Vaz-de-Mello, F. (2000) Estado atual de conhecimento dos Scarabaeinae s. str. (Coleoptera: Scarabeoidea) do Brasil. In: Martin- Piera, F., Morrone, J. &amp; Melic, A. (Eds.), Hacia un Proyecto CYTED para el inventario Y Estimacion de la Diversidad Entomologica en Iberoamerica: PrIBES- 2000. Monografias Tercer Milenio &amp; Sociedad Entomologica Aragonesa, Zaragoza, Spain, pp. 183 - 195." type="book chapter" year="2000">Vaz-de-Mello 2000: 192</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0A92BD71FFDAFFAD0830E5C8FD92FAE6" author="Krajcik, M." box="[366,539,1267,1291]" page="47" pageId="6" pageNumber="89" year="2006">
<bibRefCitation id="EFA2E691FFDAFFAD0830E5C8FD92FAE6" author="Krajcik, M." box="[366,539,1267,1291]" pageId="6" pageNumber="89" pagination="1 - 189" refId="ref9158" refString="Krajcik, M. (2006) Checklist of Scarabeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X, Supplement 3, 1 - 189." type="journal article" year="2006">Krajcik 2006: 47</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0A92BD71FFDAFFAD0BC5E5C8FC4DFAE6" author="Vaz-de-Mello, F. &amp; Edmonds, W. &amp; Ocampo, F. &amp; Schoolmeesters, P." box="[667,964,1267,1291]" page="55" pageId="6" pageNumber="89" year="2011">
<bibRefCitation id="EFA2E691FFDAFFAD0BC5E5C8FC4DFAE6" author="Vaz-de-Mello, F. &amp; Edmonds, W. &amp; Ocampo, F. &amp; Schoolmeesters, P." box="[667,964,1267,1291]" pageId="6" pageNumber="89" pagination="1 - 73" refId="ref10018" refString="Vaz-de-Mello, F., Edmonds, W., Ocampo, F. &amp; Schoolmeesters, P. (2011) A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa, 2854 (1), 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="B9474772FFDAFFAD0A70E5CFFCD2FAE6" box="[814,859,1267,1291]" italics="true" pageId="6" pageNumber="89">et al</emphasis>
. 2011: 55
</bibRefCitation>
</treatmentCitation>
, fig. 59 (figure);
<treatmentCitation id="0A92BD71FFDAFFAD0D28E5C8FAAAFAE6" author="Krajcik, M." box="[1142,1315,1267,1291]" page="88" pageId="6" pageNumber="89" year="2012">
<bibRefCitation id="EFA2E691FFDAFFAD0D28E5C8FAAAFAE6" author="Krajcik, M." box="[1142,1315,1267,1291]" pageId="6" pageNumber="89" pagination="1 - 278" refId="ref9194" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. X. Supplement 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 88</bibRefCitation>
</treatmentCitation>
(checklist);
<bibRefCitation id="EFA2E691FFDAFFAD0999E429FE29FAC7" author="Schoolmeesters, P." box="[199,416,1298,1322]" pageId="6" pageNumber="89" pagination="2405 - 8858" refId="ref9492" refString="Schoolmeesters, P. (2020) Scarabs: world Scarabaeidae database (version 2019 - 11 - 02). In: Roskov Y. et al., (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 2020 - 02 - 24. ISSN 2405 - 8858. Species 2000: Naturalis, Leiden, the Netherlands. Available from: http: // www. catalogueoflife. org / col (accessed 12 January 2022)" type="book chapter" year="2020">Schoolmeesters 2020</bibRefCitation>
(catalog);
<bibRefCitation id="EFA2E691FFDAFFAD0B52E429FD58FAC7" author="Vaz-de-Mello, F." box="[524,721,1298,1322]" pageId="6" pageNumber="89" refId="ref9975" refString="Vaz-de-Mello, F. (2021) Scarabaeinae. In: Catalogo Taxonomico da Fauna do Brasil. Available from: http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 127498 (accessed 5 March 2021)" type="url" year="2021">Vaz-de-Mello 2021</bibRefCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDAFFAD09C9E463FEA3F9EF" pageId="6" pageNumber="89" type="diagnosis">
<paragraph id="8B8C9B60FFDAFFAD09C9E463FEA3F9EF" blockId="6.[151,1437,1368,2043]" pageId="6" pageNumber="89">
<emphasis id="B9474772FFDAFFAD09C9E463FE83FA9F" bold="true" box="[151,266,1368,1394]" pageId="6" pageNumber="89">Diagnosis</emphasis>
.
<taxonomicName id="4C33E0E3FFDAFFAD0846E463FD92FA9F" authorityName=": Pereira" authorityYear="1953" box="[280,539,1368,1394]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDAFFAD0846E463FD92FA9F" box="[280,539,1368,1394]" italics="true" pageId="6" pageNumber="89">Deltepilissus infernalis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C33E0E3FFDAFFAD0A14E462FC68FA9F" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[842,993,1368,1394]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFDAFFAD0A14E462FC68FA9F" box="[842,993,1368,1394]" italics="true" pageId="6" pageNumber="89">D. diabolicus</emphasis>
</taxonomicName>
by head approximately 1.4 times wider than long (
<figureCitation id="130887E5FFDAFFAD0851E447FEEFFA7B" box="[271,358,1404,1430]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2E</figureCitation>
); punctures of central portion of pronotum spaced by at least twice their diameter (
<figureCitation id="130887E5FFDAFFAD0C5CE447FA06FA7B" box="[1282,1423,1404,1430]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Figs 2A, 2C</figureCitation>
); body length
<quantity id="4CCB3685FFDAFFAD087BE49BFE1BFA56" box="[293,402,1440,1467]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="6" pageNumber="89" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
; males with denticle on external edge of protibial spur (
<figureCitation id="130887E5FFDAFFAD0D57E49BFBDEFA57" box="[1033,1111,1440,1466]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2I</figureCitation>
), and a conspicuous denticle on inner edge of metatibial spur (
<figureCitation id="130887E5FFDAFFAD0B46E4FFFDFDFA33" box="[536,628,1476,1502]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2K</figureCitation>
); apex of parameres sightly curved inward in dorsal and ventral views (
<figureCitation id="130887E5FFDAFFAD09C1E4D3FE94F9EF" box="[159,285,1512,1538]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="6" pageNumber="89">Figs 3DE</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDAFFAC0999E737FD1DFD6F" lastPageId="7" lastPageNumber="90" pageId="6" pageNumber="89" type="description">
<paragraph id="8B8C9B60FFDAFFAC0999E737FD1DFD6F" blockId="6.[151,1437,1368,2043]" lastBlockId="7.[151,1437,151,2036]" lastPageId="7" lastPageNumber="90" pageId="6" pageNumber="89">
<emphasis id="B9474772FFDAFFAD0999E737FD6EF9CB" bold="true" box="[199,743,1548,1574]" pageId="6" pageNumber="89">Redescription. Colour and tegument sculpture</emphasis>
(
<figureCitation id="130887E5FFDAFFAD0BAAE737FCFDF9CB" box="[756,884,1548,1574]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Figs 2AD</figureCitation>
). Body, in general, black. Head and pronotum with light copper or blue sheen. Body surface densely punctate.
<emphasis id="B9474772FFDAFFAD0A69E70BFC04F9A7" bold="true" box="[823,909,1584,1610]" pageId="6" pageNumber="89">Length</emphasis>
.
<quantity id="4CCB3685FFDAFFAD0AC2E70BFB83F9A6" box="[924,1034,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="6" pageNumber="89" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
.
<emphasis id="B9474772FFDAFFAD0D4BE70BFBDCF9A7" bold="true" box="[1045,1109,1584,1610]" pageId="6" pageNumber="89">Head</emphasis>
(
<figureCitation id="130887E5FFDAFFAD0D38E70BFB37F9A7" box="[1126,1214,1584,1610]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2E</figureCitation>
). Densely punctate. Foveiform punctures approximately equally spaced to their diameter; punctures shallower, more spaced and smaller adjacent to clypeal teeth. Clypeus with two widely spaced, small teeth. Surface slightly depressed behind to clypeal teeth. Clypeo-genal suture visible.
<emphasis id="B9474772FFDAFFAD0B7BE7A7FDF7F95B" bold="true" box="[549,638,1692,1718]" pageId="6" pageNumber="89">Thorax</emphasis>
. Disc of pronotum with dense foveiform punctures spaced by at least twice their diameter; punctures denser on posterocentral portion (
<figureCitation id="130887E5FFDAFFAD0A35E7FBFC7FF937" box="[875,1014,1728,1754]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Figs 2A, 2C</figureCitation>
). Hypomera not excavate, with sparse setose punctures on anterior and posterolateral portions (
<figureCitation id="130887E5FFDAFFAD0A70E7DFFC01F913" box="[814,904,1764,1790]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2F</figureCitation>
). Hypomera with oblique carina not reaching lateral margin; carina about half hypomeral width. Mesoventrite almost smooth, with sparse punctures. Metaventrite punctures variable in size, smaller and denser medially (
<figureCitation id="130887E5FFDAFFAD0A5EE617FCF6F8AB" box="[768,895,1836,1862]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Figs 2BD</figureCitation>
).
<emphasis id="B9474772FFDAFFAD0ACEE617FC55F8AB" bold="true" box="[912,988,1836,1862]" pageId="6" pageNumber="89">Elytra</emphasis>
(
<figureCitation id="130887E5FFDAFFAD0AB4E617FBF9F8AB" box="[1002,1136,1836,1862]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2A, 2C</figureCitation>
). Punctures of elytral striae spaced by once or twice their diameter on 1st, 2nd and 3rd striae; punctures more spaced on lateral striae. Elytral striae wider, carinate margins clearly separated. Seventh stria effaced basally. Eighth and ninth striae almost fused, only visible on posterior half. Elytral interstriae with foveiform punctures spaced at least by once or twice their diameter. Pseudepipleural carina complete, visible on entire length of epipleuron.
<emphasis id="B9474772FFDAFFAD0D66E687FB24F83B" bold="true" box="[1080,1197,1980,2006]" pageId="6" pageNumber="89">Abdomen</emphasis>
(
<figureCitation id="130887E5FFDAFFAD0DE3E687FA90F83B" box="[1213,1305,1980,2006]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="6" pageNumber="89">Fig. 2G</figureCitation>
). Punctures denser on lateral surface of ventrites. Pygidium rounded apically, densely punctate, separated from propygidium by carina (
<figureCitation id="130887E5FFDBFFAC0851E1ACFEE0FF5C" box="[271,361,151,177]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2H</figureCitation>
). Pygidium with sulcus on basal and apical edges.
<emphasis id="B9474772FFDBFFAC0AFAE1ACFC55FF5C" bold="true" box="[932,988,151,177]" pageId="7" pageNumber="90">Legs</emphasis>
. Metatibial spur spatulate; apex acutely angular on inner edge (
<figureCitation id="130887E5FFDBFFAC08C5E180FE7FFF38" box="[411,502,187,213]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2K</figureCitation>
). First meso- and metatarsomeres triangular; tarsomeres 24 trapezoidal, the apical subrectangular (
<figureCitation id="130887E5FFDBFFAC0812E1E4FE4FFF14" box="[332,454,223,249]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2KL</figureCitation>
). Tarsal claws reduced (
<figureCitation id="130887E5FFDBFFAC0B8DE1E4FCC1FF14" box="[723,840,223,249]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Figs 2IL</figureCitation>
).
<emphasis id="B9474772FFDBFFAC0A05E1E5FBCAFF15" bold="true" box="[859,1091,222,248]" pageId="7" pageNumber="90">Sexual dimorphism</emphasis>
. Females can be distinguished from males by last abdominal ventrite evenly wide (
<figureCitation id="130887E5FFDBFFAC0BB1E039FCC5FEF1" box="[751,844,258,284]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2D</figureCitation>
); last abdominal ventrite of males slightly narrower medially (
<figureCitation id="130887E5FFDBFFAC0856E01DFEEBFEAD" box="[264,354,294,320]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2G</figureCitation>
). Females with protibial spur slightly emarginated apically (bifurcate), with a denticle (spiniform process) on inner and external edges (
<figureCitation id="130887E5FFDBFFAC0B10E071FD28FE89" box="[590,673,330,356]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2J</figureCitation>
); protibial spur of males with denticle on external edge only (
<figureCitation id="130887E5FFDBFFAC0C31E071FF3BFE65" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2I</figureCitation>
). Females with metatibial spur apically sinuous and bifurcate, inner and external edges with denticles (
<figureCitation id="130887E5FFDBFFAC0C31E056FF33FE46" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2L</figureCitation>
); males only with a conspicuous denticle on apical inner edge of metatibial spur (
<figureCitation id="130887E5FFDBFFAC0D23E0AAFB52FE46" box="[1149,1243,401,427]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2K</figureCitation>
). Apical third of metatibiae almost straight in females (
<figureCitation id="130887E5FFDBFFAC0B1BE08EFD12FE22" box="[581,667,437,463]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2L</figureCitation>
), distinctly curved inward in males (
<figureCitation id="130887E5FFDBFFAC0D69E08EFB1BFE22" box="[1079,1170,437,463]" captionStart="FIGURE 2" captionStartId="8.[151,250,1870,1894]" captionTargetBox="[152,1434,358,1832]" captionTargetId="figure-104@8.[151,1436,345,1847]" captionTargetPageId="8" captionText="FIGURE 2. Details of the external morphology of Deltepilissus infernalis. AB, Body, male, dorsal and ventral views, respectively; CD, body, female, dorsal and ventral views, respectively; E, head, male; F, hypomeron, male; G, abdominal ventrites, male; H, pygidium, male; IJ, detail of protibial spur, male and female, respectively; KL, right metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388913" httpUri="https://zenodo.org/record/6388913/files/figure.png" pageId="7" pageNumber="90">Fig. 2K</figureCitation>
).
<emphasis id="B9474772FFDBFFAC0DF8E08EFA90FE22" bold="true" box="[1190,1305,437,463]" pageId="7" pageNumber="90">Aedeagus</emphasis>
. Parameres symmetrical, subparallel in lateral view, abruptly pointing downward apically (
<figureCitation id="130887E5FFDBFFAC0D52E0E2FBE8FE1E" box="[1036,1121,473,499]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="7" pageNumber="90">Fig. 3F</figureCitation>
). Apex of parameres curved inward in dorsal and ventral views (
<figureCitation id="130887E5FFDBFFAC0B7BE0C7FD2AFDFA" box="[549,675,508,535]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="7" pageNumber="90">Figs 3DE</figureCitation>
).
<emphasis id="B9474772FFDBFFAC0BEBE0C6FCDAFDFA" bold="true" box="[693,851,509,535]" pageId="7" pageNumber="90">Endophallus.</emphasis>
Lamella copulatrix absent. Superior right peripheral endophallite ring shaped; lateral projection arched, rounded apically (
<figureCitation id="130887E5FFDBFFAC0AC7E31BFC78FDD7" box="[921,1009,544,570]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="7" pageNumber="90">Fig. 3H</figureCitation>
). Frontolateral peripheral endophallite elongate, irregular (
<figureCitation id="130887E5FFDBFFAC082BE37FFE4FFDB3" box="[373,454,580,606]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="7" pageNumber="90">Fig. 3J</figureCitation>
). Complex of axial and subaxial endophallites elongate, irregular (
<figureCitation id="130887E5FFDBFFAC0DE9E37FFA98FDB3" box="[1207,1297,580,606]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="7" pageNumber="90">Fig. 3N</figureCitation>
). Additional endophallites short, comma shaped (
<figureCitation id="130887E5FFDBFFAC0B6EE353FD0EFD6F" box="[560,647,616,642]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. AC, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. DF. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="7" pageNumber="90">Fig. 3L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDBFFAC0999E3B7FEF0FAC4" pageId="7" pageNumber="90" type="materials_examined">
<paragraph id="8B8C9B60FFDBFFAC0999E3B7FE15FCFC" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="90">
<emphasis id="B9474772FFDBFFAC0999E3B7FCC9FD45" bold="true" box="[199,832,651,680]" pageId="7" pageNumber="90">
Type material.
<taxonomicName id="4C33E0E3FFDBFFAC082BE3B0FD27FD4B" authority="Harold." authorityName="Harold." box="[373,686,651,678]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC082BE3B0FDC7FD4B" bold="true" box="[373,590,651,678]" italics="true" pageId="7" pageNumber="90">Canthon infernalis</emphasis>
Harold.
</taxonomicName>
<typeStatus id="548825C2FFDBFFAC0BEFE3B7FCAEFD4B" box="[689,807,652,678]" pageId="7" pageNumber="90" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D3550E9FFDBFFAC0A75E3B6FCC9FD45" box="[811,832,653,680]" pageId="7" pageNumber="90" type="male"></specimenCount>
</emphasis>
(
<emphasis id="B9474772FFDBFFAC0A13E3B7FB8CFD48" bold="true" box="[845,1029,651,678]" pageId="7" pageNumber="90">here designated</emphasis>
;
<collectionCode id="ED2203A5FFDBFFAC0D4EE3B7FBE8FD4B" box="[1040,1121,652,678]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="7" pageNumber="90">MFNB</collectionCode>
)(
<figureCitation id="130887E5FFDBFFAC0D2AE3B0FB31FD4B" box="[1140,1208,651,678]" captionStart="FIGURE 4" captionStartId="10.[151,250,1910,1934]" captionTargetBox="[278,1321,181,1886]" captionTargetId="figure-30@10.[267,1321,181,1886]" captionTargetPageId="10" captionText="FIGURE 4. Type specimens in Deltepilissus and respective label data. AB, Lectotype of Canthon diabolicus Harold, 1880: A, body, dorsal view; B, labels. CD, Lectotype of Canthon infernalis Harold, 1880: C, body, dorsal view; D, labels. EH, holotype of Deltepilissus travassosi Pereira, 1949: E, body, dorsal view; F, head; G, left protibia, dorsal view; H, protibial and tarsal claws. Photographs AD provided by Fernando Z. Vaz-de-Mello." figureDoi="http://doi.org/10.5281/zenodo.6388917" httpUri="https://zenodo.org/record/6388917/files/figure.png" pageId="7" pageNumber="90">Figs4</figureCitation>
C-D):“
<typeStatus id="548825C2FFDBFFAC0C5BE3B7FA15FD4B" box="[1285,1436,652,678]" pageId="7" pageNumber="90" type="holotype">HOLOTYPE</typeStatus>
//
<emphasis id="B9474772FFDBFFAC09F0E394FEC3FD24" box="[174,330,687,713]" italics="true" pageId="7" pageNumber="90">
<collectingCountry id="F324DBF0FFDBFFAC09F0E394FF7BFD24" box="[174,242,687,713]" name="Brazil" pageId="7" pageNumber="90">Brasil</collectingCountry>
Besuxe
</emphasis>
// 9005 // Type //
<taxonomicName id="4C33E0E3FFDBFFAC0B48E394FD08FD24" box="[534,641,687,713]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC0B48E394FD08FD24" box="[534,641,687,713]" italics="true" pageId="7" pageNumber="90">infernalis</emphasis>
</taxonomicName>
/
<emphasis id="B9474772FFDBFFAC0BC7E38BFD4FFD24" box="[665,710,688,713]" italics="true" pageId="7" pageNumber="90">Har</emphasis>
//
<typeStatus id="548825C2FFDBFFAC0BBBE394FCFFFD24" box="[741,886,687,713]" pageId="7" pageNumber="90" type="syntype">SYNTYPUS</typeStatus>
/
<taxonomicName id="4C33E0E3FFDBFFAC0AD3E394FC65FD24" authorityName="Hoffmannsegg" authorityYear="1817" box="[909,1004,687,713]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus">Canthon</taxonomicName>
/
<taxonomicName id="4C33E0E3FFDBFFAC0D5DE394FA80FD24" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" box="[1027,1289,687,713]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
infernalis
<bibRefCitation id="EFA2E691FFDBFFAC0D2AE394FA80FD24" author="Harold, E." box="[1140,1289,687,713]" pageId="7" pageNumber="90" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
</taxonomicName>
/ labelled by MNHUB 2013.
<emphasis id="B9474772FFDBFFAC0813E3E9FCC7FD02" bold="true" box="[333,846,722,751]" pageId="7" pageNumber="90">
<taxonomicName id="4C33E0E3FFDBFFAC0813E3E9FD37FD00" authority="Pereira." authorityName="Pereira." box="[333,702,722,749]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0813E3E9FDD1FD00" bold="true" box="[333,600,722,749]" italics="true" pageId="7" pageNumber="90">Deltepilissus travassosi</emphasis>
Pereira.
</taxonomicName>
<typeStatus id="548825C2FFDBFFAC0B9BE3E8FCBBFD00" box="[709,818,723,749]" pageId="7" pageNumber="90" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3550E9FFDBFFAC0A67E3EFFCC7FD02" box="[825,846,724,751]" pageId="7" pageNumber="90" type="male"></specimenCount>
</emphasis>
(
<collectionCode id="ED2203A5FFDBFFAC0A03E3E8FC20FD00" box="[861,937,723,749]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="90" type="Museum">MZSP</collectionCode>
) (
<figureCitation id="130887E5FFDBFFAC0A9EE3E8FBC9FD00" box="[960,1088,723,749]" captionStart="FIGURE 4" captionStartId="10.[151,250,1910,1934]" captionTargetBox="[278,1321,181,1886]" captionTargetId="figure-30@10.[267,1321,181,1886]" captionTargetPageId="10" captionText="FIGURE 4. Type specimens in Deltepilissus and respective label data. AB, Lectotype of Canthon diabolicus Harold, 1880: A, body, dorsal view; B, labels. CD, Lectotype of Canthon infernalis Harold, 1880: C, body, dorsal view; D, labels. EH, holotype of Deltepilissus travassosi Pereira, 1949: E, body, dorsal view; F, head; G, left protibia, dorsal view; H, protibial and tarsal claws. Photographs AD provided by Fernando Z. Vaz-de-Mello." figureDoi="http://doi.org/10.5281/zenodo.6388917" httpUri="https://zenodo.org/record/6388917/files/figure.png" pageId="7" pageNumber="90">Figs 4EH</figureCitation>
):
<collectingRegion id="49F75582FFDBFFAC0D0DE3E8FB71FD00" box="[1107,1272,723,749]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">Rio de Janeiro</collectingRegion>
, Paineiras,
<date id="FF8DBDA0FFDBFFAC0C26E3E8FF5BFCFC" pageId="7" pageNumber="90" value="1934-04">IV-1934</date>
, Dr. L. Travassos.
</paragraph>
<paragraph id="8B8C9B60FFDBFFAC0999E221FAEBFBE6" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="90">
<emphasis id="B9474772FFDBFFAC0999E221FE31FCD9" bold="true" box="[199,440,794,820]" pageId="7" pageNumber="90">Additional material.</emphasis>
<materialsCitation id="3B5B913DFFDBFFAC08E3E221FE88FCB5" ID-GBIF-Occurrence="3738408308" collectionCode="BMNH" collectorName="Brasil" country="Brazil" location="BRAZIL" pageId="7" pageNumber="90" specimenCount="1">
<emphasis id="B9474772FFDBFFAC08E3E221FDA1FCD9" bold="true" box="[445,552,794,820]" pageId="7" pageNumber="90">
<collectingCountry id="F324DBF0FFDBFFAC08E3E221FDA1FCD9" box="[445,552,794,820]" name="Brazil" pageId="7" pageNumber="90">BRAZIL</collectingCountry>
</emphasis>
: no specific locality [
<collectorName id="26C6FEB6FFDBFFAC0A4DE221FCD0FCD9" box="[787,857,794,820]" pageId="7" pageNumber="90">Brasil</collectorName>
inter], no date and collector, 1 unsexed specimen (
<collectionCode id="ED2203A5FFDBFFAC09C1E205FF70FCB5" box="[159,249,830,856]" pageId="7" pageNumber="90">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5B913DFFDBFFAC0855E27BFB1DFCB5" ID-GBIF-Occurrence="3738408305" box="[267,1172,830,859]" collectingDate="1986-09-05" collectionCode="CEMT" collectorName="R. L. C. Baptista" country="Brazil" location="Mimoso do Sul" pageId="7" pageNumber="90" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo">
<collectingRegion id="49F75582FFDBFFAC0855E27BFE62FCB5" box="[267,491,830,859]" country="Brazil" name="Espirito Santo" pageId="7" pageNumber="90">ESPÍRITO SANTO</collectingRegion>
,
<location id="8EECCDBBFFDBFFAC08ABE205FD2BFCB5" LSID="urn:lsid:plazi:treatment:039A2A76FFDAFFA309C9E2E9FDE1FEAC:8EECCDBBFFDBFFAC08ABE205FD2BFCB5" box="[501,674,830,856]" country="Brazil" name="Mimoso do Sul" pageId="7" pageNumber="90" stateProvince="Espirito Santo">Mimoso do Sul</location>
,
<date id="FF8DBDA0FFDBFFAC0BF3E205FCA6FCB4" box="[685,815,830,857]" pageId="7" pageNumber="90" value="1986-09-05">
<collectingDate id="EFC94448FFDBFFAC0BF3E205FCA6FCB4" box="[685,815,830,857]" pageId="7" pageNumber="90" value="1986-09-05">05.IX.1986</collectingDate>
</date>
,
<collectorName id="26C6FEB6FFDBFFAC0A67E205FC6EFCB5" box="[825,999,830,856]" pageId="7" pageNumber="90">R.L.C. Baptista</collectorName>
<specimenCount id="9D3550E9FFDBFFAC0D5FE205FBA0FCB4" box="[1025,1065,830,857]" pageId="7" pageNumber="90" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED2203A5FFDBFFAC0D69E205FB02FCB5" box="[1079,1163,830,856]" pageId="7" pageNumber="90">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5B913DFFDBFFAC0DC3E205FDE2FC4D" ID-GBIF-Occurrence="3738408310" collectingDate="1948-02-25" collectionCode="BMNH, MNRJ" collectorName="W. Zikan" country="Brazil" county="Fry" elevation="800" location="Serra da Caveira" municipality="Itaguai" pageId="7" pageNumber="90" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingRegion id="49F75582FFDBFFAC0DC3E205FAFFFCB5" box="[1181,1398,830,857]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">RIO DE JANEIRO</collectingRegion>
, no date and collector 1 unsexed specimen (
<collectionCode id="ED2203A5FFDBFFAC0B38E259FD35FC91" box="[614,700,866,892]" pageId="7" pageNumber="90">BMNH</collectionCode>
ex
<collectingCounty id="62EDE3ECFFDBFFAC0BA6E259FC96FC91" box="[760,799,866,892]" pageId="7" pageNumber="90">Fry</collectingCounty>
collection);
<collectingMunicipality id="6BE8011AFFDBFFAC0AF5E259FC75FC91" box="[939,1020,866,892]" pageId="7" pageNumber="90">Itaguaí</collectingMunicipality>
,
<location id="8EECCDBBFFDBFFAC0D58E259FB4BFC91" LSID="urn:lsid:plazi:treatment:039A2A76FFDAFFA309C9E2E9FDE1FEAC:8EECCDBBFFDBFFAC0D58E259FB4BFC91" box="[1030,1218,866,892]" country="Brazil" county="Fry" municipality="Itaguai" name="Serra da Caveira" pageId="7" pageNumber="90" stateProvince="Rio de Janeiro">Serra da Caveira</location>
,
<quantity id="4CCB3685FFDBFFAC0D93E259FA9CFC91" box="[1229,1301,866,892]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="90" unit="m" value="800.0">
<elevation id="001E7C53FFDBFFAC0D93E259FA9CFC91" box="[1229,1301,866,892]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="90" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FF8DBDA0FFDBFFAC0C7EE259FA11FC90" box="[1312,1432,866,893]" pageId="7" pageNumber="90" value="1948-02-25">
<collectingDate id="EFC94448FFDBFFAC0C7EE259FA11FC90" box="[1312,1432,866,893]" pageId="7" pageNumber="90" value="1948-02-25">25.II.1948</collectingDate>
</date>
,
<collectorName id="26C6FEB6FFDBFFAC09C9E2BDFF76FC72" box="[151,255,901,928]" pageId="7" pageNumber="90">W. Zikán</collectorName>
1 unsexed specimen (
<collectionCode id="ED2203A5FFDBFFAC0B4CE2BDFDEAFC4D" box="[530,611,902,928]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="90">MNRJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5B913DFFDBFFAC0B29E2BDFA11FC4D" ID-GBIF-Occurrence="3738408306" box="[631,1432,901,928]" collectingDate="1948-02-25" collectionCode="CEMT" collectorName="Zikan" country="Brazil" elevation="600" location="Serra da Caveira" municipality="Itaguai" pageId="7" pageNumber="90" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="6BE8011AFFDBFFAC0B29E2BDFD41FC4D" box="[631,712,902,928]" pageId="7" pageNumber="90">Itaguaí</collectingMunicipality>
,
<location id="8EECCDBBFFDBFFAC0B8AE2BDFC1DFC72" LSID="urn:lsid:plazi:treatment:039A2A76FFDAFFA309C9E2E9FDE1FEAC:8EECCDBBFFDBFFAC0B8AE2BDFC1DFC72" box="[724,916,901,928]" country="Brazil" municipality="Itaguai" name="Serra da Caveira" pageId="7" pageNumber="90">Serra da Caveira</location>
,
<quantity id="4CCB3685FFDBFFAC0AFEE2BEFC63FC4D" box="[928,1002,901,928]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="90" unit="m" value="600.0">
<elevation id="001E7C53FFDBFFAC0AFEE2BEFC63FC4D" box="[928,1002,901,928]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="90" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FF8DBDA0FFDBFFAC0AA9E2BEFBE7FC4D" box="[1015,1134,901,928]" pageId="7" pageNumber="90" value="1948-02-25">
<collectingDate id="EFC94448FFDBFFAC0AA9E2BEFBE7FC4D" box="[1015,1134,901,928]" pageId="7" pageNumber="90" value="1948-02-25">25.II.1948</collectingDate>
</date>
, W.
<collectorName id="26C6FEB6FFDBFFAC0DFFE2BEFB6BFC72" box="[1185,1250,901,927]" pageId="7" pageNumber="90">Zikán</collectorName>
<specimenCount id="9D3550E9FFDBFFAC0C5FE2BDFAA5FC4D" box="[1281,1324,902,928]" pageId="7" pageNumber="90" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED2203A5FFDBFFAC0C62E2BDFA19FC4D" box="[1340,1424,902,928]" pageId="7" pageNumber="90">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5B913DFFDBFFAC09C9E291FCAFFC2E" ID-GBIF-Occurrence="3738408311" box="[151,806,937,964]" collectingDate="1999-12" collectionCode="CEMT" collectorName="E. &amp; P. Grossi" country="Brazil" location="Nova Friburgo" pageId="7" pageNumber="90" specimenCount="1" specimenCount-female="1">
<location id="8EECCDBBFFDBFFAC09C9E291FEC8FC2E" LSID="urn:lsid:plazi:treatment:039A2A76FFDAFFA309C9E2E9FDE1FEAC:8EECCDBBFFDBFFAC09C9E291FEC8FC2E" box="[151,321,937,964]" country="Brazil" name="Nova Friburgo" pageId="7" pageNumber="90">Nova Friburgo</location>
,
<date id="FF8DBDA0FFDBFFAC0811E292FE3EFC2E" box="[335,439,937,963]" pageId="7" pageNumber="90" value="1999-12">
<collectingDate id="EFC94448FFDBFFAC0811E292FE3EFC2E" box="[335,439,937,963]" pageId="7" pageNumber="90" value="1999-12">XII.1999</collectingDate>
</date>
,
<collectorName id="26C6FEB6FFDBFFAC089BE291FDE5FC2E" box="[453,620,937,964]" pageId="7" pageNumber="90">E. &amp; P. Grossi</collectorName>
<specimenCount id="9D3550E9FFDBFFAC0BD2E292FD30FC29" box="[652,697,937,964]" pageId="7" pageNumber="90" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED2203A5FFDBFFAC0B94E292FC97FC2E" box="[714,798,937,963]" pageId="7" pageNumber="90">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5B913DFFDBFFAC0A6AE291FEFDFC0A" ID-GBIF-Occurrence="3738408301" collectingDate="1999-12" collectionCode="CERPE" collectorName="E. &amp; P. Grossi" country="Brazil" location="Nova Friburgo" pageId="7" pageNumber="90" specimenCount="1">
<location id="8EECCDBBFFDBFFAC0A6AE291FC57FC2E" LSID="urn:lsid:plazi:treatment:039A2A76FFDAFFA309C9E2E9FDE1FEAC:8EECCDBBFFDBFFAC0A6AE291FC57FC2E" box="[820,990,937,964]" country="Brazil" name="Nova Friburgo" pageId="7" pageNumber="90">Nova Friburgo</location>
,
<date id="FF8DBDA0FFDBFFAC0AB2E292FBDDFC2E" box="[1004,1108,937,963]" pageId="7" pageNumber="90" value="1999-12">
<collectingDate id="EFC94448FFDBFFAC0AB2E292FBDDFC2E" box="[1004,1108,937,963]" pageId="7" pageNumber="90" value="1999-12">XII.1999</collectingDate>
</date>
,
<collectorName id="26C6FEB6FFDBFFAC0D3CE291FA81FC2E" box="[1122,1288,937,964]" pageId="7" pageNumber="90">E. &amp; P. Grossi</collectorName>
1 unsexed specimen (
<collectionCode id="ED2203A5FFDBFFAC0850E2F6FEE5FC0A" box="[270,364,973,999]" pageId="7" pageNumber="90">CERPE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5B913DFFDBFFAC0823E2F6FE45FBE1" ID-GBIF-Occurrence="3738408309" collectingDate="1971-12" collectionCode="CMNC, A, AMBC" country="Brazil" location="Martinez" municipality="Howden" pageId="7" pageNumber="90" specimenCount="2" specimenCount-male="2" stateProvince="Sao Paulo">
<collectorName id="26C6FEB6FFDBFFAC0823E2F6FD96FC0A" box="[381,543,973,999]" hasSafeAffix="true" pageId="7" pageNumber="90">Rio de Janeiro</collectorName>
,
<collectorName id="26C6FEB6FFDBFFAC0B76E2F6FD3DFC0A" box="[552,692,973,999]" pageId="7" pageNumber="90">Jacarepaguá</collectorName>
, 1956, no collector
<specimenCount id="9D3550E9FFDBFFAC0AC0E2F6FC4EFC05" box="[926,967,973,1000]" pageId="7" pageNumber="90" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2203A5FFDBFFAC0A8AE2F6FBA1FC0A" box="[980,1064,973,999]" pageId="7" pageNumber="90">CMNC</collectionCode>
ex. H. &amp;
<collectionCode id="ED2203A5FFDBFFAC0DFCE2F6FB30FC0A" box="[1186,1209,973,999]" pageId="7" pageNumber="90">A</collectionCode>
.
<collectingMunicipality id="6BE8011AFFDBFFAC0D9DE2F6FAABFC0A" box="[1219,1314,973,999]" pageId="7" pageNumber="90">Howden</collectingMunicipality>
collection, ex.
<collectionCode id="ED2203A5FFDBFFAC09E0E2CAFF5CFBE6" box="[190,213,1009,1035]" pageId="7" pageNumber="90">A</collectionCode>
.
<location id="8EECCDBBFFDBFFAC09BEE2CAFECCFBE6" LSID="urn:lsid:plazi:treatment:039A2A76FFDAFFA309C9E2E9FDE1FEAC:8EECCDBBFFDBFFAC09BEE2CAFECCFBE6" box="[224,325,1009,1035]" country="Brazil" municipality="Howden" name="Martinez" pageId="7" pageNumber="90" stateProvince="Sao Paulo">Martínez</location>
collection).
</materialsCitation>
<materialsCitation id="3B5B913DFFDBFFAC088CE2C9FAD7FBE6" ID-GBIF-Occurrence="3738408302" box="[466,1374,1009,1037]" collectingDate="1971-12" collectionCode="CMNC, A, AMBC" collectorName="Ayr Bello" country="Brazil" location="Martinez" municipality="Howden" pageId="7" pageNumber="90" specimenCount="2" specimenCount-male="2" stateProvince="Sao Paulo">
<collectingRegion id="49F75582FFDBFFAC088CE2C9FDE6FBE6" box="[466,623,1009,1037]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="90">SÃO PAULO</collectingRegion>
,
<collectorName id="26C6FEB6FFDBFFAC0B24E2CAFD5EFBE6" box="[634,727,1009,1035]" pageId="7" pageNumber="90">Guarujá</collectorName>
,
<collectorName id="26C6FEB6FFDBFFAC0BBCE2CAFC37FBE6" box="[738,958,1009,1035]" pageId="7" pageNumber="90">Forte dos Andradas</collectorName>
,
<date id="FF8DBDA0FFDBFFAC0A94E2CAFBBBFBE6" box="[970,1074,1009,1035]" pageId="7" pageNumber="90" value="1971-12">
<collectingDate id="EFC94448FFDBFFAC0A94E2CAFBBBFBE6" box="[970,1074,1009,1035]" pageId="7" pageNumber="90" value="1971-12">XII.1971</collectingDate>
</date>
,
<collectorName id="26C6FEB6FFDBFFAC0D62E2CAFB20FBE6" box="[1084,1193,1009,1035]" pageId="7" pageNumber="90">Ayr Bello</collectorName>
<specimenCount id="9D3550E9FFDBFFAC0D9BE2CAFB66FBE6" box="[1221,1263,1009,1035]" pageId="7" pageNumber="90" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2203A5FFDBFFAC0DA0E2CAFADEFBE6" box="[1278,1367,1009,1035]" pageId="7" pageNumber="90">AMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8C9B60FFDBFFAC0999E52FFEF0FAC4" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="90">
<emphasis id="B9474772FFDBFFAC0999E52FFEAAFBC3" bold="true" box="[199,291,1044,1070]" pageId="7" pageNumber="90">Habitat</emphasis>
. This species inhabits the Brazilian Atlantic rainforest from southern
<collectingRegion id="49F75582FFDBFFAC0D7CE52FFB48FBC2" box="[1058,1217,1044,1071]" country="Brazil" name="Espirito Santo" pageId="7" pageNumber="90">Espírito Santo</collectingRegion>
to
<collectingRegion id="49F75582FFDBFFAC0DBCE52EFAD8FBC3" box="[1250,1361,1044,1071]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="90">São Paulo</collectingRegion>
states, mainly at low elevation along the coastal parts of the Atlantic rainforest, from
<quantity id="4CCB3685FFDBFFAC0D7CE503FB03FBBE" box="[1058,1162,1080,1107]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="7" pageNumber="90" unit="m" value="400.0" valueMax="800.0" valueMin="0.0">
0
<specimenCount id="9D3550E9FFDBFFAC0D60E503FB03FBBE" box="[1086,1162,1080,1107]" pageId="7" pageNumber="90" type="generic">800 m</specimenCount>
</quantity>
.
<collectionCode id="ED2203A5FFDBFFAC0DC8E503FB20FBBF" box="[1174,1193,1080,1106]" pageId="7" pageNumber="90">A</collectionCode>
single specimen was collected by Renner Baptista in a soil block (manure and roots) dug up in a cattle ranch in
<collectingRegion id="49F75582FFDBFFAC0DFFE567FACCFB9B" box="[1185,1349,1116,1142]" country="Brazil" name="Espirito Santo" pageId="7" pageNumber="90">Espírito Santo</collectingRegion>
,
<collectingCountry id="F324DBF0FFDBFFAC0C0FE567FA10FB9B" box="[1361,1433,1116,1142]" name="Brazil" pageId="7" pageNumber="90">Brazil</collectingCountry>
.
<collectionCode id="ED2203A5FFDBFFAC09C9E5BBFF23FB77" box="[151,170,1152,1178]" pageId="7" pageNumber="90">A</collectionCode>
specimen was also collected within debris (leaves, seaweed, broken shells, insects and some plastic garbage) deposited on beach sand in Guarujá,
<collectingRegion id="49F75582FFDBFFAC0B71E59FFD2AFB50" box="[559,675,1187,1214]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="90">São Paulo</collectingRegion>
,
<collectingCountry id="F324DBF0FFDBFFAC0BF3E598FD78FB50" box="[685,753,1187,1213]" name="Brazil" pageId="7" pageNumber="90">Brazil</collectingCountry>
(Ayr Bello, personal communication). All those records were made from manual collect method. However, the data presented above are not enough to define the food habits of
<taxonomicName id="4C33E0E3FFDBFFAC09C9E5D7FEADFAE8" box="[151,292,1259,1285]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC09C9E5D7FEADFAE8" box="[151,292,1259,1285]" italics="true" pageId="7" pageNumber="90">D. infernalis</emphasis>
</taxonomicName>
. As already mentioned for
<taxonomicName id="4C33E0E3FFDBFFAC0B0FE5D7FD6EFAE8" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[593,743,1259,1285]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFDBFFAC0B0FE5D7FD6EFAE8" box="[593,743,1259,1285]" italics="true" pageId="7" pageNumber="90">D. diabolicus</emphasis>
</taxonomicName>
, the baits traditionally used for collecting dung beetles did not attract
<taxonomicName id="4C33E0E3FFDBFFAC09BAE434FEF8FAC4" box="[228,369,1295,1321]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC09BAE434FEF8FAC4" box="[228,369,1295,1321]" italics="true" pageId="7" pageNumber="90">D. infernalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDBFFAC0999E409FD92FA55" pageId="7" pageNumber="90" type="distribution">
<paragraph id="8B8C9B60FFDBFFAC0999E409FD92FA55" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="90">
<emphasis id="B9474772FFDBFFAC0999E409FEE8FAA1" bold="true" box="[199,353,1330,1356]" pageId="7" pageNumber="90">Distribution.</emphasis>
Known from southeastern Brazil (
<collectingRegion id="49F75582FFDBFFAC0BABE409FC14FAA1" box="[757,925,1330,1356]" country="Brazil" name="Espirito Santo" pageId="7" pageNumber="90">Espírito Santo</collectingRegion>
,
<collectingRegion id="49F75582FFDBFFAC0AF5E409FBDDFAA1" box="[939,1108,1330,1356]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">Rio de Janeiro</collectingRegion>
and
<collectingRegion id="49F75582FFDBFFAC0DCFE409FA82FAA0" box="[1169,1291,1330,1357]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="90">São Paulo</collectingRegion>
).
<emphasis id="B9474772FFDBFFAC0C7EE409FEE5FA9D" bold="true" pageId="7" pageNumber="90">
Literature records.
<collectingCountry id="F324DBF0FFDBFFAC085FE46DFEE5FA9D" box="[257,364,1366,1392]" name="Brazil" pageId="7" pageNumber="90">BRAZIL</collectingCountry>
</emphasis>
:
<collectingRegion id="49F75582FFDBFFAC0820E46DFDD7FA9D" box="[382,606,1366,1393]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">RIO DE JANEIRO</collectingRegion>
, Itaguaí (
<bibRefCitation id="EFA2E691FFDBFFAC0B94E46DFCE9FA9C" author="Pereira, F." box="[714,864,1366,1393]" pageId="7" pageNumber="90" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">Pereira 1953</bibRefCitation>
),
<collectingRegion id="49F75582FFDBFFAC0A29E46DFB94FA9D" box="[887,1053,1366,1392]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">Rio de Janeiro</collectingRegion>
[Paineiras] (
<bibRefCitation id="EFA2E691FFDBFFAC0DEFE46DFACFFA9D" author="Pereira, F." box="[1201,1350,1366,1392]" pageId="7" pageNumber="90" pagination="231 - 246" refId="ref9277" refString="Pereira, F. (1949) Escarabeideos americanos II (Coleopt. - Scarabaeidae). Arquivos do Museu Paranaense, 7, 231 - 246." type="journal article" year="1949">Pereira 1949</bibRefCitation>
,
<bibRefCitation id="EFA2E691FFDBFFAC0C0AE46DFA06FA9C" author="Pereira, F." box="[1364,1423,1366,1393]" pageId="7" pageNumber="90" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">1953</bibRefCitation>
),
<collectingRegion id="49F75582FFDBFFAC09C9E441FEB2FA79" box="[151,315,1402,1428]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">Rio de Janeiro</collectingRegion>
[cited as “Guanabara”, the state-level name by which the city was known between 1960 and 1975] (
<bibRefCitation id="EFA2E691FFDBFFAC09C1E4A6FE38FA55" author="Vulcano, M. A. &amp; Pereira, F." box="[159,433,1437,1464]" pageId="7" pageNumber="90" pagination="570 - 685" refId="ref10315" refString="Vulcano, M. A. &amp; Pereira, F. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey, 15, 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964</bibRefCitation>
) (
<figureCitation id="130887E5FFDBFFAC0896E4A6FD84FA55" box="[456,525,1437,1464]" captionStart="FIGURE 5" captionStartId="11.[151,250,1103,1127]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-19@11.[151,1436,181,1080]" captionTargetPageId="11" captionText="FIGURE 5. Known distribution of species of Deltepilissus. A, Records plotted on terrestrial biomes layer; B, records plotted on elevational range layer." figureDoi="http://doi.org/10.5281/zenodo.6388921" httpUri="https://zenodo.org/record/6388921/files/figure.png" pageId="7" pageNumber="90">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFDBFFA30999E4FAFDE1FEAC" lastPageId="8" lastPageNumber="91" pageId="7" pageNumber="90" type="discussion">
<paragraph id="8B8C9B60FFDBFFAC0999E4FAFD65F888" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="90">
<emphasis id="B9474772FFDBFFAC0999E4FAFEB2FA36" bold="true" box="[199,315,1473,1499]" pageId="7" pageNumber="90">Remarks.</emphasis>
<bibRefCitation id="EFA2E691FFDBFFAC081CE4FAFE68FA36" author="Harold, E." box="[322,481,1473,1499]" pageId="7" pageNumber="90" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold (1880)</bibRefCitation>
described
<taxonomicName id="4C33E0E3FFDBFFAC0B05E4FAFCA4FA36" box="[603,813,1473,1499]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC0B05E4FAFCA4FA36" box="[603,813,1473,1499]" italics="true" pageId="7" pageNumber="90">Canthon infernalis</emphasis>
</taxonomicName>
from one or more specimens collected by von Olfers, a German naturalist, in
<collectingCountry id="F324DBF0FFDBFFAC08D1E4DEFE5EFA12" box="[399,471,1509,1535]" name="Brazil" pageId="7" pageNumber="90">Brazil</collectingCountry>
. A type specimen was found and photographed by Fernando Vaz-de-Mello in MFNB (see material and methods section).
<bibRefCitation id="EFA2E691FFDBFFAC0B6CE732FD5CF9CE" author="Pereira, F." box="[562,725,1545,1571]" pageId="7" pageNumber="90" pagination="231 - 246" refId="ref9277" refString="Pereira, F. (1949) Escarabeideos americanos II (Coleopt. - Scarabaeidae). Arquivos do Museu Paranaense, 7, 231 - 246." type="journal article" year="1949">Pereira (1949)</bibRefCitation>
described
<taxonomicName id="4C33E0E3FFDBFFAC0A0CE732FBD4F9CE" box="[850,1117,1545,1571]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0A0CE732FBD4F9CE" box="[850,1117,1545,1571]" italics="true" pageId="7" pageNumber="90">Deltepilissus travassosi</emphasis>
</taxonomicName>
on a specimen from
<collectingRegion id="49F75582FFDBFFAC0C0FE732FF62F9AB" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="F324DBF0FFDBFFAC09A7E717FEB4F9AB" box="[249,317,1580,1606]" name="Brazil" pageId="7" pageNumber="90">Brazil</collectingCountry>
[
<typeStatus id="548825C2FFDBFFAC0810E717FE3AF9AB" box="[334,435,1580,1606]" pageId="7" pageNumber="90" type="holotype">holotype</typeStatus>
from Parque Nacional da Tijuca, Paineiras,
<collectingRegion id="49F75582FFDBFFAC0AE7E717FBECF9AB" box="[953,1125,1580,1606]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="90">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="F324DBF0FFDBFFAC0D2DE717FB32F9AB" box="[1139,1211,1580,1606]" name="Brazil" pageId="7" pageNumber="90">Brazil</collectingCountry>
; housed in MZSP]. Later,
<bibRefCitation id="EFA2E691FFDBFFAC09BFE76BFE0AF986" author="Pereira, F." box="[225,387,1616,1643]" pageId="7" pageNumber="90" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">Pereira (1953)</bibRefCitation>
considered
<taxonomicName id="4C33E0E3FFDBFFAC0B50E76AFD2AF987" box="[526,675,1617,1642]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0B50E76AFD2AF987" box="[526,675,1617,1642]" italics="true" pageId="7" pageNumber="90">D. travassosi</emphasis>
</taxonomicName>
a junior subjective synonym of
<taxonomicName id="4C33E0E3FFDBFFAC0D4BE76BFB60F987" box="[1045,1257,1616,1642]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC0D4BE76BFB60F987" box="[1045,1257,1616,1642]" italics="true" pageId="7" pageNumber="90">Canthon infernalis</emphasis>
</taxonomicName>
and transferred the latter to the genus
<taxonomicName id="4C33E0E3FFDBFFAC08CEE74FFDA8F963" authorityName="Pereira" authorityYear="1949" box="[400,545,1652,1678]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDBFFAC08CEE74FFDA8F963" box="[400,545,1652,1678]" italics="true" pageId="7" pageNumber="90">Deltepilissus</emphasis>
</taxonomicName>
. In this study, we examined the
<typeStatus id="548825C2FFDBFFAC0AD2E74FFC67F963" box="[908,1006,1652,1678]" pageId="7" pageNumber="90" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C33E0E3FFDBFFAC0D4DE74EFB21F963" box="[1043,1192,1653,1678]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0D4DE74EFB21F963" box="[1043,1192,1653,1678]" italics="true" pageId="7" pageNumber="90">D. travassosi</emphasis>
</taxonomicName>
and compared it with the photographs of the type of
<taxonomicName id="4C33E0E3FFDBFFAC0B5AE7A3FD50F95F" box="[516,729,1688,1714]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC0B5AE7A3FD50F95F" box="[516,729,1688,1714]" italics="true" pageId="7" pageNumber="90">Canthon infernalis</emphasis>
</taxonomicName>
, and we did not find significant differences in their external morphology. Unfortunately, we could not examine the genitalia of the
<typeStatus id="548825C2FFDBFFAC0A94E780FBA5F938" box="[970,1068,1723,1749]" pageId="7" pageNumber="90" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C33E0E3FFDBFFAC0D09E787FB66F938" box="[1111,1263,1724,1749]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0D09E787FB66F938" box="[1111,1263,1724,1749]" italics="true" pageId="7" pageNumber="90">D. travassosi</emphasis>
</taxonomicName>
, as the current policies of the MZSP do not allow the dissection of name-bearing type specimens (
<emphasis id="B9474772FFDBFFAC0D09E7DBFBF3F914" box="[1111,1146,1760,1785]" italics="true" pageId="7" pageNumber="90">i.e.</emphasis>
,
<typeStatus id="548825C2FFDBFFAC0DD4E7E4FB73F914" box="[1162,1274,1759,1785]" pageId="7" pageNumber="90" type="holotype">holotypes</typeStatus>
,
<typeStatus id="548825C2FFDBFFAC0C59E7E4FAF3F917" box="[1287,1402,1759,1786]" pageId="7" pageNumber="90" type="lectotype">lectotypes</typeStatus>
) if there are no other available types in that collection (
<emphasis id="B9474772FFDBFFAC0B89E63FFD73F8F0" box="[727,762,1796,1821]" italics="true" pageId="7" pageNumber="90">i.e.</emphasis>
,
<typeStatus id="548825C2FFDBFFAC0A59E638FCFCF8F0" box="[775,885,1795,1821]" pageId="7" pageNumber="90" type="paratype">paratypes</typeStatus>
,
<typeStatus id="548825C2FFDBFFAC0A21E638FBAFF8F3" box="[895,1062,1795,1822]" pageId="7" pageNumber="90" type="paralectotype">paralectotypes</typeStatus>
). However, because
<taxonomicName id="4C33E0E3FFDBFFAC0C57E63FFF6FF8AC" authority="Pereira" authorityName="Pereira" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0C57E63FFA14F8F0" box="[1289,1437,1796,1821]" italics="true" pageId="7" pageNumber="90">D. travassosi</emphasis>
Pereira
</taxonomicName>
was synonymized with
<taxonomicName id="4C33E0E3FFDBFFAC08ACE61CFC91F8AC" authority="Harold" authorityName="Harold" box="[498,792,1831,1857]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC08ACE61CFD4AF8AC" box="[498,707,1831,1857]" italics="true" pageId="7" pageNumber="90">Canthon infernalis</emphasis>
Harold
</taxonomicName>
by its author (
<bibRefCitation id="EFA2E691FFDBFFAC0AEBE61CFBCEF8AC" author="Pereira, F." box="[949,1095,1830,1857]" pageId="7" pageNumber="90" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">Pereira 1953</bibRefCitation>
), we agree with
<bibRefCitation id="EFA2E691FFDBFFAC0DA2E61CFA15F8AC" author="Pereira, F." box="[1276,1436,1830,1857]" pageId="7" pageNumber="90" pagination="387 - 402" refId="ref9305" refString="Pereira, F. (1953) Notas sinonimicas (Col. Scarabaeidae). Dusenia, 4, 387 - 402." type="journal article" year="1953">Pereira (1953)</bibRefCitation>
to consider
<taxonomicName id="4C33E0E3FFDBFFAC0844E670FE27F889" box="[282,430,1867,1892]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="travassosi">
<emphasis id="B9474772FFDBFFAC0844E670FE27F889" box="[282,430,1867,1892]" italics="true" pageId="7" pageNumber="90">D. travassosi</emphasis>
</taxonomicName>
as synonym of
<taxonomicName id="4C33E0E3FFDBFFAC0B01E670FD6CF889" box="[607,741,1866,1892]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC0B01E670FD6CF889" box="[607,741,1866,1892]" italics="true" pageId="7" pageNumber="90">D infernalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8C9B60FFDBFFAC0999E655FE82F819" blockId="7.[151,1437,151,2036]" pageId="7" pageNumber="90">
<emphasis id="B9474772FFDBFFAC0999E655FE49F865" bold="true" box="[199,448,1902,1928]" pageId="7" pageNumber="90">Conservation status.</emphasis>
Over the past 25 years, the several collecting events in Atlantic rainforest remnants yielded only
<specimenCount id="9D3550E9FFDBFFAC0872E6A9FE5DF841" box="[300,468,1938,1964]" pageId="7" pageNumber="90" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C33E0E3FFDBFFAC08A2E6A8FD03F841" box="[508,650,1938,1964]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFDBFFAC08A2E6A8FD03F841" box="[508,650,1938,1964]" italics="true" pageId="7" pageNumber="90">D. infernalis</emphasis>
</taxonomicName>
(see “additional material”). Other known records for this species are older, with specimens collected between the early 19
<superScript id="7C463628FFDBFFAC0BB7E68EFD7FF82E" attach="left" box="[745,758,1973,1987]" fontSize="6" pageId="7" pageNumber="90">th</superScript>
century (18181819 by von Olfers, see
<bibRefCitation id="EFA2E691FFDBFFAC0DE2E68DFAEDF83D" author="Papavero, N." box="[1212,1380,1974,2000]" pageId="7" pageNumber="90" refId="ref9218" refString="Papavero, N. (1971) Essays on the history of Neotropical dipterology: with special reference to collectors: 1750 - 1905: Volume I. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, Brazil, 216 pp. https: // doi. org / 10.5962 / bhl. title. 101715" type="book" year="1971">Papavero 1971</bibRefCitation>
) and the 1980s.
</paragraph>
<paragraph id="8B8C9B60FFD4FFA30999E1ACFDE1FEAC" blockId="8.[151,1436,150,321]" pageId="8" pageNumber="91">
The extent of occurrence of this species was estimated as up to
<quantity id="4CCB3685FFD4FFA30AE3E1ACFB9CFF5C" box="[957,1045,151,177]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="91" unit="km" value="100.0">100 km</quantity>
<superScript id="7C463628FFD4FFA30D4BE1ADFB94FF49" attach="left" box="[1045,1053,150,164]" fontSize="6" pageId="8" pageNumber="91">2</superScript>
, with large surrounding areas of unsuitable habitat in southeastern
<collectingCountry id="F324DBF0FFD4FFA30B47E180FDE8FF38" box="[537,609,187,213]" name="Brazil" pageId="8" pageNumber="91">Brazil</collectingCountry>
. According to the International Union for Conservation of Nature (2012) criteria,
<taxonomicName id="4C33E0E3FFD4FFA309A8E1E4FE0CFF14" box="[246,389,223,249]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFD4FFA309A8E1E4FE0CFF14" box="[246,389,223,249]" italics="true" pageId="8" pageNumber="91">D. infernalis</emphasis>
</taxonomicName>
may be assessed as Critically Endangered (CR) under criterion B1ab(iii)+B2ab(iii), mainly due to high fragmentation and destruction of the Atlantic rainforest and low number of specimens collected over more than 140 years since its description.
</paragraph>
</subSubSection>
</treatment>
</document>