449 lines
57 KiB
XML
449 lines
57 KiB
XML
<document id="D776212DE10E8BB4C69F85C2C82136FC" ID-CLB-Dataset="255197" ID-DOI="10.11646/zootaxa.5271.1.4" ID-GBIF-Dataset="a18e1635-652e-4a9f-9c01-145260e7de24" ID-ISSN="1175-5326" ID-Zenodo-Dep="7864357" ID-ZooBank="82D6F656-55DD-4DEB-84D8-BBB888E7B22E" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1682331210569" checkinUser="plazi" docAuthor="Dantas, Galileu P. S., Hamada, Neusa & Giłka, Wojciech" docDate="2023" docId="A41C0621FFBBD757FF68FA2BE6A9FE4D" docLanguage="en" docName="zootaxa.5271.1.4.pdf" docOrigin="Zootaxa 5271 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5271.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus aries Dantas & Hamada & Giłka 2023, sp. nov." docType="treatment" docUuid="19965FA3-4F73-4F16-BAAC-9EC8C72608EA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="120" masterDocId="58257E59FFBAD752FFFFFFA8E26FFFB8" masterDocTitle="Tanytarsus van der Wulp (Chironomidae, Diptera): new species from the western Amazon region in Peru and Brazil, new records from the Neotropics, and remarks on the taxonomy of the genus" masterLastPageNumber="139" masterPageNumber="115" pageNumber="116" updateTime="1699514847129" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="64C6B29CA489C7DFE53F0B258C684419">Tanytarsus van der Wulp (Chironomidae, Diptera): new species from the western Amazon region in Peru and Brazil, new records from the Neotropics, and remarks on the taxonomy of the genus</mods:title>
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<mods:namePart id="80726A954057FC76C3A11417F02CE98C">Dantas, Galileu P. S.</mods:namePart>
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<mods:affiliation id="C296E0F8F67D3552DA56D71FFF222213">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt); Av. André Araújo, 2936, 69067 - 375, Manaus, Amazonas, Brazil. & galileu. psd @ gmail. com; https: // orcid. org / 0000 - 0002 - 9155 - 533 X</mods:affiliation>
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<mods:nameIdentifier id="509BE0B225666174B483AE2CF3A00C8B" type="email">galileu.psd@gmail.com</mods:nameIdentifier>
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<mods:namePart id="686BE43AADA747F0CA05B9E41E3C418E">Hamada, Neusa</mods:namePart>
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<mods:affiliation id="16869E6584249744A0BFEE5F9F9169E6">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade (CoBio), Divisão de Curso em Entomologia (DiEnt); Av. André Araújo, 2936, 69067 - 375, Manaus, Amazonas, Brazil. & nhamada @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 3526 - 5426</mods:affiliation>
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<mods:nameIdentifier id="759673B036E4E67840913B9020565768" type="email">nhamada@inpa.gov.br</mods:nameIdentifier>
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<mods:namePart id="3DFB33D21A0C605FF13C06510AD8A379">Giłka, Wojciech</mods:namePart>
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<mods:affiliation id="C0BC9313EC02BF04DF344247C60B9856">University of Gdańsk, Faculty of Biology, Department of Invertebrate Zoology and Parasitology, Laboratory of Systematic Zoology; Wita Stwosza 59, 80 - 308 Gdańsk, Poland. wojciech. gilka @ ug. edu. pl; https: // orcid. org / 0000 - 0002 - 8403 - 5432</mods:affiliation>
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<mods:nameIdentifier id="470875A5187681589BEF9D16963F248C" type="email">wojciech.gilka@ug.edu.pl</mods:nameIdentifier>
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<mods:title id="337D69B0FD4A9C393748EB9B2BBBB6FE">Zootaxa</mods:title>
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<mods:date id="C1F8612BC713C657D2051C86EA0744F7">2023</mods:date>
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<taxonomicName id="EBB5CCB4FFBBD753FF68FA2BE331FA25" ID-CoL="BDMMC" authority="Dantas & Hamada & Giłka, 2023" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[151,350,1411,1437]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="aries" status="sp. nov.">
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<emphasis id="1EC16B25FFBBD753FF68FA2BE331FA25" bold="true" box="[151,350,1411,1437]" italics="true" pageId="1" pageNumber="116">Tanytarsus aries</emphasis>
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https://zoobank.org/
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(
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)
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<emphasis id="1EC16B25FFBBD753FF68F9BBE32BF994" bold="true" box="[151,324,1554,1581]" pageId="1" pageNumber="116">Type material.</emphasis>
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<typeStatus id="F30E0995FFBBD753FEB5F9BBE3DDF995" box="[330,434,1555,1581]" pageId="1" pageNumber="116" type="holotype">Holotype</typeStatus>
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♁,
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,
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<collectingRegion id="EE7179D5FFBBD753FDCFF9BBE016F995" box="[560,633,1555,1581]" country="Peru" name="Cusco" pageId="1" pageNumber="116">Cusco</collectingRegion>
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,
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,
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<location id="296AE1ECFFBBD753FCFAF9BBE1ECF995" LSID="urn:lsid:plazi:treatment:A41C0621FFBBD757FF68FA2BE6A9FE4D:296AE1ECFFBBD753FCFAF9BBE1ECF995" box="[773,899,1555,1581]" country="Peru" latitude="-13.336111" longLatPrecision="21" longitude="-70.84917" municipality="Quincemil" name="Araza river" pageId="1" pageNumber="116" stateProvince="Cusco">Araza river</location>
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tributary,
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,
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a.s.l.
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</elevation>
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,
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<date id="580B91F7FFBBD753FF68F99FE324F9E9" box="[151,331,1591,1617]" pageId="1" pageNumber="116" value="2012-08-23" valueMax="2012-08-31" valueMin="2012-08-23">
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<collectingDate id="484F681FFFBBD753FF68F99FE324F9E9" box="[151,331,1591,1617]" pageId="1" pageNumber="116" value="2012-08-23" valueMax="2012-08-31" valueMin="2012-08-23">23–31.viii.2012</collectingDate>
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</date>
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,
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<collectingMethod id="F5F4CF20FFBBD753FEA8F99FE38AF9E9" box="[343,485,1591,1617]" pageId="1" pageNumber="116">Malaise trap</collectingMethod>
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,
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<collectorName id="8140D2E1FFBBD753FE0EF99FE01DF9E9" box="[497,626,1591,1617]" pageId="1" pageNumber="116">J.A. Rafael</collectorName>
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,
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<collectionCode id="4AA42FF2FFBBD753FD81F99FE0FBF9E9" box="[638,660,1591,1617]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="116">R</collectionCode>
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.
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<collectionCode id="4AA42FF2FFBBD753FD68F99FE0C2F9E9" box="[663,685,1591,1617]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="116">R</collectionCode>
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.
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<collectorName id="8140D2E1FFBBD753FD47F99FE11CF9E9" box="[696,883,1591,1617]" pageId="1" pageNumber="116">Cavichioli, D.M</collectorName>
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. Takiya (
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<collectionCode id="4AA42FF2FFBBD753FC26F99FE65AF9E9" box="[985,1077,1591,1617]" pageId="1" pageNumber="116">MUSM</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="9CDDBD6AFFBBD753FBB7F99FE387F9CD" ID-GBIF-Occurrence="4090939329" collectingDate="2012-08-23" collectingDateMax="2012-08-31" collectingDateMin="2012-08-23" collectionCode="MUSM, INPA" collectorName="J. A. Rafael & Cavichioli, D. M" country="Peru" elevation="874" latitude="-13.336111" location="Araza river" longLatPrecision="21" longitude="-70.84917" municipality="Quincemil" pageId="1" pageNumber="116" specimenCount="1" stateProvince="Cusco" typeStatus="paratype">
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<typeStatus id="F30E0995FFBBD753FBB7F99FE6D6F9E9" box="[1096,1209,1591,1617]" pageId="1" pageNumber="116" type="paratype">Paratypes</typeStatus>
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: 5 ♁♁ (2
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<collectionCode id="4AA42FF2FFBBD753FAD5F99FE7EDF9E9" box="[1322,1410,1591,1617]" pageId="1" pageNumber="116">MUSM</collectionCode>
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, 3
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<collectionCode id="4AA42FF2FFBBD753FF68F9F3E2B4F9CD" box="[151,219,1627,1653]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="116">INPA</collectionCode>
|
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), same data as holotype
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||
</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="64AFE4BCFFBBD753FF38F9D7E109F905" pageId="1" pageNumber="116" type="etymology">
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<paragraph id="2C0AB737FFBBD753FF38F9D7E109F905" blockId="1.[151,1437,1554,1905]" pageId="1" pageNumber="116">
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<emphasis id="1EC16B25FFBBD753FF38F9D7E3CFF921" bold="true" box="[199,416,1663,1689]" pageId="1" pageNumber="116">
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<emphasis id="1EC16B25FFBBD753FF38F9D7E3F6F921" bold="true" box="[199,409,1663,1689]" italics="true" pageId="1" pageNumber="116">Derivatio nominis</emphasis>
|
||
.
|
||
</emphasis>
|
||
From Latin, in reference to the hypopygial anal point, in the lateral aspect resembling a horned head of a male sheep/ram (
|
||
<figureCitation id="B48EABB2FFBBD753FDE6F90BE01BF905" box="[537,628,1699,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="1" pageNumber="116">Fig. 2D</figureCitation>
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||
). Noun in apposition.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64AFE4BCFFBBD753FF38F96EE672F8C9" pageId="1" pageNumber="116" type="diagnosis">
|
||
<paragraph id="2C0AB737FFBBD753FF38F96EE672F8C9" blockId="1.[151,1437,1554,1905]" pageId="1" pageNumber="116">
|
||
<emphasis id="1EC16B25FFBBD753FF38F96EE32DF958" bold="true" box="[199,322,1734,1760]" pageId="1" pageNumber="116">Diagnosis.</emphasis>
|
||
AR ≤ 0.30. Tergite IX with microtrichia-free area near base of anal point, tergite bands Y-shaped. Anal point stout, triangular, crests broad, flanking large horn-like bars curved and turned up in proximal sections. Superior volsella subrectangular, with posteromedian corner slightly projected, bearing small ventral lip; digitus finger-like, not reaching margin of superior volsella. Median volsella with several setiform and single small foliate lamella. Inferior volsella with posteromedially directed head bearing dorsal flap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="78CAE7BFFFB8D750FF68F830E1A2F808" ID-DOI="http://doi.org/10.5281/zenodo.7864361" ID-Zenodo-Dep="7864361" box="[151,973,1944,1968]" httpUri="https://zenodo.org/record/7864361/files/figure.png" pageId="2" pageNumber="117" startId="2.[151,250,1944,1968]" targetBox="[151,1435,179,1918]" targetPageId="2" targetType="figure">
|
||
<paragraph id="2C0AB737FFB8D750FF68F830E1A2F808" blockId="2.[151,973,1944,1968]" box="[151,973,1944,1968]" pageId="2" pageNumber="117">
|
||
<emphasis id="1EC16B25FFB8D750FF68F830E37CF808" bold="true" box="[151,275,1944,1968]" pageId="2" pageNumber="117">FIGURE 1.</emphasis>
|
||
Sites of sampling the type material in Peru (A, B) and Brazil (C, D).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="64AFE4BCFFB9D757FF38FF3EE6A9FE4D" lastPageId="5" lastPageNumber="120" pageId="3" pageNumber="118" type="description">
|
||
<paragraph id="2C0AB737FFB9D751FF38FF3EE040FF09" blockId="3.[151,1437,150,825]" box="[199,559,150,177]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FF3EE336FF08" bold="true" box="[199,345,150,176]" pageId="3" pageNumber="118">Description.</emphasis>
|
||
Adult male (n = 6)
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FF13E096FF41" blockId="3.[151,1437,150,825]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FF13E39FFF6C" box="[199,496,187,213]" italics="true" pageId="3" pageNumber="118">Body size and proportions</emphasis>
|
||
. Total length
|
||
<quantity id="EB4D1AD2FFB9D751FD70FF12E159FF6D" box="[655,822,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" metricValueMax="2.21" metricValueMin="2.05" pageId="3" pageNumber="118" unit="mm" value="2.13" valueMax="2.21" valueMin="2.05">2.05–2.21 mm</quantity>
|
||
. Wing length
|
||
<quantity id="EB4D1AD2FFB9D751FC2AFF13E613FF6D" box="[981,1148,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.055" metricValueMax="1.07" metricValueMin="1.04" pageId="3" pageNumber="118" unit="mm" value="1.055" valueMax="1.07" valueMin="1.04">1.04–1.07 mm</quantity>
|
||
. Total length/wing length 1.97–2.06. Wing length/length of profemur 2.12–2.30.
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FEABE73AFEDD" blockId="3.[151,1437,150,825]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FEABE33FFEA5" box="[199,336,259,285]" italics="true" pageId="3" pageNumber="118">Colouration</emphasis>
|
||
. Head capsule and palps yellow to light brown, eyes mostly pale brown, basal portion black, antenna brown. Scutal vittae and postnotum light brown, ground colour of thorax, scutellum, sternum, and haltere yellow to faint brown. Legs yellow to light brown. Wing veins yellow, membrane pale. Abdomen yellowish.
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FEC7E720FE4D" blockId="3.[151,1437,150,825]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FEC7E36BFE31" box="[199,260,367,393]" italics="true" pageId="3" pageNumber="118">Head</emphasis>
|
||
. Eyes bare, with well-developed dorsomedian extensions. Antenna with 13 flagellomeres; ultimate flagellomere 122–125 μm long; AR 0.28–0.30. Frontal tubercles 7–8 μm long. Tentorium 93–100 μm long, with elongate digitiform apex. Temporal setae 7–9 on each side. Clypeus with 10–13 setae. Lengths of palpomeres 1–5 (in μm): 20–25, 23–26, 78–85, 92–94, 143; third palpomere with 2 sensilla clavata subapically, 12 μm long.
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FE57E1A8FD85" blockId="3.[151,1437,150,825]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FE57E377FDA1" box="[199,280,511,537]" italics="true" pageId="3" pageNumber="118">Thorax</emphasis>
|
||
. Ac 14–18, restricted to anterior region of scutum; Dc 5–6 on each side, uniserial; Pa 1 on each side; Scts 4. Scutum projected and rounded anteriorly, overreaching antepronotum.
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FDEFE3F9FD11" blockId="3.[151,1437,150,825]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FDEFE36EFDD8" box="[199,257,583,608]" italics="true" pageId="3" pageNumber="118">Wing</emphasis>
|
||
. Obovate, with anal lobe strongly reduced. Almost all veins (except subcosta) and entire membrane posterior to radial veins area (except 1/5 basal of m and cu cells) covered with macrotrichia. Brachiolum with 1 seta. VRCu 1.40–1.46.
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FD1BE134FC81" blockId="3.[151,1437,150,825]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FD1BE293FD74" box="[199,252,691,716]" italics="true" pageId="3" pageNumber="118">Legs</emphasis>
|
||
. Foreleg tibia with short lanceolate spur 16–19 μm long. Tibial combs of mid and hind legs separated; spurs of mid leg unequal: one apically curved, 20–22 μm long, second straight, 12–16 μm long; spurs of hind leg unequal: one apically curved, 25–26 μm long, second straight, 16–17 μm long. Basitarsus of mid leg without sensilla chaetica. Lengths and proportions of legs as in
|
||
<tableCitation id="6137828CFFB9D751FCFDFCB7E13AFC81" box="[770,853,799,825]" captionStart="TABLE 1" captionStartId="3.[152,244,866,890]" captionText="TABLE 1. Lengths (in μm) and proportions of leg segments of Tanytarsus aries sp. nov., male (n = 6)." pageId="3" pageNumber="118">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="78CAE7BFFFB9D751FF67FCCAE689FCC2" box="[152,1254,866,890]" pageId="3" pageNumber="118" startId="3.[152,244,866,890]" targetType="table">
|
||
<paragraph id="2C0AB737FFB9D751FF67FCCAE689FCC2" blockId="3.[152,1254,866,890]" box="[152,1254,866,890]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF67FCCAE360FCC2" bold="true" box="[152,271,866,890]" pageId="3" pageNumber="118">TABLE 1.</emphasis>
|
||
Lengths (in μm) and proportions of leg segments of
|
||
<taxonomicName id="EBB5CCB4FFB9D751FCBCFCCBE19CFCC1" authority="Dantas & Hamada & Giłka, 2023" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[835,1011,866,890]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="aries" status="sp. nov.">
|
||
<emphasis id="1EC16B25FFB9D751FCBCFCCBE19CFCC1" box="[835,1011,866,890]" italics="true" pageId="3" pageNumber="118">Tanytarsus aries</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1EC16B25FFB9D751FC05FCCAE620FCC2" bold="true" box="[1018,1103,866,890]" pageId="3" pageNumber="118">
|
||
<taxonomicNameLabel id="05F2D65EFFB9D751FC05FCCAE620FCC2" box="[1018,1103,866,890]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (n = 6).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2C0AB737FFB9D751FF27FC22E7E8FBA3" pageId="3" pageNumber="118">
|
||
<table id="5EB54597FFB928ADFF59FC22E7E7FB99" box="[166,1416,906,1057]" colsContinueIn="7.[166,1414,192,343]" gridcols="11" gridrows="4" pageId="3" pageNumber="118">
|
||
<tr id="9285B575FFB928ADFF59FC22E7E7FC11" box="[166,1416,906,937]" gridrow="0" pageId="3" pageNumber="118" rowspan-0="1">
|
||
<th id="D154DC09FFB928ADFF27FC22E35EFC11" box="[216,305,906,937]" gridcol="1" gridrow="0" pageId="3" pageNumber="118">fe</th>
|
||
<th id="D154DC09FFB928ADFEABFC22E3C3FC11" box="[340,428,906,937]" gridcol="2" gridrow="0" pageId="3" pageNumber="118">ti</th>
|
||
<th id="D154DC09FFB928ADFE33FC22E04BFC11" box="[460,548,906,937]" gridcol="3" gridrow="0" pageId="3" pageNumber="118">ta1</th>
|
||
<th id="D154DC09FFB928ADFDBCFC22E0F3FC11" box="[579,668,906,937]" gridcol="4" gridrow="0" pageId="3" pageNumber="118">ta2</th>
|
||
<th id="D154DC09FFB928ADFD45FC22E17CFC11" box="[698,787,906,937]" gridcol="5" gridrow="0" pageId="3" pageNumber="118">ta3</th>
|
||
<th id="D154DC09FFB928ADFCCEFC22E1E6FC11" box="[817,905,906,937]" gridcol="6" gridrow="0" pageId="3" pageNumber="118">ta4</th>
|
||
<th id="D154DC09FFB928ADFC59FC22E191FC11" box="[934,1022,906,937]" gridcol="7" gridrow="0" pageId="3" pageNumber="118">ta 5</th>
|
||
<th id="D154DC09FFB928ADFBE3FC22E6EEFC11" box="[1052,1153,906,937]" gridcol="8" gridrow="0" pageId="3" pageNumber="118">LR</th>
|
||
<th id="D154DC09FFB928ADFB62FC22E76CFC11" box="[1181,1283,906,937]" gridcol="9" gridrow="0" pageId="3" pageNumber="118">BV</th>
|
||
<th id="D154DC09FFB928ADFADDFC22E7E7FC11" box="[1314,1416,906,937]" gridcol="10" gridrow="0" pageId="3" pageNumber="118">SV</th>
|
||
</tr>
|
||
<tr id="9285B575FFB928ADFF59FC1AE7E7FC68" box="[166,1416,946,976]" gridrow="1" pageId="3" pageNumber="118">
|
||
<th id="D154DC09FFB928ADFF59FC1AE2D6FC68" box="[166,185,946,976]" gridcol="0" gridrow="1" pageId="3" pageNumber="118">p1</th>
|
||
<td id="D154DC09FFB928ADFF27FC1AE35EFC68" box="[216,305,946,976]" gridcol="1" gridrow="1" pageId="3" pageNumber="118">468–489</td>
|
||
<td id="D154DC09FFB928ADFEABFC1AE3C3FC68" box="[340,428,946,976]" gridcol="2" gridrow="1" pageId="3" pageNumber="118">240–247</td>
|
||
<td id="D154DC09FFB928ADFE33FC1AE04BFC68" box="[460,548,946,976]" gridcol="3" gridrow="1" pageId="3" pageNumber="118">648–672</td>
|
||
<td id="D154DC09FFB928ADFDBCFC1AE0F3FC68" box="[579,668,946,976]" gridcol="4" gridrow="1" pageId="3" pageNumber="118">300–341</td>
|
||
<td id="D154DC09FFB928ADFD45FC1AE17CFC68" box="[698,787,946,976]" gridcol="5" gridrow="1" pageId="3" pageNumber="118">250–273</td>
|
||
<td id="D154DC09FFB928ADFCCEFC1AE1E6FC68" box="[817,905,946,976]" gridcol="6" gridrow="1" pageId="3" pageNumber="118">190–203</td>
|
||
<td id="D154DC09FFB928ADFC59FC1AE191FC68" box="[934,1022,946,976]" gridcol="7" gridrow="1" pageId="3" pageNumber="118">100–104</td>
|
||
<td id="D154DC09FFB928ADFBE3FC1AE6EEFC68" box="[1052,1153,946,976]" gridcol="8" gridrow="1" pageId="3" pageNumber="118">2.62–2.80</td>
|
||
<td id="D154DC09FFB928ADFB62FC1AE76CFC68" box="[1181,1283,946,976]" gridcol="9" gridrow="1" pageId="3" pageNumber="118">1.50–1.64</td>
|
||
<td id="D154DC09FFB928ADFADDFC1AE7E7FC68" box="[1314,1416,946,976]" gridcol="10" gridrow="1" pageId="3" pageNumber="118">1.05–1.14</td>
|
||
</tr>
|
||
<tr id="9285B575FFB928ADFF59FC73E7E7FC41" box="[166,1416,987,1017]" gridrow="2" pageId="3" pageNumber="118">
|
||
<th id="D154DC09FFB928ADFF59FC73E2D6FC41" box="[166,185,987,1017]" gridcol="0" gridrow="2" pageId="3" pageNumber="118">p 2</th>
|
||
<td id="D154DC09FFB928ADFF27FC73E35EFC41" box="[216,305,987,1017]" gridcol="1" gridrow="2" pageId="3" pageNumber="118">515–541</td>
|
||
<td id="D154DC09FFB928ADFEABFC73E3C3FC41" box="[340,428,987,1017]" gridcol="2" gridrow="2" pageId="3" pageNumber="118">386–395</td>
|
||
<td id="D154DC09FFB928ADFE33FC73E04BFC41" box="[460,548,987,1017]" gridcol="3" gridrow="2" pageId="3" pageNumber="118">225–230</td>
|
||
<td id="D154DC09FFB928ADFDBCFC73E0F3FC41" box="[579,668,987,1017]" gridcol="4" gridrow="2" pageId="3" pageNumber="118">100–102</td>
|
||
<td id="D154DC09FFB928ADFD45FC73E17CFC41" box="[698,787,987,1017]" gridcol="5" gridrow="2" pageId="3" pageNumber="118">65–68</td>
|
||
<td id="D154DC09FFB928ADFCCEFC73E1E6FC41" box="[817,905,987,1017]" gridcol="6" gridrow="2" pageId="3" pageNumber="118">37–40</td>
|
||
<td id="D154DC09FFB928ADFC59FC73E191FC41" box="[934,1022,987,1017]" gridcol="7" gridrow="2" pageId="3" pageNumber="118">36–38</td>
|
||
<td id="D154DC09FFB928ADFBE3FC73E6EEFC41" box="[1052,1153,987,1017]" gridcol="8" gridrow="2" pageId="3" pageNumber="118">0.58–0.60</td>
|
||
<td id="D154DC09FFB928ADFB62FC73E76CFC41" box="[1181,1283,987,1017]" gridcol="9" gridrow="2" pageId="3" pageNumber="118">4.70–4.72</td>
|
||
<td id="D154DC09FFB928ADFADDFC73E7E7FC41" box="[1314,1416,987,1017]" gridcol="10" gridrow="2" pageId="3" pageNumber="118">4.00–4.07</td>
|
||
</tr>
|
||
<tr id="9285B575FFB928ADFF59FBABE7E7FB99" box="[166,1416,1027,1057]" gridrow="3" pageId="3" pageNumber="118">
|
||
<th id="D154DC09FFB928ADFF59FBABE2D6FB99" box="[166,185,1027,1057]" gridcol="0" gridrow="3" pageId="3" pageNumber="118">p3</th>
|
||
<td id="D154DC09FFB928ADFF27FBABE35EFB99" box="[216,305,1027,1057]" gridcol="1" gridrow="3" pageId="3" pageNumber="118">553–560</td>
|
||
<td id="D154DC09FFB928ADFEABFBABE3C3FB99" box="[340,428,1027,1057]" gridcol="2" gridrow="3" pageId="3" pageNumber="118">460–465</td>
|
||
<td id="D154DC09FFB928ADFE33FBABE04BFB99" box="[460,548,1027,1057]" gridcol="3" gridrow="3" pageId="3" pageNumber="118">300–305</td>
|
||
<td id="D154DC09FFB928ADFDBCFBABE0F3FB99" box="[579,668,1027,1057]" gridcol="4" gridrow="3" pageId="3" pageNumber="118">185–190</td>
|
||
<td id="D154DC09FFB928ADFD45FBABE17CFB99" box="[698,787,1027,1057]" gridcol="5" gridrow="3" pageId="3" pageNumber="118">174–178</td>
|
||
<td id="D154DC09FFB928ADFCCEFBABE1E6FB99" box="[817,905,1027,1057]" gridcol="6" gridrow="3" pageId="3" pageNumber="118">102–111</td>
|
||
<td id="D154DC09FFB928ADFC59FBABE191FB99" box="[934,1022,1027,1057]" gridcol="7" gridrow="3" pageId="3" pageNumber="118">58–63</td>
|
||
<td id="D154DC09FFB928ADFBE3FBABE6EEFB99" box="[1052,1153,1027,1057]" gridcol="8" gridrow="3" pageId="3" pageNumber="118">0.65–0.67</td>
|
||
<td id="D154DC09FFB928ADFB62FBABE76CFB99" box="[1181,1283,1027,1057]" gridcol="9" gridrow="3" pageId="3" pageNumber="118">2.46–2.52</td>
|
||
<td id="D154DC09FFB928ADFADDFBABE7E7FB99" box="[1314,1416,1027,1057]" gridcol="10" gridrow="3" pageId="3" pageNumber="118">3.39–3.42</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FBE4E3D3FA51" blockId="3.[151,1437,1099,2042]" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38FBE4E339FBDD" box="[199,342,1100,1125]" italics="true" pageId="3" pageNumber="118">Hypopygium</emphasis>
|
||
. Tergite IX covered with dense short microtrichia except for bare area near base of anal point, with two simple median setae; lateral teeth small, bilobed; tergite bands Y-shaped, fused part ~20–25 μm long, reaching anal point base (
|
||
<figureCitation id="B48EABB2FFB9D751FE3FFB39E072FB13" box="[448,541,1169,1195]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="3" pageNumber="118">Fig. 2A</figureCitation>
|
||
). Anal point stout, triangular, lateral margins with 3–4 setae, crests broad and round, flanking large (27–33 μm long) horn-like bars—strongly curved and turned up in proximal sections (
|
||
<figureCitation id="B48EABB2FFB9D751FA90FB1CE36FFB4A" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="3" pageNumber="118">Fig. 2A, C–E</figureCitation>
|
||
). Superior volsella 26–29 μm long, subrectangular, posteromedian corner slightly projected, with small ventral lip; 4–5 setae dorsally, 2 setae on median margin and 1 seta on anteroventral tubercle, microtrichia on dorsal surface absent; digitus finger-like, 12–14 μm long, not reaching median margin of superior volsella (
|
||
<figureCitation id="B48EABB2FFB9D751FAE5FAB6E2A4FAE3" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="3" pageNumber="118">Fig. 2A, B, F, G</figureCitation>
|
||
). Stem of median volsella simple, 12–13 μm long, with three setiform and one small foliate lamella (
|
||
<figureCitation id="B48EABB2FFB9D751FABFFAE9E28CFAC7" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="3" pageNumber="118">Fig. 2B, F, H, I</figureCitation>
|
||
). Inferior volsella 45–52 μm long, slightly curved, with posteromedially directed head bearing dorsal flap (
|
||
<figureCitation id="B48EABB2FFB9D751FF60FA20E375FA1A" box="[159,282,1416,1442]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="3" pageNumber="118">Fig. 2A, B</figureCitation>
|
||
). Phallapodeme 50–56 μm long; transverse sternapodeme 35–38 μm long, with small oral projections. Gonocoxite 75–82 μm long. Gonostylus 52–55 μm long, slightly swollen at mid length, tapering to slender tip. HR 1.38–1.58, HV 3.90–4.25.
|
||
</paragraph>
|
||
<paragraph id="2C0AB737FFB9D751FF38FA59E0E4F9B3" blockId="3.[151,1437,1099,2042]" box="[199,651,1521,1548]" pageId="3" pageNumber="118">Female and immature stages. Unknown.</paragraph>
|
||
<paragraph id="2C0AB737FFB9D757FF38F9BDE3E5FEA5" blockId="3.[151,1437,1099,2042]" lastBlockId="5.[151,1437,151,501]" lastPageId="5" lastPageNumber="120" pageId="3" pageNumber="118">
|
||
<emphasis id="1EC16B25FFB9D751FF38F9BDE325F997" bold="true" box="[199,330,1557,1583]" pageId="3" pageNumber="118">Taxonomy.</emphasis>
|
||
<bibRefCitation id="4824CAC6FFB9D751FEAFF9BCE06EF997" author="Sawedal, L." box="[336,513,1556,1583]" pageId="3" pageNumber="118" pagination="123 - 143" refId="ref14767" refString="Sawedal, L. (1981) Amazonian Tanytarsini II. Description of Caladomyia n. gen. and eight new species (Diptera: Chironomidae). Entomologica scandinavica, 12, 123 - 143. https: // doi. org / 10.1163 / 187631281794709890" type="journal article" year="1981">Säwedal (1981)</bibRefCitation>
|
||
proposed
|
||
<taxonomicName id="EBB5CCB4FFB9D751FD8AF9BCE091F996" authorityName="Sawedal" authorityYear="1981" box="[629,766,1556,1582]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FD8AF9BCE091F996" box="[629,766,1556,1582]" italics="true" pageId="3" pageNumber="118">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
for 18 species known at that time, which have been divided into two groups,
|
||
<emphasis id="1EC16B25FFB9D751FEA1F991E3FCF9EA" box="[350,403,1593,1618]" italics="true" pageId="3" pageNumber="118">spixi</emphasis>
|
||
and
|
||
<emphasis id="1EC16B25FFB9D751FE30F990E05AF9EA" box="[463,565,1592,1618]" italics="true" pageId="3" pageNumber="118">orellanai</emphasis>
|
||
. The division has been, however, considered unwarranted (
|
||
<bibRefCitation id="4824CAC6FFB9D751FB1BF990E70CF9EA" author="Reiff, N." box="[1252,1379,1592,1618]" pageId="3" pageNumber="118" pagination="175 - 198" refId="ref14143" refString="Reiff, N. (2000) Review of the mainly Neotropical genus Caladomyia Sawedal, 1981, with descriptions of seven new species (Insecta, Diptera, Chironomidae, Tanytarsini). Spixiana, 23, 175 - 198." type="journal article" year="2000">Reiff 2000</bibRefCitation>
|
||
) and ceased to be used (
|
||
<bibRefCitation id="4824CAC6FFB9D751FE99F9F3E01FF9CD" author="Trivinho-Strixino, S." box="[358,624,1627,1653]" pageId="3" pageNumber="118" pagination="1 - 41" refId="ref15000" refString="Trivinho-Strixino, S. (2012) A systematic review of Neotropical Caladomyia Sawedal (Diptera: Chironomidae). Zootaxa, 3495 (1), 1 - 41. https: // doi. org / 10.11646 / zootaxa. 3495.1.1" type="journal article" year="2012">Trivinho-Strixino 2012</bibRefCitation>
|
||
). Moreover, difficulties in diagnosing and delimiting or remarks on close relations between
|
||
<taxonomicName id="EBB5CCB4FFB9D751FEA2F9D6E389F920" authorityName="Sawedal" authorityYear="1981" box="[349,486,1662,1688]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FEA2F9D6E389F920" box="[349,486,1662,1688]" italics="true" pageId="3" pageNumber="118">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EBB5CCB4FFB9D751FDE9F9D7E0FFF920" box="[534,656,1663,1688]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FDE9F9D7E0FFF920" box="[534,656,1663,1688]" italics="true" pageId="3" pageNumber="118">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
have been raised (e.g.,
|
||
<bibRefCitation id="4824CAC6FFB9D751FC76F9D6E66EF920" author="Reiff, N." box="[905,1025,1662,1688]" pageId="3" pageNumber="118" pagination="175 - 198" refId="ref14143" refString="Reiff, N. (2000) Review of the mainly Neotropical genus Caladomyia Sawedal, 1981, with descriptions of seven new species (Insecta, Diptera, Chironomidae, Tanytarsini). Spixiana, 23, 175 - 198." type="journal article" year="2000">Reiff 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4824CAC6FFB9D751FBF7F9D6E6BFF920" author="Sanseverino, A. M." box="[1032,1232,1662,1688]" pageId="3" pageNumber="118" refId="ref14639" refString="Sanseverino, A. M. (2006) A review of the genus Tanytarsus van der Wulp, 1874 (Insecta, Diptera, Chironomidae) from the Neotropical Region. Dissertation in: Ludwig-Maximilians-Universitat Munchen, Fakultat fur Biologie, 312 pp. Available from: http: // edoc. ub. uni-muenchen. de / 4975 (accessed 30 February 2023)" type="book" year="2006">Sanseverino 2006</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4824CAC6FFB9D751FB27F9D6E2BCF903" author="Trivinho-Strixino, S." pageId="3" pageNumber="118" pagination="1 - 41" refId="ref15000" refString="Trivinho-Strixino, S. (2012) A systematic review of Neotropical Caladomyia Sawedal (Diptera: Chironomidae). Zootaxa, 3495 (1), 1 - 41. https: // doi. org / 10.11646 / zootaxa. 3495.1.1" type="journal article" year="2012">Trivinho-Strixino 2012</bibRefCitation>
|
||
), also based on the fossil record (
|
||
<bibRefCitation id="4824CAC6FFB9D751FDB2F909E111F903" author="Zakrzewska, M. & Gilka, W." box="[589,894,1697,1723]" pageId="3" pageNumber="118" pagination="397 - 403" refId="ref15254" refString="Zakrzewska, M. & Gilka, W. (2013) In the Eocene, the extant genus Caladomyia occurred in the Palaearctic (Diptera: Chironomidae: Tanytarsini). Polish Journal of Entomology, 82, 397 - 403. https: // doi. org / 10.2478 / pjen- 2013 - 0003" type="journal article" year="2013">Zakrzewska & Giłka 2013</bibRefCitation>
|
||
). Recently, after synonymizing
|
||
<taxonomicName id="EBB5CCB4FFB9D751FB1CF909E703F903" authorityName="Sawedal" authorityYear="1981" box="[1251,1388,1697,1723]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FB1CF909E703F903" box="[1251,1388,1697,1723]" italics="true" pageId="3" pageNumber="118">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EBB5CCB4FFB9D751FF68F96DE37EF966" box="[151,273,1733,1758]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FF68F96DE37EF966" box="[151,273,1733,1758]" italics="true" pageId="3" pageNumber="118">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
, all former
|
||
<taxonomicName id="EBB5CCB4FFB9D751FE6FF96CE076F966" authorityName="Sawedal" authorityYear="1981" box="[400,537,1732,1758]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FE6FF96CE076F966" box="[400,537,1732,1758]" italics="true" pageId="3" pageNumber="118">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
have been proposed to be placed in the
|
||
<emphasis id="1EC16B25FFB9D751FC37F96DE663F966" box="[968,1036,1733,1758]" italics="true" pageId="3" pageNumber="118">ortoni</emphasis>
|
||
group, recognized as monophyletic, although DNA sequences of only three described and named species (among 30) + those of
|
||
<specimenCount id="3AB37CBEFFB9D751FB2EF940E714F8BA" box="[1233,1403,1768,1794]" pageId="3" pageNumber="118" type="generic">two specimens</specimenCount>
|
||
of unknown
|
||
<taxonomicName id="EBB5CCB4FFB9D751FEF9F8A3E3E0F89D" authorityName="Sawedal" authorityYear="1981" box="[262,399,1803,1829]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FEF9F8A3E3E0F89D" box="[262,399,1803,1829]" italics="true" pageId="3" pageNumber="118">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
have been used in the molecular analysis (
|
||
<bibRefCitation id="4824CAC6FFB9D751FC95F8A3E67BF89D" author="Lin, X. & Stur, E. & Ekrem, T." box="[874,1044,1803,1829]" pageId="3" pageNumber="118" pagination="659 - 677" refId="ref14007" refString="Lin, X., Stur, E. & Ekrem, T. (2018) Molecular phylogeny and temporal diversification of Tanytarsus van der Wulp (Diptera: Chironomidae) support generic synonymies, a new classification and centre of origin. Systematic Entomology, 43 (4), 659 - 677. https: // doi. org / 10.1111 / syen. 12292" type="journal article" year="2018">
|
||
Lin
|
||
<emphasis id="1EC16B25FFB9D751FC66F8A4E1BEF89D" box="[921,977,1803,1829]" italics="true" pageId="3" pageNumber="118">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Phylogenetic relationships within these species still need support using integrative methods, those of molecular, based on at least the majority of described species, and their morphology. The significant heteromorphism in former
|
||
<taxonomicName id="EBB5CCB4FFB9D751FBA8F8F9E68FF8D3" authorityName="Sawedal" authorityYear="1981" box="[1111,1248,1873,1899]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FBA8F8F9E68FF8D3" box="[1111,1248,1873,1899]" italics="true" pageId="3" pageNumber="118">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
seems to reflect the full range of structural diversity found in
|
||
<taxonomicName id="EBB5CCB4FFB9D751FD56F8DDE14CF836" box="[681,803,1909,1934]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFB9D751FD56F8DDE14CF836" box="[681,803,1909,1934]" italics="true" pageId="3" pageNumber="118">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
, from relatively simple to the most sophisticated, thus the concept of one group for all these species can be perceived as tentative. Hence, we do not include
|
||
<taxonomicName id="EBB5CCB4FFB9D751FAD6F831E711F80A" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[1321,1406,1945,1970]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="aries">
|
||
<emphasis id="1EC16B25FFB9D751FAD6F831E711F80A" box="[1321,1406,1945,1970]" italics="true" pageId="3" pageNumber="118">T. aries</emphasis>
|
||
</taxonomicName>
|
||
to the
|
||
<emphasis id="1EC16B25FFB9D751FF3DF814E369F86D" box="[194,262,1980,2005]" italics="true" pageId="3" pageNumber="118">ortoni</emphasis>
|
||
group and refrain from proposing a possible division in the cluster(s) of these taxa (not an aim of this study), but we present the new species that extends the knowledge on the structural diversity. Regarding the shape of the gonostylus, anal tergite and volsellae,
|
||
<taxonomicName id="EBB5CCB4FFBFD757FD71FF3FE08DFF08" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[654,738,151,176]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="aries">
|
||
<emphasis id="1EC16B25FFBFD757FD71FF3FE08DFF08" box="[654,738,151,176]" italics="true" pageId="5" pageNumber="120">T. aries</emphasis>
|
||
</taxonomicName>
|
||
slightly resembles
|
||
<taxonomicName id="EBB5CCB4FFBFD757FC42FF3FE761FF09" authority="(Sawedal, 1981)" baseAuthorityName="Sawedal" baseAuthorityYear="1981" box="[957,1294,151,177]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="humboldti">
|
||
<emphasis id="1EC16B25FFBFD757FC42FF3FE624FF09" box="[957,1099,151,177]" italics="true" pageId="5" pageNumber="120">T. humboldti</emphasis>
|
||
(
|
||
<bibRefCitation id="4824CAC6FFBFD757FBA5FF3FE76AFF09" author="Sawedal, L." box="[1114,1285,151,177]" pageId="5" pageNumber="120" pagination="123 - 143" refId="ref14767" refString="Sawedal, L. (1981) Amazonian Tanytarsini II. Description of Caladomyia n. gen. and eight new species (Diptera: Chironomidae). Entomologica scandinavica, 12, 123 - 143. https: // doi. org / 10.1163 / 187631281794709890" type="journal article" year="1981">Säwedal, 1981</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, it differs from all former
|
||
<taxonomicName id="EBB5CCB4FFBFD757FE5EFF13E045FF6D" authorityName="Sawedal" authorityYear="1981" box="[417,554,187,213]" class="Insecta" family="Chironomidae" genus="Caladomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFBFD757FE5EFF13E045FF6D" box="[417,554,187,213]" italics="true" pageId="5" pageNumber="120">Caladomyia</emphasis>
|
||
</taxonomicName>
|
||
and other
|
||
<taxonomicName id="EBB5CCB4FFBFD757FD59FF13E14FFF6C" box="[678,800,187,212]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EC16B25FFBFD757FD59FF13E14FFF6C" box="[678,800,187,212]" italics="true" pageId="5" pageNumber="120">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the anal point, bearing broad crests and large, strongly curved horn-like bars (
|
||
<figureCitation id="B48EABB2FFBFD757FDBAFF77E0E3FF41" box="[581,652,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1910]" captionTargetBox="[172,1415,181,1862]" captionTargetId="figure-20@4.[172,1415,181,1862]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus aries sp. nov., male. A–C: hypopygium in dorsal (A), ventral (B) and lateral aspect (C); D, E: anal point in lateral (D) and dorsal aspect (E); F, G: superior volsella and digitus photographed (F) and drawn (G); H, I: median volsellae photographed (H) and drawn (I); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green)." figureDoi="http://doi.org/10.5281/zenodo.7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="5" pageNumber="120">Fig. 2</figureCitation>
|
||
). The low antennal ratio (AR 0.3 or less) is a supplementation of the diagnosis for
|
||
<taxonomicName id="EBB5CCB4FFBFD757FECFFEABE3ECFEA4" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[304,387,259,284]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="aries">
|
||
<emphasis id="1EC16B25FFBFD757FECFFEABE3ECFEA4" box="[304,387,259,284]" italics="true" pageId="5" pageNumber="120">T. aries</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="78CAE7BFFFBED756FF68F8F6E0B7F85B" ID-DOI="http://doi.org/10.5281/zenodo.7864365" ID-Zenodo-Dep="7864365" httpUri="https://zenodo.org/record/7864365/files/figure.png" pageId="4" pageNumber="119" startId="4.[151,250,1886,1910]" targetBox="[172,1415,181,1862]" targetPageId="4" targetType="figure">
|
||
<paragraph id="2C0AB737FFBED756FF68F8F6E0B7F85B" blockId="4.[151,1437,1886,2019]" pageId="4" pageNumber="119">
|
||
<emphasis id="1EC16B25FFBED756FF68F8F6E37AF8CE" bold="true" box="[151,277,1886,1910]" pageId="4" pageNumber="119">FIGURE 2.</emphasis>
|
||
<taxonomicName id="EBB5CCB4FFBED756FEE2F8F7E3A9F8CE" authority="Dantas & Hamada & Giłka, 2023" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[285,454,1887,1910]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="aries" status="sp. nov.">
|
||
<emphasis id="1EC16B25FFBED756FEE2F8F7E3A9F8CE" box="[285,454,1887,1910]" italics="true" pageId="4" pageNumber="119">Tanytarsus aries</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1EC16B25FFBED756FE31F8F7E04FF8CF" bold="true" box="[462,544,1887,1911]" pageId="4" pageNumber="119">
|
||
<taxonomicNameLabel id="05F2D65EFFBED756FE31F8F7E04FF8CF" box="[462,544,1887,1911]" pageId="4" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis id="1EC16B25FFBED756FD90F8F6E0CFF8CE" bold="true" box="[623,672,1886,1910]" pageId="4" pageNumber="119">A–C</emphasis>
|
||
: hypopygium in dorsal (
|
||
<emphasis id="1EC16B25FFBED756FC5DF8F6E1DBF8CE" bold="true" box="[930,948,1886,1910]" pageId="4" pageNumber="119">A</emphasis>
|
||
), ventral (
|
||
<emphasis id="1EC16B25FFBED756FBDDF8F7E65CF8CF" bold="true" box="[1058,1075,1887,1911]" pageId="4" pageNumber="119">B</emphasis>
|
||
) and lateral aspect (
|
||
<emphasis id="1EC16B25FFBED756FAF8F8F6E776F8CE" bold="true" box="[1287,1305,1886,1910]" pageId="4" pageNumber="119">C</emphasis>
|
||
);
|
||
<emphasis id="1EC16B25FFBED756FACEF8F7E72CF8CF" bold="true" box="[1329,1347,1887,1911]" pageId="4" pageNumber="119">D</emphasis>
|
||
,
|
||
<emphasis id="1EC16B25FFBED756FAADF8F7E70CF8CF" bold="true" box="[1362,1379,1887,1911]" pageId="4" pageNumber="119">E</emphasis>
|
||
: anal point in lateral (
|
||
<emphasis id="1EC16B25FFBED756FEC1F82BE33FF823" bold="true" box="[318,336,1923,1947]" pageId="4" pageNumber="119">D</emphasis>
|
||
) and dorsal aspect (
|
||
<emphasis id="1EC16B25FFBED756FDDDF82BE05CF823" bold="true" box="[546,563,1923,1947]" pageId="4" pageNumber="119">E</emphasis>
|
||
);
|
||
<emphasis id="1EC16B25FFBED756FDB5F82BE036F823" bold="true" box="[586,601,1923,1947]" pageId="4" pageNumber="119">F</emphasis>
|
||
,
|
||
<emphasis id="1EC16B25FFBED756FD97F82AE013F822" bold="true" box="[616,636,1922,1946]" pageId="4" pageNumber="119">G</emphasis>
|
||
: superior volsella and digitus photographed (
|
||
<emphasis id="1EC16B25FFBED756FBAEF82BE60FF823" bold="true" box="[1105,1120,1923,1947]" pageId="4" pageNumber="119">F</emphasis>
|
||
) and drawn (
|
||
<emphasis id="1EC16B25FFBED756FB13F82AE76FF822" bold="true" box="[1260,1280,1922,1946]" pageId="4" pageNumber="119">G</emphasis>
|
||
);
|
||
<emphasis id="1EC16B25FFBED756FAE8F82BE744F823" bold="true" box="[1303,1323,1923,1947]" pageId="4" pageNumber="119">H</emphasis>
|
||
,
|
||
<emphasis id="1EC16B25FFBED756FAC6F82BE72CF823" bold="true" box="[1337,1347,1923,1947]" pageId="4" pageNumber="119">I</emphasis>
|
||
: median volsellae photographed (
|
||
<emphasis id="1EC16B25FFBED756FE6BF80FE3C7F807" bold="true" box="[404,424,1959,1983]" pageId="4" pageNumber="119">H</emphasis>
|
||
) and drawn (
|
||
<emphasis id="1EC16B25FFBED756FDCDF80FE053F807" bold="true" box="[562,572,1959,1983]" pageId="4" pageNumber="119">I</emphasis>
|
||
); D–I magnified × 1.5–3 relative to A and B; superior volsella in the drawings (red) with anteroventral tubercle (yellow), and digitus (green).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2C0AB737FFBFD757FF38FE8EE6A9FE4D" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="120">
|
||
<emphasis id="1EC16B25FFBFD757FF38FE8EE0A8FEF8" bold="true" box="[199,711,294,321]" pageId="5" pageNumber="120">Geographical distribution and bionomics.</emphasis>
|
||
<taxonomicName id="EBB5CCB4FFBFD757FD2DFE8FE1E0FEF8" authorityName="Dantas & Hamada & Giłka" authorityYear="2023" box="[722,911,295,320]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="aries">
|
||
<emphasis id="1EC16B25FFBFD757FD2DFE8FE1E0FEF8" box="[722,911,295,320]" italics="true" pageId="5" pageNumber="120">Tanytarsus aries</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="F30E0995FFBFD757FB41FE8FE681FEF9" box="[1214,1262,295,321]" pageId="5" pageNumber="120">type</typeStatus>
|
||
locality in the highlands of Amazonian Forest in
|
||
<collectingCountry id="54A2F7A7FFBFD757FDDCFEE3E039FEDD" box="[547,598,331,357]" name="Peru" pageId="5" pageNumber="120">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="B48EABB2FFBFD757FD97FEE3E08AFEDD" box="[616,741,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1968]" captionTargetBox="[151,1435,179,1918]" captionTargetId="figure-20@2.[151,1435,179,1918]" captionTargetPageId="2" captionText="FIGURE 1. Sites of sampling the type material in Peru (A, B) and Brazil (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7864361" httpUri="https://zenodo.org/record/7864361/files/figure.png" pageId="5" pageNumber="120">Fig. 1A, B</figureCitation>
|
||
). All specimens were collected using a Malaise trap set over a small rocky-bottomed stream surrounded by dense vegetation. This region is known for its numerous long and narrow valleys, mountain streams and warm, humid, and rainy weather (
|
||
<bibRefCitation id="4824CAC6FFBFD757FC1EFE3BE6D9FE15" author="Pulgar-Vidal, J." box="[993,1206,403,429]" pageId="5" pageNumber="120" refId="ref14107" refString="Pulgar-Vidal, J. (1996) Geografia del Peru. Las ocho regiones naturales. La Regionalizacion Transversal. La Sabiduria Ecologica Tradicional. 10 ª Edition. Peisa, Lima, 218 pp." type="book" year="1996">Pulgar-Vidal 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4824CAC6FFBFD757FB3BFE3BE2BBFE69" author="Brack, A. E. & Mendiola, C. V." pageId="5" pageNumber="120" refId="ref13380" refString="Brack, A. E. & Mendiola, C. V. (2004) Ecologia del Peru. Ed. Bruno, Lima, 495 pp." type="book" year="2004">Brack & Mendiola 2004</bibRefCitation>
|
||
). As noted by
|
||
<bibRefCitation id="4824CAC6FFBFD757FE87FE1FE0CEFE69" author="Brack, A. E. & Mendiola, C. V." box="[376,673,439,465]" pageId="5" pageNumber="120" refId="ref13380" refString="Brack, A. E. & Mendiola, C. V. (2004) Ecologia del Peru. Ed. Bruno, Lima, 495 pp." type="book" year="2004">Brack & Mendiola (2004)</bibRefCitation>
|
||
, this ecoregion is a significant centre of endemism; however, it has been rapidly degraded by human activities, particularly those related to occupation along roads.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |