treatments-xml/data/91/5A/87/915A87D4476AB26CFF2EFD49FB685CD4.xml
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<document id="3D1D6843A6E3D4EF2DCE2614F63A7D9E" ID-CLB-Dataset="7216" ID-DOI="10.11646/zootaxa.4963.1.12" ID-GBIF-Dataset="e2e5f76c-f669-444b-a6de-48c0b68856bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="4711150" ID-ZooBank="7AE51A9D-9519-4D71-B678-05B2E9B55077" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618558294478" checkinUser="plazi" docAuthor="Čiampor, Fedor, Kodada, Ján, Bozáňová, Jana &amp; Čiamporová-Zaťovičová, Zuzana" docDate="2021" docId="915A87D4476AB26CFF2EFD49FB685CD4" docLanguage="en" docName="zootaxa.4963.1.12.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Disersus otongachi Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová 2021, sp.nov." docType="treatment" docVersion="7" lastPageNumber="197" masterDocId="6D63FFAC476BB268FFB9FF81FFC25B2D" masterDocTitle="Disersus otongachi a new species of Larainae riffle beetles from Ecuador (Coleoptera: Elmidae)" masterLastPageNumber="199" masterPageNumber="193" pageNumber="194" updateTime="1698937122776" updateUser="ExternalLinkService">
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<mods:title id="B9F4A06BE812DAC8E8BC940E968C77C3">Disersus otongachi a new species of Larainae riffle beetles from Ecuador (Coleoptera: Elmidae)</mods:title>
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<mods:namePart id="549FE22519F663BF1DF3C0E24E386E18">Čiampor, Fedor</mods:namePart>
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<mods:namePart id="8FB9EE29939E44F12D2C01CCAE65B2E9">Kodada, Ján</mods:namePart>
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<mods:namePart id="C31F3CA0F3242DF3EC8AFBFD4DA8EAC4">Bozáňová, Jana</mods:namePart>
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<mods:namePart id="CF2E43E6F413BEC601C6D3F2786742F4">Čiamporová-Zaťovičová, Zuzana</mods:namePart>
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<paragraph id="194C36C2476AB269FF2EFD49FE1F59CF" blockId="1.[151,477,711,738]" box="[151,477,711,738]" pageId="1" pageNumber="194">
<heading id="420481AE476AB269FF2EFD49FE1F59CF" bold="true" box="[151,477,711,738]" fontSize="11" level="1" pageId="1" pageNumber="194" reason="1">
<emphasis id="2B87EAD0476AB269FF2EFD49FE1F59CF" bold="true" box="[151,477,711,738]" pageId="1" pageNumber="194">
<taxonomicName id="DEF34D41476AB269FF2EFD49FE4259CF" authority="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová, 2021" authorityName="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová" authorityYear="2021" box="[151,384,711,738]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="otongachi" status="sp. nov.">
<emphasis id="2B87EAD0476AB269FF2EFD49FE4259CF" bold="true" box="[151,384,711,738]" italics="true" pageId="1" pageNumber="194">Disersus otongachi</emphasis>
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</emphasis>
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</paragraph>
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<subSubSection id="51E96549476AB269FF2EFC91FC3E5897" pageId="1" pageNumber="194" type="materials_examined">
<paragraph id="194C36C2476AB269FF2EFC91FDAE5863" blockId="1.[151,1437,784,1998]" pageId="1" pageNumber="194">
<emphasis id="2B87EAD0476AB269FF2EFC91FEF35807" bold="true" box="[151,305,784,810]" pageId="1" pageNumber="194">
<typeStatus id="C6488860476AB269FF2EFC91FF105807" box="[151,210,784,810]" pageId="1" pageNumber="194">Type</typeStatus>
locality
</emphasis>
: Rio Toachi, which is small river ca
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wide, has fast flowing white water, is slightly eutrophied, and has riffles and large boulders (
<figureCitation id="81C82A47476AB269FDA3FCB5FD9C5863" box="[538,606,820,846]" captionStart="FIGURES 59" captionStartId="3.[151,264,1981,2005]" captionTargetBox="[222,1368,202,1909]" captionTargetId="figure-21@3.[203,1384,181,1957]" captionTargetPageId="3" captionText="FIGURES 59. Disersus otongachi sp.nov.: 5) aedeagus dorsal view, 6) aedeagus ventral view, 7) aedeagus lateral view, 8) ovipositor ventral view, 9) male metatibia lateral view. Scale bars: 58: 0.1mm, 9: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711158" httpUri="https://zenodo.org/record/4711158/files/figure.png" pageId="1" pageNumber="194">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="194C36C2476AB269FF7EFCD9FC3E5897" blockId="1.[151,1437,784,1998]" pageId="1" pageNumber="194">
<emphasis id="2B87EAD0476AB269FF7EFCD9FEB2585F" bold="true" box="[199,368,856,882]" pageId="1" pageNumber="194">Type material</emphasis>
:
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<typeStatus id="C6488860476AB269FE39FCD9FE2F585F" box="[384,493,856,882]" pageId="1" pageNumber="194" type="holotype">Holotype</typeStatus>
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<specimenCount id="0FF5FD4B476AB269FE4FFCD8FDC9585E" box="[502,523,857,883]" pageId="1" pageNumber="194" type="male"></specimenCount>
(
<collectionCode id="7FE2AE07476AB269FDA5FCD9FDA9585F" box="[540,619,856,882]" pageId="1" pageNumber="194">PUCE</collectionCode>
): „
<collectingCountry id="61E47652476AB269FD30FCD9FD29585F" box="[649,747,856,882]" name="Ecuador" pageId="1" pageNumber="194">Ecuador</collectingCountry>
,
<collectingRegion id="DB37F820476AB269FD41FCD9FB6E585E" box="[760,1196,856,883]" country="Ecuador" name="Santo Domingo de los Tsachilas" pageId="1" pageNumber="194">Santo Domingo de los Tsachilas prov.</collectingRegion>
,
<collectingMunicipality id="F928ACB8476AB269FB00FCD9FAEC585F" box="[1209,1326,856,882]" pageId="1" pageNumber="194">Otongachi</collectingMunicipality>
env.,
<location id="1C2C6019476AB269FACDFCD9FF2458BB" LSID="urn:lsid:plazi:treatment:915A87D4476AB26CFF2EFD49FB685CD4:1C2C6019476AB269FACDFCD9FF2458BB" country="Ecuador" municipality="Otongachi" name="Rio Toachi" pageId="1" pageNumber="194" stateProvince="Santo Domingo de los Tsachilas">Río Toachi</location>
, 00˚1914.8” S, 78˚5708.4” W,
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a.s.l.
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,
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<collectingDate id="7D09E9EA476AB269FD4BFCFDFCA758BB" box="[754,869,892,918]" pageId="1" pageNumber="194" value="2013-10-08">10.8.2013</collectingDate>
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, light, Čiampor &amp; Čiamporová-Zaťovičová lgt.“
</materialsCitation>
,
<materialsCitation id="A99B3C9F476AB269FF2EFC21FC3B5897" ID-GBIF-Occurrence="3083772302" box="[151,1017,928,955]" collectingDate="2013-10-08" collectionCode="PUCE, CCB" country="Ecuador" elevation="861" location="Rio Toachi" municipality="Otongachi" pageId="1" pageNumber="194" specimenCount="26" specimenCount-female="15" specimenCount-male="11" stateProvince="Santo Domingo de los Tsachilas" typeStatus="paratype">
<emphasis id="2B87EAD0476AB269FF2EFC21FECD5897" bold="true" box="[151,271,928,954]" pageId="1" pageNumber="194">
<typeStatus id="C6488860476AB269FF2EFC21FECD5897" box="[151,271,928,954]" pageId="1" pageNumber="194" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="0FF5FD4B476AB269FEA4FC21FE8D5897" box="[285,335,928,954]" pageId="1" pageNumber="194" type="male">11♂</specimenCount>
,
<specimenCount id="0FF5FD4B476AB269FEE2FC21FE515896" box="[347,403,928,955]" pageId="1" pageNumber="194" type="female">15 ♀</specimenCount>
(
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,
<collectionCode id="7FE2AE07476AB269FE40FC21FDF15897" box="[505,563,928,954]" country="India" httpUri="http://grbio.org/cool/e3tf-9780" name="Central College, Bangalore" pageId="1" pageNumber="194">CCB</collectionCode>
) with the same locality data as
<typeStatus id="C6488860476AB269FC2AFC21FC3B5897" box="[915,1017,928,954]" pageId="1" pageNumber="194" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="51E96549476AB26CFF7EFC45FCFA5D2F" lastPageId="4" lastPageNumber="197" pageId="1" pageNumber="194" type="description">
<paragraph id="194C36C2476AB269FF7EFC45FB365F0B" blockId="1.[151,1437,784,1998]" pageId="1" pageNumber="194">
<emphasis id="2B87EAD0476AB269FF7EFC45FE9B58F3" bold="true" box="[199,345,964,990]" pageId="1" pageNumber="194">Description.</emphasis>
Body elongate (
<figureCitation id="81C82A47476AB269FDAEFC45FDBC58F3" box="[535,638,964,990]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="1" pageNumber="194">Figs 1, 3</figureCitation>
), CL: males ca 9.5 (8.8910.46) mm, females ca 10.2 (910.91) mm, ca 3.2 times as long as wide (CL/EW), dorsum convex, surface hairy. Body dark brown to black; scape, pedicel yellowish, trochanters paler; mesotibiae pale brown and ventral side lighter due to pale pubescence.
</paragraph>
<paragraph id="194C36C2476AB269FF7EFBB1FC1F5EA3" blockId="1.[151,1437,784,1998]" pageId="1" pageNumber="194">
Head partly retractable into thorax, densely setose, HW: males 1.56 (
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), female 1.66 (1.631.77) mm, ID: males 0.93 (0.841) mm, females 0.97 (0.931.05) mm. Labrum wider and longer than clypeus, densely covered with long yellow setae, anterior margin only very feebly emarginate in the middle, lateral angles broadly rounded; clypeuswithlongersetaealong anterior margin; frontoclypeal suture straight. Eyes large, protruding beyond outline of head, suboval in lateral view, bordered by long black setae from inner side; frons simple, without grooves or sutures. Maxillary palpus distinct, terminal palpomere longest and widest, truncate apically with microreticulate distal, flat surface. Labium membranous apically; labial palpus distinctly smaller than maxillary palpus, terminal segment robust, rounded apically, segments 1-2 with long yellow setae. Antennae with 11 antennomeres, setose, long, almost reaching base of pronotum; scape ca twice as long as pedicel or other segments, almost straight; segments 311 subequal in size, subtriangular, not forming elongated club.
</paragraph>
<paragraph id="194C36C2476AB269FF7EFA19FDAF5CE3" blockId="1.[151,1437,784,1998]" pageId="1" pageNumber="194">
<emphasis id="2B87EAD0476AB269FF7EFA19FEE55E9F" bold="true" box="[199,295,1432,1458]" pageId="1" pageNumber="194">Thorax.</emphasis>
Pronotum (
<figureCitation id="81C82A47476AB269FE10FA19FDC85E9F" box="[425,522,1432,1458]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="1" pageNumber="194">Figs 1, 3</figureCitation>
) widest at base, anterior margin straight, PW: males 2.52 (2.41-2.74) mm, females 2.64 (2.362.91) mm, PL: males 1.7 (1.621.92) mm, females 1.66 (1.451.82) mm; base broadly sinuate on each side and narrowly so in front of scutellum; posterolateral angles strongly produced, acute; two short prescutellar pits; disc convex, without median groove and sublateral carinae, sublateral grooves very finely indicated at base; sides convex, lateral margins very narrowly explanate; admedian prebasal pits shallow, slightly elongated. Hypomeron sinuate, wider at base and anterior end, constricted around procoxa. Prosternal process (
<figureCitation id="81C82A47476AB269FBC9F9CDFB175D4B" box="[1136,1237,1612,1638]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="1" pageNumber="194">Figs 2, 4</figureCitation>
) triangular, about as long as wide at base, sides slightly constricted before rounded apex; prosternum in front of coxae extremely short; surface of prosternum densely setose. Mesoventrite (
<figureCitation id="81C82A47476AB269FCFEF915FC715D83" box="[839,947,1684,1710]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="1" pageNumber="194">Figs 2, 4</figureCitation>
) short and wide, with deep narrow mesal depression; sides around coxae rised; basal margin bisinuate. Metaventrite (
<figureCitation id="81C82A47476AB269FC4AF939FB9B5DFF" box="[1011,1113,1720,1746]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="1" pageNumber="194">Figs 2, 4</figureCitation>
) smooth, discrimen distinct, reaching anterior and posterior margin. Elytra (
<figureCitation id="81C82A47476AB269FD17F95DFCD15DDB" box="[686,787,1756,1782]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="1" pageNumber="194">Figs 1, 3</figureCitation>
) with 10 rows of punctures largest on disc, sides parallel in anterior ca ¾, then more or less sharply tapering toward apices, which are spinose, around 2.2 times as long as wide; EL: males 6.54 (6.087.31) mm, females 7.09 (6.187.45) mm; EW: males 2.93 (
<quantity id="DE0B9B27476AB269FB37F8A5FAF55C12" box="[1166,1335,1828,1855]" metricMagnitude="-3" metricUnit="m" metricValue="2.98" metricValueMax="3.15" metricValueMin="2.81" pageId="1" pageNumber="194" unit="mm" value="2.98" valueMax="3.15" valueMin="2.81">2.813.15 mm</quantity>
, females 3.18 (2.823.45) mm; disc without tubercles or accessory rows; epipleuron wider along metaventrite, than narrow, setose. Scutellum subtriangular, all three sides arcuate. Legs long and slender; coxae moderately widely separated; femora and tibiae flattened; profemora widest; mesotibiae pale, metatibiae longest, with pale curved setae in males (
<figureCitation id="81C82A47476AB269FF19F835FF265CE3" box="[160,228,1972,1998]" captionStart="FIGURES 59" captionStartId="3.[151,264,1981,2005]" captionTargetBox="[222,1368,202,1909]" captionTargetId="figure-21@3.[203,1384,181,1957]" captionTargetPageId="3" captionText="FIGURES 59. Disersus otongachi sp.nov.: 5) aedeagus dorsal view, 6) aedeagus ventral view, 7) aedeagus lateral view, 8) ovipositor ventral view, 9) male metatibia lateral view. Scale bars: 58: 0.1mm, 9: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711158" httpUri="https://zenodo.org/record/4711158/files/figure.png" pageId="1" pageNumber="194">Fig. 9</figureCitation>
); tarsal claws large and acuminate.
</paragraph>
<caption id="4D8C664A4769B26AFF2EF83CFEFF5CD7" ID-DOI="http://doi.org/10.5281/zenodo.4711156" ID-Zenodo-Dep="4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="2" pageNumber="195" startId="2.[151,264,1981,2005]" targetBox="[198,1357,201,1938]" targetPageId="2">
<paragraph id="194C36C24769B26AFF2EF83CFEFF5CD7" blockId="2.[151,1437,1981,2042]" pageId="2" pageNumber="195">
<emphasis id="2B87EAD04769B26AFF2EF83CFEF95CF8" bold="true" box="[151,315,1981,2005]" pageId="2" pageNumber="195">FIGURES 14.</emphasis>
Habitus of
<taxonomicName id="DEF34D414769B26AFE0DF83FFDBA5CF8" authority="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová, 2021" authorityName="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová" authorityYear="2021" box="[436,632,1981,2005]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="otongachi" status="sp. nov.">
<emphasis id="2B87EAD04769B26AFE0DF83FFDBA5CF8" box="[436,632,1981,2005]" italics="true" pageId="2" pageNumber="195">Disersus otongachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B457AB4769B26AFDC6F83FFD095CF8" box="[639,715,1982,2005]" pageId="2" pageNumber="195" rank="species">sp.nov.</taxonomicNameLabel>
: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm.
</paragraph>
</caption>
<caption id="4D8C664A4768B26BFF2EF83CFBDA5CD7" ID-DOI="http://doi.org/10.5281/zenodo.4711158" ID-Zenodo-Dep="4711158" httpUri="https://zenodo.org/record/4711158/files/figure.png" pageId="3" pageNumber="196" startId="3.[151,264,1981,2005]" targetBox="[222,1368,202,1909]" targetPageId="3">
<paragraph id="194C36C24768B26BFF2EF83CFBDA5CD7" blockId="3.[151,1436,1981,2042]" pageId="3" pageNumber="196">
<emphasis id="2B87EAD04768B26BFF2EF83CFEFF5CF8" bold="true" box="[151,317,1981,2005]" pageId="3" pageNumber="196">FIGURES 59.</emphasis>
<taxonomicName id="DEF34D414768B26BFEFCF83FFDC85CF8" authority="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová, 2021" authorityName="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová" authorityYear="2021" box="[325,522,1981,2005]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="196" phylum="Arthropoda" rank="species" species="otongachi" status="sp. nov.">
<emphasis id="2B87EAD04768B26BFEFCF83FFDC85CF8" box="[325,522,1981,2005]" italics="true" pageId="3" pageNumber="196">Disersus otongachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B457AB4768B26BFDAAF83FFD9D5CF8" box="[531,607,1982,2005]" pageId="3" pageNumber="196" rank="species">sp.nov.</taxonomicNameLabel>
: 5) aedeagus dorsal view, 6) aedeagus ventral view, 7) aedeagus lateral view, 8) ovipositor ventral view, 9) male metatibia lateral view. Scale bars: 58: 0.1mm, 9: 1mm.
</paragraph>
</caption>
<caption id="4D8C664A476FB26CFF2EFC2FFE9E58C7" ID-DOI="http://doi.org/10.5281/zenodo.4711160" ID-Zenodo-Dep="4711160" httpUri="https://zenodo.org/record/4711160/files/figure.png" pageId="4" pageNumber="197" startId="4.[151,250,942,966]" targetBox="[177,1411,181,918]" targetPageId="4">
<paragraph id="194C36C2476FB26CFF2EFC2FFE9E58C7" blockId="4.[151,1437,942,1002]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF2EFC2FFEE358EB" bold="true" box="[151,289,942,966]" pageId="4" pageNumber="197">FIGURE 10.</emphasis>
Type locality of
<taxonomicName id="DEF34D41476FB26CFE6BFC2EFD5558EB" authority="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová, 2021" authorityName="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová" authorityYear="2021" box="[466,663,942,966]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="otongachi" status="sp. nov.">
<emphasis id="2B87EAD0476FB26CFE6BFC2EFD5558EB" box="[466,663,942,966]" italics="true" pageId="4" pageNumber="197">Disersus otongachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30B457AB476FB26CFD26FC2FFD2858E8" box="[671,746,942,965]" pageId="4" pageNumber="197" rank="species">sp.nov.</taxonomicNameLabel>
, Toachi River, Otongachi Reserve, province Santo Domingo de los Tsachilas, Ecuador.
</paragraph>
</caption>
<paragraph id="194C36C2476FB26CFF7EFB92FC0F5EB8" blockId="4.[151,1437,1043,2042]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF7EFB92FEFE5F00" bold="true" box="[199,316,1043,1069]" pageId="4" pageNumber="197">Abdomen</emphasis>
(
<figureCitation id="81C82A47476FB26CFEF5FB92FE705F00" box="[332,434,1043,1069]" captionStart="FIGURES 14" captionStartId="2.[151,264,1981,2005]" captionTargetBox="[198,1357,201,1938]" captionTargetId="figure-23@2.[193,1393,179,1955]" captionTargetPageId="2" captionText="FIGURES 14. Habitus of Disersus otongachi sp.nov.: 1) male, dorsal, 2) male, ventral, 3) female, dorsal, 4) female, ventral. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711156" httpUri="https://zenodo.org/record/4711156/files/figure.png" pageId="4" pageNumber="197">Figs 2, 4</figureCitation>
) setose; intercoxal process of first ventrite triangular, carinae of ventrite 1 behind coxae absent; ventrites transverse, ventrites 3 and 4 with posterolateral angles slightly produced, ventrite 5 about as long as ventrite 2, triangular, with apical tuft of dense yellowish setae. Aedeagus (
<figureCitation id="81C82A47476FB26CFC6AFBDAFBF55F5B" box="[979,1079,1115,1142]" captionStart="FIGURES 59" captionStartId="3.[151,264,1981,2005]" captionTargetBox="[222,1368,202,1909]" captionTargetId="figure-21@3.[203,1384,181,1957]" captionTargetPageId="3" captionText="FIGURES 59. Disersus otongachi sp.nov.: 5) aedeagus dorsal view, 6) aedeagus ventral view, 7) aedeagus lateral view, 8) ovipositor ventral view, 9) male metatibia lateral view. Scale bars: 58: 0.1mm, 9: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711158" httpUri="https://zenodo.org/record/4711158/files/figure.png" pageId="4" pageNumber="197">Figs 57</figureCitation>
) elongate, fibula absent, corona membranous in basal 1/3 of penis; parameres ca. 0.4 times as long as median lobe, widest at middle, narrowed toward rounded apex (lateral view); median lobe slender, in dorsal / ventral view asymmetrical, constricted behind middle, narrowed behind constriction, distal ca 0.6 parallel-sided, apex acuminate; in lateral view median lobe bisinuate, narrow in distal half, basal half wider; phallobase long, ca 1.3 times longer than median lobe, straight, parallel-sided in dorsal / ventral view, narrowed distally in lateral view. Ovipositor (
<figureCitation id="81C82A47476FB26CFBF5FA8EFB4D5E04" box="[1100,1167,1295,1321]" captionStart="FIGURES 59" captionStartId="3.[151,264,1981,2005]" captionTargetBox="[222,1368,202,1909]" captionTargetId="figure-21@3.[203,1384,181,1957]" captionTargetPageId="3" captionText="FIGURES 59. Disersus otongachi sp.nov.: 5) aedeagus dorsal view, 6) aedeagus ventral view, 7) aedeagus lateral view, 8) ovipositor ventral view, 9) male metatibia lateral view. Scale bars: 58: 0.1mm, 9: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4711158" httpUri="https://zenodo.org/record/4711158/files/figure.png" pageId="4" pageNumber="197">Fig. 8</figureCitation>
) with terminal segment (stylus) short, straight, slightly widened towards base; preterminal segment stout, abruptly widened basally, apically densely setose, ca 2.7 times as long as terminal segment; distal segment produced medio-basally; basal segment ca. twice as long as preterminal and distal segments combined, spurs curved.
</paragraph>
<paragraph id="194C36C2476FB26CFF7EFA1EFCFA5D2F" blockId="4.[151,1437,1043,2042]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF7EFA1EFE705E94" bold="true" box="[199,434,1439,1465]" pageId="4" pageNumber="197">Sexual dimorphism</emphasis>
: Males differ from females by slightly smaller size, longer antennae, depressed disc of metaventrite, protibia much more widened, metatibia with stout pale setae, abdominal intercoxal process depressed in males (convex in females) and excised apex of ventrite 5.
</paragraph>
</subSubSection>
<subSubSection id="51E96549476FB26CFF7EF98AFF325D64" pageId="4" pageNumber="197" type="distribution">
<paragraph id="194C36C2476FB26CFF7EF98AFF325D64" blockId="4.[151,1437,1043,2042]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF7EF98AFE9B5D08" bold="true" box="[199,345,1547,1573]" pageId="4" pageNumber="197">Distribution</emphasis>
:
<taxonomicName id="DEF34D41476FB26CFED4F98DFDC65D08" authorityName="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová" authorityYear="2021" box="[365,516,1547,1573]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="otongachi">
<emphasis id="2B87EAD0476FB26CFED4F98DFDC65D08" box="[365,516,1547,1573]" italics="true" pageId="4" pageNumber="197">D. otongachi</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="C6488860476FB26CFC8FF98AFCA45D08" box="[822,870,1547,1573]" pageId="4" pageNumber="197">type</typeStatus>
locality (
<figureCitation id="81C82A47476FB26CFC63F98AFBF35D08" box="[986,1073,1547,1573]" captionStart="FIGURE 10" captionStartId="4.[151,250,942,966]" captionTargetBox="[177,1411,181,918]" captionTargetId="figure-23@4.[177,1411,181,918]" captionTargetPageId="4" captionText="FIGURE 10. Type locality of Disersus otongachi sp.nov., Toachi River, Otongachi Reserve, province Santo Domingo de los Tsachilas, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.4711160" httpUri="https://zenodo.org/record/4711160/files/figure.png" pageId="4" pageNumber="197">Fig. 10</figureCitation>
), Rio Toachi in the Otongachi reserve.
</paragraph>
</subSubSection>
<subSubSection id="51E96549476FB26CFF7EF9D2FD2C5DBC" pageId="4" pageNumber="197" type="diagnosis">
<paragraph id="194C36C2476FB26CFF7EF9D2FD2C5DBC" blockId="4.[151,1437,1043,2042]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF7EF9D2FE055D40" bold="true" box="[199,455,1619,1645]" pageId="4" pageNumber="197">Differential diagnosis</emphasis>
.
<taxonomicName id="DEF34D41476FB26CFE6FF9D5FDAB5D40" authorityName="Čiampor &amp; Kodada &amp; Bozáňová &amp; Čiamporová-Zaťovičová" authorityYear="2021" box="[470,617,1619,1645]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="otongachi">
<emphasis id="2B87EAD0476FB26CFE6FF9D5FDAB5D40" box="[470,617,1619,1645]" italics="true" pageId="4" pageNumber="197">D. otongachi</emphasis>
</taxonomicName>
can be easily distinguished from congeners by its long antennae and the unique asymetrical median lobe in the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="51E96549476FB26CFF7EF91AFB685CD4" pageId="4" pageNumber="197" type="etymology">
<paragraph id="194C36C2476FB26CFF7EF91AFB785D98" blockId="4.[151,1437,1043,2042]" box="[199,1210,1691,1717]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF7EF91AFE8A5D98" bold="true" box="[199,328,1691,1717]" pageId="4" pageNumber="197">Etymology</emphasis>
: The new species was named after Reserva Otongachi, where it was collected.
</paragraph>
<paragraph id="194C36C2476FB26CFF7EF93EFB685CD4" blockId="4.[151,1437,1043,2042]" pageId="4" pageNumber="197">
<emphasis id="2B87EAD0476FB26CFF7EF93EFF3D5DF4" bold="true" box="[199,255,1727,1753]" pageId="4" pageNumber="197">Note</emphasis>
: Of the
<specimenCount id="0FF5FD4B476FB26CFEDEF93EFDDD5DF4" box="[359,543,1727,1753]" pageId="4" pageNumber="197" type="generic">three specimens</specimenCount>
used for molecular analysis, DNA was successfully extracted from two, the sequences of the barcoding fragment are freely available in www.boldsystems.org, records ELMSA049-16 and ELMSA398-19. Besides
<taxonomicName id="DEF34D41476FB26CFE0FF889FD4C5C0C" authority="DNA" authorityName="DNA" box="[438,654,1799,1825]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="otongachi">
<emphasis id="2B87EAD0476FB26CFE0FF889FD885C0C" box="[438,586,1799,1825]" italics="true" pageId="4" pageNumber="197">D. otongachi</emphasis>
DNA
</taxonomicName>
barcodes, five
<taxonomicName id="DEF34D41476FB26CFCF9F889FC605C0C" authorityName="Sharp" authorityYear="1882" box="[832,930,1800,1825]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="2B87EAD0476FB26CFCF9F889FC605C0C" box="[832,930,1800,1825]" italics="true" pageId="4" pageNumber="197">Disersus</emphasis>
</taxonomicName>
barcoding fragment (5 COI) sequences are currently published and freely available in databases. However, according to the voucher photo (www.boldsystems. org), two represent other taxa (MZCRI3369-13 is a larva of
<taxonomicName id="DEF34D41476FB26CFC93F8CEFC745C44" box="[810,950,1871,1897]" class="Insecta" family="Psephenidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="family">Psephenidae</taxonomicName>
and MZCRI3368-13 is an unspecified larva of
<taxonomicName id="DEF34D41476FB26CFF0CF8F2FED75CA0" authorityName="Curtis" authorityYear="1830" box="[181,277,1907,1933]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
, but it is certainly not a
<taxonomicName id="DEF34D41476FB26CFD99F8F5FD405CA0" authorityName="Sharp" authorityYear="1882" box="[544,642,1908,1933]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="2B87EAD0476FB26CFD99F8F5FD405CA0" box="[544,642,1908,1933]" italics="true" pageId="4" pageNumber="197">Disersus</emphasis>
</taxonomicName>
larva). The remaining three sequences represent 2
<taxonomicName id="DEF34D41476FB26CFB03F8F5FAB55CA0" box="[1210,1399,1907,1933]" class="Insecta" family="Elmidae" genus="Disersus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2B87EAD0476FB26CFB03F8F5FADE5CA0" box="[1210,1308,1908,1933]" italics="true" pageId="4" pageNumber="197">Disersus</emphasis>
species
</taxonomicName>
,
<taxonomicName id="DEF34D41476FB26CFA38F8F5FF055C9C" authorityName="Spangler &amp; Santiago-Fragoso" authorityYear="1987" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="2B87EAD0476FB26CFA38F8F5FF055C9C" italics="true" pageId="4" pageNumber="197">D. inca</emphasis>
</taxonomicName>
from
<collectingCountry id="61E47652476FB26CFEB6F816FEAE5C9C" box="[271,364,1943,1969]" name="Ecuador" pageId="4" pageNumber="197">Ecuador</collectingCountry>
(ELMSA045-16) and
<taxonomicName id="DEF34D41476FB26CFDC9F819FD385C9C" box="[624,762,1943,1969]" class="Insecta" family="Elmidae" genus="Disersus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2B87EAD0476FB26CFDC9F819FD315C9C" box="[624,755,1944,1969]" italics="true" pageId="4" pageNumber="197">Disersus sp</emphasis>
.
</taxonomicName>
from
<collectingCountry id="61E47652476FB26CFCFBF816FC7F5C9C" box="[834,957,1943,1969]" name="Costa Rica" pageId="4" pageNumber="197">Costa Rica</collectingCountry>
(MZCRI570-13, MZCRI577-13).
<taxonomicName id="DEF34D41476FB26CFAF1F819FA5E5C9C" authorityName="Spangler &amp; Santiago-Fragoso" authorityYear="1987" box="[1352,1436,1944,1969]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="2B87EAD0476FB26CFAF1F819FA5E5C9C" box="[1352,1436,1944,1969]" italics="true" pageId="4" pageNumber="197">D. inca</emphasis>
</taxonomicName>
is significantly phylogenetically distant from
<taxonomicName id="DEF34D41476FB26CFD2EF83DFCE85CF8" box="[663,810,1979,2005]" class="Insecta" family="Elmidae" genus="Disersus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="otongachi">
<emphasis id="2B87EAD0476FB26CFD2EF83DFCE85CF8" box="[663,810,1979,2005]" italics="true" pageId="4" pageNumber="197">D. otongachi</emphasis>
</taxonomicName>
(~ 13% K2P distance).
<taxonomicName id="DEF34D41476FB26CFB81F83DFB005CF8" box="[1080,1218,1979,2005]" class="Insecta" family="Elmidae" genus="Disersus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="197" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2B87EAD0476FB26CFB81F83DFB585CF8" box="[1080,1178,1980,2005]" italics="true" pageId="4" pageNumber="197">Disersus</emphasis>
sp.
</taxonomicName>
from
<collectingCountry id="61E47652476FB26CFABEF83AFA405CF8" box="[1287,1410,1979,2005]" name="Costa Rica" pageId="4" pageNumber="197">Costa Rica</collectingCountry>
is clearly more related (2.3-3.6%), but still sufficiently genetically different at the species level.
</paragraph>
</subSubSection>
</treatment>
</document>