182 lines
15 KiB
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182 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1087.64382" ID-GBIF-Dataset="9c9b7f37-a72e-4610-8ac5-908f601d1121" ID-PMC="PMC8888540" ID-Pensoft-Pub="1313-2970-1087-19" ID-Pensoft-UUID="ECD9B4DC2A3357AABC04DB88FB7D40B1" ID-PubMed="35437365" ID-ZooBank="94F2384E640E4A58B8B4D9D06675D2C2" ModsDocID="1313-2970-1087-19" checkinTime="1645627321867" checkinUser="pensoft" docAuthor="Nupponen 1, Kari & Sihvonen, Pasi" docDate="2022" docId="6ACCFE2530C751BC84266E5F96FC3F46" docLanguage="en" docName="ZooKeys 1087: 19-104" docOrigin="ZooKeys 1087" docPubDate="2022-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1087.64382" docTitle="Scythris directiphallella Nupponen 2022, sp. nov." docType="treatment" docUuid="24086F7E-BD7F-400B-B3D0-8D165AAEB9B6" docUuidSource="ZooBank" docVersion="4" id="ECD9B4DC2A3357AABC04DB88FB7D40B1" lastPageNumber="19" masterDocId="ECD9B4DC2A3357AABC04DB88FB7D40B1" masterDocTitle="Revision of Neotropical Scythrididae moths and descriptions of 22 new species from Argentina, Chile, and Peru (Lepidoptera, Gelechioidea)" masterLastPageNumber="104" masterPageNumber="19" pageNumber="19" updateTime="1668151506650" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Neotropical Scythrididae moths and descriptions of 22 new species from Argentina, Chile, and Peru (Lepidoptera, Gelechioidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nupponen 1, Kari</mods:namePart>
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<mods:affiliation>Merenneidontie 19 D, FI- 02320 Espoo, Finland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sihvonen, Pasi</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2237-9325</mods:nameIdentifier>
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<mods:affiliation>Finnish Museum of Natural History, P. O. Box 17, Pohjoinen Rautatiekatu 13, 00014 University of Helsinki, Finland</mods:affiliation>
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<mods:nameIdentifier type="email">pasi.sihvonen@helsinki.fi</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1087</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>104</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1087.64382</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1087.64382</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1087-19</mods:identifier>
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<mods:identifier type="ZooBank">94F2384E640E4A58B8B4D9D06675D2C2</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ECD9B4DC2A3357AABC04DB88FB7D40B1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="193585887" LSID="urn:lsid:zoobank.org:act:24086F7E-BD7F-400B-B3D0-8D165AAEB9B6" httpUri="http://treatment.plazi.org/id/6ACCFE2530C751BC84266E5F96FC3F46" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="http://zoobank.org/24086F7E-BD7F-400B-B3D0-8D165AAEB9B6" authority="Nupponen" authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris directiphallella" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="directiphallella" status="sp. nov.">Scythris directiphallella Nupponen</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">
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<figureCitation captionStart="Figures 12–17" captionStartId="F3" captionText="Figures 12 - 17. Scythrididae adults, genus Scythris. 12 S. directiphallella Nupponen, sp. nov., male, holotype 13 A S. furciphallella Nupponen sp. nov., male, holotype 13 B S. furciphallella Nupponen sp. nov., male, paratype 14 S. manchaoensis Nupponen, sp. nov., male holotype 15 A S. salinasgrandensis Nupponen sp. nov., female, holotype 15 B S. salinasgrandensis Nupponen sp. nov., female, paratype 16 S. angustivalvella Nupponen sp. nov., male, holotype 17 S. zeugmatica Meyrick, 1916, male, holotype." figureDoi="10.3897/zookeys.1087.64382.figures12-17" httpUri="https://binary.pensoft.net/fig/649317" pageId="0" pageNumber="19">Figs 12</figureCitation>
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<figureCitation captionStart="Figures 43–44" captionStartId="F11" captionText="Figures 43 - 44. Male genitalia of Landryia and Scythris 43 L. chilensis Nupponen sp. nov., genus combination incertae sedis, holotype, slide 3 / 18 Dec. 2019 KN 44 S. directiphallella Nupponen sp. nov., holotype; slide 3 / 28 Dec. 2019 KN." figureDoi="10.3897/zookeys.1087.64382.figures43-44" httpUri="https://binary.pensoft.net/fig/649325" pageId="0" pageNumber="19">, 44</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type material">
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<paragraph pageId="0" pageNumber="19">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Holotype</emphasis>
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.
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</emphasis>
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Argentina • ♂; prov. Santiago del Estero, Pozo Honda village S, by salt lake;
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<geoCoordinate degrees="27" direction="south" minutes="17.2" orientation="latitude" precision="92" value="-27.286667">27°17.2'S</geoCoordinate>
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,
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<geoCoordinate degrees="64" direction="west" minutes="28.0" orientation="longitude" precision="92" value="-64.46667">64°28.0'W</geoCoordinate>
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; 260 m a.s.l.; 19 Sep. 2017; K. Nupponen & R. Haverinen leg.; [BOLD sample ID] KN01052; [genitalia slide] K. Nupponen prep. no. 3/28 Dec. 2019; coll. NUPP (MZH).
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Paratype</emphasis>
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.
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</emphasis>
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Argentina • 1 ♂; same data as for holotype; coll. NUPP.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Wings grey, impossible to separate externally from
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<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. furciphallella" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="furciphallella">
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<emphasis italics="true" pageId="0" pageNumber="19">S. furciphallella</emphasis>
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</taxonomicName>
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. The male genitalia of
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<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. directiphallella" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="directiphallella">
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<emphasis italics="true" pageId="0" pageNumber="19">S. directiphallella</emphasis>
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</taxonomicName>
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are by having narrow valvae with a ventral thorn-like process apically, a straight phallus and pincer-like extensions on posterior margin of male sternite VIII.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Wingspan 9-10.5 mm. Head, collar, neck tuft, haustellum, tegula and thorax grey. Scape grey, ventrally mixed with dirty white; pecten grey and as long as diameter of scape. Flagellum fuscous, 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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length of forewing, ciliate, sensillae ~ 1/2 as long as diameter of flagellum. Labial palp: palpomere I dirty white, palpomeres II and III fuscous mixed with dirty white. Legs fuscous, more or less suffused with dirty white. Abdomen dorsally pale grey, ventrally a little paler, anal tuft cream. Forewing grey, over the wing sparsely scattered dark fuscous scales. Hindwing pale fuscous.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Male genitalia</emphasis>
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.
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</emphasis>
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Gnathos (homology interpretation of gnathos and uncus based on
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<bibRefCitation DOI="https://doi.org/10.4039/entm123160fv" author="Landry, JF" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="19" pagination="1 - 341" refId="B22" refString="Landry, JF, 1991. Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species. Memoirs of the Entomological Society of Canada 160: 1 - 341, DOI: https://doi.org/10.4039/entm123160fv" title="Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species." url="https://doi.org/10.4039/entm123160fv" volume="160" year="1991">Landry (1991)</bibRefCitation>
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) base broad, weakly sclerotised belt; distally long, slender and bent downwards, tip pointed. Uncus bilobed plate, posterior shanks subapically with small nipple-like extensions. Tegumen elongated hood, dorsally widely open. Phallus 0.75
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<normalizedToken originalValue="×">x</normalizedToken>
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length of valva, straight, tip pointed. Valva long and narrow; apex setose and slightly incurved; apically with robust ventral thorn-like process. Saccus short, triangular. Juxta narrow, 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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length of phallus. Sternum VIII pentagonal; posteriorly bifurcate, shanks short, bent inwards with tips pointed. Tergum VIII trapezoid, anterior margin widely concave and weakly sclerotised, posterior margin convex.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="etymology">
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<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="19">Diminutive noun in apposition. The species name refers to the straight phallus of the male, which is a diagnostic character of the species.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="distribution">
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<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="19">NW Argentina.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="habitat">
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<paragraph pageId="0" pageNumber="19">Habitat.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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The collecting site is a dry, shrubby area near a salt lake shore (Fig.
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<figureCitation captionStart="Figure 77" captionStartId="F35" captionText="Figure 77. Collecting site of Rhamphura depressa Meyrick, R. pozohondaensis sp. nov., R. subdimota sp. nov., R. curvisociella sp. nov., Scythris directiphallella sp. nov., S. angustivalvella sp. nov., Landryia ankylosauroides sp. nov. Argentina, Pozo Honda vill. S (259 m), 19 Sep. 2017." figureDoi="10.3897/zookeys.1087.64382.figure77" httpUri="https://binary.pensoft.net/fig/649349" pageId="0" pageNumber="19">77</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="genetic data">
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<paragraph pageId="0" pageNumber="19">Genetic data.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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BIN: BOLD:ADY7318 (
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<emphasis italics="true" pageId="0" pageNumber="19">n</emphasis>
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= 2 from Argentina). Nearest neighbour:
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<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris salinasgrandensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="salinasgrandensis">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris salinasgrandensis</emphasis>
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</taxonomicName>
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Nupponen, sp. nov. (BIN: BOLD:ADY7738, 4.49%).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="remarks">
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<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Female unknown. Based on COI maximum likelihood phylogeny, the South American taxa
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<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris salinasgrandensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="salinasgrandensis">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris salinasgrandensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris furciphallella" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="furciphallella">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris furciphallella</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris manchaoensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="manchaoensis">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris manchaoensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris angustivalvella" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="angustivalvella">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris angustivalvella</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris directiphallella" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="directiphallella">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris directiphallella</emphasis>
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</taxonomicName>
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group together, associating within a clade, whose taxa are classified in apparently non-monophyletic
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<taxonomicName authorityName="Nupponen 1 & Sihvonen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris</emphasis>
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</taxonomicName>
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on BOLD (Suppl. material 2). We classify these taxa in
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<taxonomicName authorityName="Nupponen 1 & Sihvonen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Scythris</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |