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<document ID-CLB-Dataset="33275" ID-DOI="http://dx.doi.org/10.3897/zookeys.666.9191" ID-PMC="PMC5534527" ID-Pensoft-Pub="1313-2970-666-1" ID-Pensoft-UUID="C43EEB41A94B930FFE439D1FAD29FF9C" ID-PubMed="28769631" ID-Zenodo-Dep="579453" ID-ZooBank="B3C377E8BBB14F328AECA2C22D1E625A" ModsDocID="1313-2970-666-1" checkinTime="1491551283800" checkinUser="pensoft" docAuthor="Moore, Matthew R., Jameson, Mary L., Garner, Beulah H., Audibert, Cedric, Smith, Andrew B. T. &amp; Seidel, Matthias" docDate="2017" docId="BB4E8D2516F7508B2C0FD7CF814D037D" docLanguage="en" docName="C43EEB41A94B930FFE439D1FAD29FF9C" docOrigin="ZooKeys 666" docPubDate="2017-04-06" docSource="http://dx.doi.org/10.3897/zookeys.666.9191" docTitle="Pelidnota thiliezi Soula 2009" docType="treatment" docVersion="9" id="C43EEB41A94B930FFE439D1FAD29FF9C" lastPageNumber="1" masterDocId="C43EEB41A94B930FFE439D1FAD29FF9C" masterDocTitle="Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies" masterLastPageNumber="349" masterPageNumber="1" pageNumber="1" updateTime="1719493172494" updateUser="carolina">
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<mods:titleInfo id="A3A87C19FF460CDBE119A58E1851A4E8">
<mods:title id="61A7932F5214C9DA94EC04D96DE578ED">Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies</mods:title>
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<mods:name id="E8377C2AE04F78710A8605BEFDA9CC2B" type="personal">
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<mods:namePart id="46FC1D76AC7B90434BF30E647A07989B">Moore, Matthew R.</mods:namePart>
<mods:affiliation id="9BFF465B5F9BA34AB5DA14D3DADDA2D4">Department of Entomology and Nematology, University of Florida Building 1881 Natural Drive Area, Steinmetz Hall, Box 110620, Gainesville, FL 32611 - 0620, USA</mods:affiliation>
</mods:name>
<mods:name id="18D523E26220D10FF85D27D3110AE721" type="personal">
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<mods:namePart id="AC1DD72F8FF8674A4B4024B705B76B37">Jameson, Mary L.</mods:namePart>
<mods:affiliation id="BE33405E4CDB1E3FA26B23E08A981F1E">Department of Biological Sciences, Wichita State University 1845 Fairmount, Box 26, Wichita, KS 67260 - 0026, USA</mods:affiliation>
<mods:nameIdentifier id="E670823B35BE9C2B692FA0725138F8B7" type="email">maryliz.jameson@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F3D5D76F909707E6481BF0F7A9848CED" type="personal">
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<mods:roleTerm id="ECDFF753FFD5FCFA93710837638BABFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C6832CF10D53FA6891AF1AA44E39EAD">Garner, Beulah H.</mods:namePart>
<mods:affiliation id="5633E4079F8B86D80BF8CC9AF854A858">Natural History Museum, Insects Division, Department of Life Sciences, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="BE39B5D33FFCFDB3C9C331182F21C1CA" type="personal">
<mods:role id="5B5E46D0C37A4ACCEF57E7C164987207">
<mods:roleTerm id="4D03FD133D5E76D0D7873E5652ED333D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8D4DB47F29E5599056C9D30A91B4660">Audibert, Cedric</mods:namePart>
<mods:affiliation id="9B7830F76B21A9A10E19CA0066EA910D">Musee des Confluences, Centre de Conservation et d'Etude des Collections, 13 A Rue Bancel, F- 69007 Lyon, France</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="A0728A7917D4F6F4F21796E56C90C8A6">Smith, Andrew B. T.</mods:namePart>
<mods:affiliation id="6FCB4EC9A7F6B42FD20B559EAFD108E9">Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada</mods:affiliation>
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<mods:namePart id="9739EC71E2AB452C853B3D146B47537A">Seidel, Matthias</mods:namePart>
</mods:name>
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<mods:date id="59DE3B69CA253B7869CD04DF5B47C779">2017</mods:date>
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<mods:number id="61E634F6B2D3B31354E8B790106CD593">2017-04-06</mods:number>
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<mods:number id="D0B6A66F13B921AC8F8F503E8326184C">666</mods:number>
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<mods:classification id="1F768B858CC31383CE94AA430633A306">journal article</mods:classification>
<mods:identifier id="0CEEC86DBB99F2E84824D63F553903C7" type="DOI">http://dx.doi.org/10.3897/zookeys.666.9191</mods:identifier>
<mods:identifier id="AE4B4FF58EF86096201E37FADD25E392" type="Pensoft-Pub">1313-2970-666-1</mods:identifier>
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<mods:identifier id="2EADBA0CEDE907EFCF6BD253A4AD81B2" type="Zenodo-Dep">579453</mods:identifier>
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<treatment id="BB4E8D2516F7508B2C0FD7CF814D037D" ID-GBIF-Taxon="128344314" LSID="urn:lsid:plazi:treatment:BB4E8D2516F7508B2C0FD7CF814D037D" httpUri="http://treatment.plazi.org/id/BB4E8D2516F7508B2C0FD7CF814D037D" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="A4885E174859860B94336E3F3AADB69D" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="F0D121B3993FA5D34ED8F4272B196AD8" pageId="0" pageNumber="1">
<taxonomicName id="18EA58D7E14E70337116DB0486777B74" ID-CoL="75ZYD" LSID="BB4E8D25-16F7-508B-2C0F-D7CF814D037D" authority="Soula, 2009" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">Pelidnota thiliezi Soula, 2009</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8E032E26FB60AF9B86378BB1E23B57A6" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="D5E19F5C8FEA8F351C5BC6AB4BAD7C89" pageId="0" pageNumber="1">
<taxonomicName id="5859073ECE893EFA47F523C7B9238621" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">Pelidnota thiliezi</taxonomicName>
Soula, 2009: 34, 112-113 [original combination].
</paragraph>
</subSubSection>
<subSubSection id="F21CE9D18F973438A36668BFB0BE7D2A" pageId="0" pageNumber="1" type="distribution">
<paragraph id="F8A1E21DCCB7866CEEB53E927CB188DD" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="33EAC3A5E13196FB08F40269916D7F9B" pageId="0" pageNumber="1">
BRAZIL:
<normalizedToken id="013DDB0B8BA7D1306A30640313B74707" originalValue="Goiás">Goias</normalizedToken>
(
<bibRefCitation id="058C46D4EF878691CCDC32DE6A486263" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="97DD3AB430D0A6F3C56F0AE7F273C0A8" pageId="0" pageNumber="1" type="types">
<paragraph id="D61CE666EA72BB345BFB5AD146E07025" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="3A75562FF23124FCDE1474188979ABD3" pageId="0" pageNumber="1">
The following specimens are deposited at CCECL. 1 ♂ neotype, 25 ♂ paratypes, 9 ♀ paratypes, 2 probable ♂ paratypes: 21 paratypes with identical label data: &quot;Goiana, Goyas
<normalizedToken id="9E688E722451E88BE020243703232BBB" originalValue="Brésil">Bresil</normalizedToken>
, IX-X/95//Paratype
<taxonomicName id="0C01961E441770261EDDAC4AEDB23648" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="E61FF9715D1C6E47A0D15807685464ED" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009&quot; (47030732 to 47030748, exch44 to exch47); &quot;Goiana, Goyas
<normalizedToken id="A3B71B8CE1A4A5EB3EB5D076B6BC7F23" originalValue="Brésil">Bresil</normalizedToken>
, IX-X/95//Paratype
<taxonomicName id="98A183209F85FA95E1DEC2891F54B0AD" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota grossiorum" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="DE631D118C70F6F4C661C80776A11B2A" italics="true" pageId="0" pageNumber="1">Pelidnota grossiorum</emphasis>
</taxonomicName>
S. 2008-2009//Probable
<taxonomicName id="5EC4408C14C81DE5D134B128B84C044F" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="431DE91B902BF10A21E918D2234B7DEB" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
C. Audibert 2016&quot; (47030749); &quot;Goias
<normalizedToken id="4CAC51E441115570B244D8B91E0765F2" originalValue="Goiañia">Goiania</normalizedToken>
coll. - SOULA//Paratype
<taxonomicName id="E91326BF90271E780C0D3DFB6AC8EAF5" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota grossiorum" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="B4ED76DAB2FA1BC80EA24AD7DA078CF9" italics="true" pageId="0" pageNumber="1">Pelidnota grossiorum</emphasis>
</taxonomicName>
S. 2008-2009//Probable
<taxonomicName id="FF88B66CE082A2432AD0828F70522C31" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="37C42903894800D53B31DF2E10070A54" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
C. Audibert 2016&quot; (47030750); five paratypes with identical label data: &quot;Goias
<normalizedToken id="9D62A09A88B799A84FDF5803E5241A9E" originalValue="Goiañia">Goiania</normalizedToken>
coll. - SOULA//Paratype
<taxonomicName id="3C22D467A8581AA72CF1DA9348F38AD8" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="EB51A03F0C06A4D537404ABD08DC5A65" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009&quot; (47030751 to 47030755); three paratypes with identical label data: &quot;Goias
<normalizedToken id="E1C6B2A175BDCD049B0B41741B8563C3" originalValue="Goiañia">Goiania</normalizedToken>
11/93 coll. - SOULA//Paratype
<taxonomicName id="E09AB26AF7C42353B7C54F43B355F77E" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="E477BCB73C698D0732084666B4560940" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009&quot; (47030756 to 47030758); &quot;Goiar K.P. Klausen 30/11 15//ZOOL. MUSEUM DK COPENHAGEN//Paratype
<taxonomicName id="6D0DE4E40F825614264FFFD35DE387ED" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="A7BB2C9BD0B099E169E97C2C256774D7" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009&quot; (47030759); two paratypes with identical label data: &quot;Goiar K.P. Klausen 30/11 1915//ZOOL. MUSEUM DK COPENHAGEN//Paratype
<taxonomicName id="77B625C4CCF113B6EA1806D4D0F0B3A3" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="C4BB07B169E236A2CCF6116422051C3A" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009&quot; (47030760 to 47030761).
<emphasis id="129CF7DAD2945DF54A8E8DD94C0D13CD" bold="true" pageId="0" pageNumber="1">Neotype here designated and deposited at CCECL</emphasis>
: &quot;Goiana, Goyas
<normalizedToken id="624206AEA18AB9F7027502532D550410" originalValue="Brésil">Bresil</normalizedToken>
, IX-X/95//Paratype
<taxonomicName id="1FAD7BB94330091FAF06B015B6F39261" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="459C443C3F15C1DABCFD5CA611D15548" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009//Neotype
<taxonomicName id="1DB3ABF93C0672CBAD507AEC44C9C132" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="A445C194E496FDD2240036DC7F08F12B" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
<bibRefCitation id="84669AEA12C6447DC31F64A6655FC0B9" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
C. Audibert des.&quot; (47030731) (Fig.
<figureCitation id="FA5DBB1637209AC98620F2FD9F21349F" captionStart="Figure 90" captionStartId="F90" captionText="Figure 90. Pelidnota thiliezi male, neotype from CCECL. A Dorsal habitus B Lateral habitus C Specimen labels D Male genitalia, lateral view E Male genitalia, dorsal view." figureDoi="10.3897/zookeys.666.9191.figure90" httpUri="https://binary.pensoft.net/fig/130518" pageId="0" pageNumber="1">90</figureCitation>
). Genitalia card-mounted underneath the male neotype and eight male paratypes. Box 4618680 SOULA. The following specimens are deposited at CMNC. 5 ♂ paratypes: &quot;BRASIL GOIAS Jatay Oliveira - leg. Coll.
<normalizedToken id="C6055ED8B69A41CE0DF313B3E65890B0" originalValue="Martínez">Martinez</normalizedToken>
. Nov.-972//H. &amp; A. HOWDEN COLLECTION
<emphasis id="E9483B6D832579BD15461CC4225E302E" italics="true" pageId="0" pageNumber="1">ex.</emphasis>
A. Martinez coll.//Paratype
<taxonomicName id="D599C00705A38F1B0221283ECE0E02FC" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="4B48389A6486F1B5DC7DA416264AFC54" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
S. 2008-2009&quot;.
</paragraph>
<paragraph id="D657ECFFC06E7B20168BAF93957BA72B" pageId="0" pageNumber="1">
<emphasis id="230CB72118D3652C826B0D5305DB120F" bold="true" pageId="0" pageNumber="1">Diagnosis.</emphasis>
<bibRefCitation id="6B512427AC0899C2E0B4AE8A802EF9A1" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
described
<taxonomicName id="4092B06D5A71BA95B82E1711AAC7CA15" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="390F2045F0F0F656BE30690E6070E901" italics="true" pageId="0" pageNumber="1">P. thiliezi</emphasis>
</taxonomicName>
as a
<normalizedToken id="6EEE66431ADFF98E8EC10DF14F965BEB" originalValue="“population”">&quot;population&quot;</normalizedToken>
from
<normalizedToken id="E7582C3409C8E6DE595C2A900A663C9D" originalValue="Goías">Goias</normalizedToken>
, Brazil that is &quot;close to&quot;
<taxonomicName id="3BA8C4C81453FFCF2172FF9C01FB9A39" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. estebanabadiei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estebanabadiei">
<emphasis id="B9A0A7D63A6D4C5585234A56925F787F" italics="true" pageId="0" pageNumber="1">P. estebanabadiei</emphasis>
</taxonomicName>
from Brazil, Ecuador, and Colombia.
<bibRefCitation id="7C64557E1D88713EE9815525B690AC99" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
stated that
<taxonomicName id="78FEF28DAD390FA2BF578AA4CED4766D" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="03F185D2ACB496294AA76EE2ECF51BA9" italics="true" pageId="0" pageNumber="1">P. thiliezi</emphasis>
</taxonomicName>
shares similarities with
<taxonomicName id="F3306D08130F8433E6ACCFDE7B9E6313" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. ancilla" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ancilla">
<emphasis id="A4DCF9BC45CE71369DF81A59648CBD9B" italics="true" pageId="0" pageNumber="1">P. ancilla</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="5E616EAF753B2E5A01F81367126E9BA5" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pelidnota ancilla F. Bates male holotype from BMNH. A Dorsal habitus B Lateral habitus C Specimen labels and male genitalia D Male genitalia, lateral view E Male parameres, dorsal view." figureDoi="10.3897/zookeys.666.9191.figure53" httpUri="https://binary.pensoft.net/fig/130481" pageId="0" pageNumber="1">53</figureCitation>
), and it differs only in a few characters which we consider to be highly variable within species. According to
<bibRefCitation id="7194DBB23FE5EE31182ABD2A7C6642EE" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
,
<taxonomicName id="C0C764B27CE954501A74240423C93979" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="2AB34DAA34B4BFF430460665E87E304B" italics="true" pageId="0" pageNumber="1">P. thiliezi</emphasis>
</taxonomicName>
is darker than
<taxonomicName id="E270246047C68266AD0574CC30CBADD1" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. ancilla" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ancilla">
<emphasis id="FA5E28E4497D48BC42D6966A3C2E8652" italics="true" pageId="0" pageNumber="1">P. ancilla</emphasis>
</taxonomicName>
. In comparison to
<taxonomicName id="4BBB63F3E620850DB591DBD2BE58CB7C" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. ancilla" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ancilla">
<emphasis id="6373AC0BEC705D0D7BDE394F5BFD73F6" italics="true" pageId="0" pageNumber="1">P. ancilla</emphasis>
</taxonomicName>
, the clypeus of the male is &quot;rather short, subtrapezoidal, with a subtruncate anterior margin, somewhat large, a little reflexed, and quite distinctly concave from behind&quot; and the clypeus of the female is &quot;more elongated, broadly parabolic rather than truncate&quot; (
<bibRefCitation id="1C201C7687F613786F17ABC48E527EF1" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
: 112, translated from French). In addition,
<bibRefCitation id="DD2AA361B3588A2CBD82DD1A3941EB6D" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
stated that the mandibular teeth of the male are smaller than those of
<taxonomicName id="FA5F7E3CD26E859868FD46F77FA6690F" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. estebanabadiei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estebanabadiei">
<emphasis id="6DD7F8E7EE5BB90CA2F11AEBB5708E3B" italics="true" pageId="0" pageNumber="1">P. estebanabadiei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8C1257DD543A9759E18E0895934FD372" pageId="0" pageNumber="1" type="remarks">
<paragraph id="3429A5C26CE07E57A9337FAA09B32F11" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="73095C508BE52E7B16CF8D9E2D00E375" pageId="0" pageNumber="1">
<bibRefCitation id="67F8EFF4BA0DA9CA7579A931AAF95245" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
indicated that he deposited the holotype specimen of
<taxonomicName id="B9629CFFC3936FB84E1C1945F3236780" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="0108503D292326023B03A6529AB898CA" italics="true" pageId="0" pageNumber="1">P. thiliezi</emphasis>
</taxonomicName>
at CCECL. Our study of the pelidnotine specimens at CCECL revealed that the holotype of this species is missing and it is presumed lost. We designated a neotype at CCECL (data above) from
<bibRefCitation id="68F2A709D779B80338C5E1BF93F8F6CC" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">
<normalizedToken id="0056BFFFAC9AA169D1FA73C5EDAADDE6" originalValue="Soula’s">Soula's</normalizedToken>
(2009)
</bibRefCitation>
paratype series in order to clarify the taxonomic status of
<taxonomicName id="E8F8DC30BF42DF2AEF205DAC0D23F52D" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="0A6C5AEED8E0D8973E75A1216BC7D892" italics="true" pageId="0" pageNumber="1">P. thiliezi</emphasis>
</taxonomicName>
and to secure the stability of nomenclature. The neotype is from the type locality of
<taxonomicName id="5620AA3064A959B3AEFA143245B17337" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="534B3D3CC6E8C5206143406E06BA01C3" italics="true" pageId="0" pageNumber="1">P. thiliezi</emphasis>
</taxonomicName>
(&quot;Goiana, Goias&quot;) (
<bibRefCitation id="1651A212B691C6A801A2E897DF81774E" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
). We examined all of the available
<taxonomicName id="B751048D49EF3F944E8677824EA9B938" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="83DDB5F0DAD2463C0E445E72E919AC8E" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
paratype material at CCECL. Features of the neotype specimen correspond closely to the original description (
<bibRefCitation id="1F279DF41D1E97C85DB167F80AA1BB90" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
), and the specimen shares the clypeal shape, coloration, and paramere morphology of the lost holotype.
</paragraph>
<caption id="E087187EDBAA26DD783E783D8DD8026A" ID-Zenodo-Dep="982489" doi="10.3897/zookeys.666.9191.figure90" httpUri="https://binary.pensoft.net/fig/130518" pageId="0" pageNumber="1" start="Figure 90" startId="F90">
<paragraph id="F5A36FF09051EEF6D1D88DB949BF8AE6" pageId="0" pageNumber="1">
<emphasis id="0AD9441A7160EA01E9F0F7E7507F87FD" bold="true" pageId="0" pageNumber="1">Figure 90.</emphasis>
<taxonomicName id="B4C1FE26D1B9AF0E34E844540ECDB3B8" authorityName="Soula" authorityYear="2009" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota thiliezi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thiliezi">
<emphasis id="33FF7ACAFE6832F85DCD18899CF38073" italics="true" pageId="0" pageNumber="1">Pelidnota thiliezi</emphasis>
</taxonomicName>
male, neotype from CCECL.
<emphasis id="5F628F16193D86255E0C7AC7E4756104" bold="true" pageId="0" pageNumber="1">A</emphasis>
Dorsal habitus
<emphasis id="9AC22AD0CEEC0B4CEFB2630E0529FC1C" bold="true" pageId="0" pageNumber="1">B</emphasis>
Lateral habitus
<emphasis id="E7F7FD9DE86DC0B9209765C1CEEE76AB" bold="true" pageId="0" pageNumber="1">C</emphasis>
Specimen labels
<emphasis id="8E5522EA3983CA046268DA1A4D4A5A95" bold="true" pageId="0" pageNumber="1">D</emphasis>
Male genitalia, lateral view
<emphasis id="E238C498092736784CE5BFDEC2B46D5C" bold="true" pageId="0" pageNumber="1">E</emphasis>
Male genitalia, dorsal view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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