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<document id="FF73EC004AD957D80EE7ECD6BA53E9CC" ID-DOI="10.5281/zenodo.293984" ID-GBIF-Dataset="15c86811-b9bf-40de-b89d-3fd161964b2c" ID-ISSN="1175-5326" ID-Zenodo-Dep="293984" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460260703053" checkinUser="plazi" docAuthor="Kondo, Takumasa &amp; Gullan, Penny J." docDate="2010" docId="1F5487B7FFD0FFA429F7CA79B84D8E1C" docLanguage="en" docName="zt02560p015.pdf" docOrigin="Zootaxa 2560" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pulvinaria drimyswinteri Kondo &amp; Gullan, sp. nov." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="E36DFFCFFFD5FFAC2960CD43B944883D" masterDocTitle="The Coccidae (Hemiptera: Coccoidea) of Chile, with descriptions of three new species and transfer of Lecanium resinatum Kieffer &amp; Herbst to the Kerriidae" masterLastPageNumber="15" masterPageNumber="1" pageNumber="6" updateTime="1698244323444" updateUser="plazi">
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<mods:title id="329C5FEFBA84FDFDB9905E3888BA1EE9">The Coccidae (Hemiptera: Coccoidea) of Chile, with descriptions of three new species and transfer of Lecanium resinatum Kieffer &amp; Herbst to the Kerriidae</mods:title>
</mods:titleInfo>
<mods:name id="1BB2F0C43662E250C995CBBED140D3DA" type="personal">
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<mods:namePart id="65AC963900BF470DB215F0EA02EAD2E0">Kondo, Takumasa</mods:namePart>
</mods:name>
<mods:name id="1A91B72AB053BAD8A64E4631D669E12F" type="personal">
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<mods:namePart id="A5D7C84CE6F6A921408B5817C64867E6">Gullan, Penny J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="944636E9460CC7E6CA4710AB7BEBC889">text</mods:typeOfResource>
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<mods:title id="3DC4CDCC159F5FAF091552B18A4B98A5">Zootaxa</mods:title>
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<mods:part id="2EEA854F7BD579060D88912BA7E1B023">
<mods:date id="7F2CA226FA0F3D97830D1DFBD4070361">2010</mods:date>
<mods:detail id="AE08CB50D17FAC962EEFF16EA513BEBC" type="volume">
<mods:number id="67E4623D08AD38C7C7CF464B18F03F5C">2560</mods:number>
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<mods:identifier id="98D98F25E2AA3AF96A18EE73E2B57F13" type="DOI">10.5281/zenodo.293984</mods:identifier>
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<treatment id="1F5487B7FFD0FFA429F7CA79B84D8E1C" ID-DOI="http://doi.org/10.5281/zenodo.5323159" ID-GBIF-Taxon="119393190" ID-Zenodo-Dep="5323159" LSID="urn:lsid:zoobank.org:act:05315AA1-C8EC-4797-B195-73E259684C96" httpUri="http://treatment.plazi.org/id/1F5487B7FFD0FFA429F7CA79B84D8E1C" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection id="DFE7652AFFD0FFA929F7CA79BA878FDC" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="974236A1FFD0FFA929F7CA79BA468F69" blockId="5.[151,770,1850,1914]" box="[151,770,1850,1876]" pageId="5" pageNumber="6">
<heading id="CC0A81CDFFD0FFA929F7CA79BA468F69" bold="true" box="[151,770,1850,1876]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="A589EAB3FFD0FFA929F7CA79BA468F69" bold="true" box="[151,770,1850,1876]" pageId="5" pageNumber="6">
<taxonomicName id="50FD4D22FFD0FFA929F7CA79BBDA8F69" authority="Kondo &amp; Gullan" authorityName="Kondo &amp; Gullan" box="[151,670,1850,1876]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="drimyswinteri" status="sp. nov.">
<emphasis id="A589EAB3FFD0FFA929F7CA79B8818F69" bold="true" box="[151,453,1850,1876]" italics="true" pageId="5" pageNumber="6">Pulvinaria drimyswinteri</emphasis>
Kondo &amp; Gullan
</taxonomicName>
<taxonomicNameLabel id="BEBA57C8FFD0FFA92BC5CA79BA468F69" box="[677,770,1850,1876]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="974236A1FFD0FFA929F7CA23B81A8F47" blockId="5.[151,770,1850,1914]" box="[151,350,1888,1914]" pageId="5" pageNumber="6">
(
<figureCitation id="0FC62A24FFD0FFA929FFCA23B9A78F47" box="[159,227,1888,1914]" captionStart="FIGURE 1" captionStartId="3.[151,255,870,894]" captionTargetBox="[153,1433,181,853]" captionTargetId="figure@3.[151,1436,165,868]" captionTargetPageId="3" captionText="FIGURE 1. Adult female coccids: A, Cryptinglisia chilensis; B, Pulvinaria drimyswinteri, young adults; C, Pulvinaria drimyswinteri, adult with ovisac (centre); D, Stictolecanium cranstoni, young adult; E, Stictolecanium cranstoni, mature adult; F. Pulvinariella mesembryanthemi." httpUri="https://zenodo.org/record/293985/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
B &amp; C, 3).
</paragraph>
<paragraph id="974236A1FFD0FFA929F7CAE9BA878FDC" blockId="5.[151,1436,1962,2017]" pageId="5" pageNumber="6">
<emphasis id="A589EAB3FFD0FFA929F7CAE9B8EB8FFC" box="[151,431,1962,1985]" italics="true" pageId="5" pageNumber="6">Proposed common names</emphasis>
. Spanish: Escama blanda algodonosa del canelo; English: Canelo cottony scale. (The host plant of this scale is known by the common name of “canelo” in
<collectingCountry id="EFEA7631FFD0FFA92A19CA89BAF38FDC" box="[889,951,1994,2017]" name="Chile" pageId="5" pageNumber="6">Chile</collectingCountry>
.)
</paragraph>
</subSubSection>
<subSubSection id="DFE7652AFFD3FFAB29F7CDD4BAF68FEA" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="974236A1FFD3FFAA29F7CDD4B880891F" blockId="6.[151,1438,151,290]" pageId="6" pageNumber="7">
<emphasis id="A589EAB3FFD3FFAA29F7CDD4B8B5888C" bold="true" box="[151,497,151,177]" pageId="6" pageNumber="7">
<typeStatus id="48468803FFD3FFAA29F7CDD4B99F888C" box="[151,219,151,177]" pageId="6" pageNumber="7">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="48468803FFD3FFAA289ACDDBBB33888F" box="[506,631,152,178]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
, adult female.
<collectingCountry id="EFEA7631FFD3FFAA2A5CCDDBBAC3888F" box="[828,903,152,178]" name="Chile" pageId="6" pageNumber="7">Chile</collectingCountry>
, Xth region, Huerquehue National Park, 39°10ˏ00˝N, 75°43ˏ29˝N,
<quantity id="50059B44FFD3FFAA28AECDFEBB5D88EA" box="[462,537,189,215]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="7" unit="m" value="750.0">750 m</quantity>
a.s.l.,
<date id="E3431061FFD3FFAA2B07CDFEBBA488EA" box="[615,736,189,215]" pageId="6" pageNumber="7" value="2006-02-21">21.ii.2006</date>
, coll. Takumasa Kondo, ex
<taxonomicName id="50FD4D22FFD3FFAA2D4ACDFEB99A88C1" authority="J.R. Forst. &amp; G. Forst." authorityName="J.R. Forst. &amp; G. Forst." class="Magnoliopsida" family="Winteraceae" genus="Drimys" kingdom="Plantae" order="Canellales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="winteri">
<emphasis id="A589EAB3FFD3FFAA2D4ACDFEBD9F88EB" box="[1066,1243,189,214]" italics="true" pageId="6" pageNumber="7">Drimys winteri</emphasis>
J.R. Forst. &amp; G. Forst.
</taxonomicName>
, 1(1) (
<collectionCode id="F1ECAE64FFD3FFAA284ACDA1B8C188C1" LSID="urn:lsid:biocol.org:col:34871" box="[298,389,226,252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
).
<typeStatus id="48468803FFD3FFAA28F9CDA1BB4A88C1" box="[409,526,226,252]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
, same data as
<typeStatus id="48468803FFD3FFAA2BD6CDA1BA5B88C1" box="[694,799,226,252]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
, 15 (15) (
<quantity id="50059B44FFD3FFAA2AF1CDA1BAFB88C1" box="[913,959,226,252]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="7" unit="in" value="2.0">2 in</quantity>
BME including voucher No. TK0241,
<quantity id="50059B44FFD3FFAA2CEDCDA1B9EA891F" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="7" unit="in" value="3.0">3 in</quantity>
<collectionCode id="F1ECAE64FFD3FFAA29D5CC4BB854891F" box="[181,272,264,290]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="6" pageNumber="7">MNNC</collectionCode>
,
<quantity id="50059B44FFD3FFAA287CCC4BB813891F" box="[284,343,264,290]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="7" unit="in" value="10.0">10 in</quantity>
<collectionCode id="F1ECAE64FFD3FFAA283ECC4BB8FC891F" LSID="urn:lsid:biocol.org:col:34871" box="[350,440,264,290]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7">USNM</collectionCode>
).
</paragraph>
<caption id="C3826629FFD3FFAA29F7CA8ABAC48FDC" box="[151,896,1993,2017]" httpUri="https://zenodo.org/record/293987/files/figure.png" pageId="6" pageNumber="7" targetBox="[186,1397,397,1992]" targetPageId="6">
<paragraph id="974236A1FFD3FFAA29F7CA8ABAC48FDC" blockId="6.[151,896,1993,2017]" box="[151,896,1993,2017]" pageId="6" pageNumber="7">
<emphasis id="A589EAB3FFD3FFAA29F7CA8AB85D8FDC" bold="true" box="[151,281,1993,2017]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
<taxonomicName id="50FD4D22FFD3FFAA2840CA89BBAD8FDC" authority="Kondo &amp; Gullan" authorityName="Kondo &amp; Gullan" box="[288,745,1994,2017]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="drimyswinteri">
<emphasis id="A589EAB3FFD3FFAA2840CA89BB688FDC" box="[288,556,1994,2017]" italics="true" pageId="6" pageNumber="7">Pulvinaria drimyswinteri</emphasis>
Kondo &amp; Gullan
</taxonomicName>
, adult female.
</paragraph>
</caption>
<paragraph id="974236A1FFD2FFAB29A6CDD4BBA1888F" blockId="7.[151,1438,151,2007]" box="[198,741,151,178]" pageId="7" pageNumber="8">
<emphasis id="A589EAB3FFD2FFAB29A6CDD4B82C888C" bold="true" box="[198,360,151,177]" pageId="7" pageNumber="8">Adult female</emphasis>
(measurements based on n=14).
</paragraph>
<paragraph id="974236A1FFD2FFAB29A6CDFEB8E2898A" blockId="7.[151,1438,151,2007]" pageId="7" pageNumber="8">
<emphasis id="A589EAB3FFD2FFAB29A6CDFEB88B88EA" bold="true" box="[198,463,189,215]" pageId="7" pageNumber="8">Unmounted material</emphasis>
(
<figureCitation id="0FC62A24FFD2FFAB2881CDFEBB6E88EA" box="[481,554,189,215]" captionStart="FIGURE 1" captionStartId="3.[151,255,870,894]" captionTargetBox="[153,1433,181,853]" captionTargetId="figure@3.[151,1436,165,868]" captionTargetPageId="3" captionText="FIGURE 1. Adult female coccids: A, Cryptinglisia chilensis; B, Pulvinaria drimyswinteri, young adults; C, Pulvinaria drimyswinteri, adult with ovisac (centre); D, Stictolecanium cranstoni, young adult; E, Stictolecanium cranstoni, mature adult; F. Pulvinariella mesembryanthemi." httpUri="https://zenodo.org/record/293985/files/figure.png" pageId="7" pageNumber="8">Figs 1</figureCitation>
B &amp; C). Adult female in life about
<quantity id="50059B44FFD2FFAB2A81CDFEBD3588EA" box="[993,1137,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="6.2" metricValueMin="2.5" pageId="7" pageNumber="8" unit="mm" value="4.35" valueMax="6.2" valueMin="2.5">2.56.2 mm</quantity>
long, 2.0
<quantity id="50059B44FFD2FFAB2D94CDFEBC1588EA" box="[1268,1361,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="8" unit="mm" value="4.2">4.2 mm</quantity>
wide, oval, moderately convex, covered by a thin waxy layer. Brown to yellowish brown; some specimens with submarginal areas lighter, yellowish, with mid-dorsal area darker, reddish-brown. Younger specimens especially tessellated, tessellations becoming less clear in older specimens; some older specimens with a clear darker mid-dorsal longitudinal line running from area just anterior to anal plates up to head margin. Immature females reddish brown. Ovisac, in either a straight or curved line, present on mature adult females, up to 4 times length of female.
</paragraph>
<paragraph id="974236A1FFD2FFAB29A6CC81BC2089E1" blockId="7.[151,1438,151,2007]" box="[198,1380,450,476]" pageId="7" pageNumber="8">
<emphasis id="A589EAB3FFD2FFAB29A6CC81B8AA89E1" bold="true" box="[198,494,450,476]" pageId="7" pageNumber="8">Slide-mounted material</emphasis>
(
<figureCitation id="0FC62A24FFD2FFAB289ECC81BB0089E1" box="[510,580,450,476]" captionStart="FIGURE 3" captionStartId="6.[151,255,1993,2017]" captionTargetBox="[186,1397,397,1992]" captionTargetId="figure@6.[151,1436,343,2006]" captionTargetPageId="6" captionText="FIGURE 3. Pulvinaria drimyswinteri Kondo &amp; Gullan, adult female." httpUri="https://zenodo.org/record/293987/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
). Body
<quantity id="50059B44FFD2FFAB2BC2CC81BA7489E1" box="[674,816,450,476]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="6.1" metricValueMin="2.3" pageId="7" pageNumber="8" unit="mm" value="4.199999999999999" valueMax="6.1" valueMin="2.3">2.36.1 mm</quantity>
long,
<quantity id="50059B44FFD2FFAB2A19CC81BD4389E1" box="[889,1031,450,476]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="4.1" metricValueMin="1.7" pageId="7" pageNumber="8" unit="mm" value="2.9" valueMax="4.1" valueMin="1.7">1.74.1 mm</quantity>
wide, elongate-oval in shape.
</paragraph>
<paragraph id="974236A1FFD2FFAB29A5CCA4B9AE8B6F" blockId="7.[151,1438,151,2007]" pageId="7" pageNumber="8">
<emphasis id="A589EAB3FFD2FFAB29A5CCA4B8758A3C" bold="true" box="[197,305,487,513]" pageId="7" pageNumber="8">Dorsum.</emphasis>
Derm membranous. Dorsal setae sharply spinose, rather short, each
<quantity id="50059B44FFD2FFAB2D30CCABBDAF8A3F" box="[1104,1259,488,514]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.25" metricValueMin="0.75" pageId="7" pageNumber="8" unit="mm" value="10.0" valueMax="12.5" valueMin="7.5">7.512.5 mm</quantity>
long, scattered evenly on dorsum. Dorsal microducts not detected. Simple pores each 1.8 μm wide, scarce, scattered over dorsum. Dorsal tubular ducts, dorsal tubercles and pocket-like sclerotizations absent. Preopercular pores small, each 4.06.0 μm wide, circular in shape, present along midline from area just anterior to anal plates and extending anteriorly up to about mesothorax. Anal plates together quadrate, with rounded outer angles, plates located at about 1/5 to 1/4 of body length from posterior margin, each plate 215225 μm long, 105125 μm wide, anterolateral margin 160175 μm long, posterolateral margin 150165 μm long; with about 3 slender setae on dorsal surface along posterior margin of each plate; 1 pair of fringe setae; about 4 ventral subapical setae on each side and 2 pairs of hypopygial setae. Anal ring with 10 setae. Sclerotic area around anal plates absent.
</paragraph>
<paragraph id="974236A1FFD2FFAB29A5CE1EBDF38BDA" blockId="7.[151,1438,151,2007]" pageId="7" pageNumber="8">
<emphasis id="A589EAB3FFD2FFAB29A5CE1EB86A8B4A" bold="true" box="[197,302,861,887]" pageId="7" pageNumber="8">Margin.</emphasis>
Marginal setae sharply spinose, straight to slightly bent, each 1330 μm long, arranged in a single row, with 1021 on each side between anterior and posterior stigmatic areas. Stigmatic clefts very shallow or absent. Stigmatic spine totalling 3 per stigmatic area, median seta longest, each 3265 μm long; lateral setae shorter, each 1043 μm long. Eyes 2225 µm wide, located on dorsal margin.
</paragraph>
<paragraph id="974236A1FFD2FFAB29A5CEB1BAF68FEA" blockId="7.[151,1438,151,2007]" pageId="7" pageNumber="8">
<emphasis id="A589EAB3FFD2FFAB29A5CEB1B8588C31" bold="true" box="[197,284,1010,1036]" pageId="7" pageNumber="8">Venter.</emphasis>
Derm entirely membranous. Perivulvar pores each 810 μm wide, with 58 loculi, abundant on perivulvar region, more or less in single rows across mid areas of all abdominal and thoracic segments, abundant around metathoracic coxae, with a fewer around prothoracic and mesothoracic coxae. Spiracular pores each 6.07.0 μm wide, usually with 5 loculi, an occasional pore with 3, 4 or 6 loculi, present in a narrow band as wide as peritreme or narrower (about 2 or 3 pores wide), with line of pores extending laterally from each spiracle to body margin. Ventral microducts scattered evenly throughout, each about 3.0 μm wide. Ventral tubular ducts of 3
<typeStatus id="48468803FFD2FFAB28BBC991BB648CD1" box="[475,544,1234,1260]" pageId="7" pageNumber="8">types</typeStatus>
: (i) large size tubular ducts present on each side of abdomen in an inner submarginal band from area posterior to vulvar region near anal folds up to area just posterior to metathoracic spiracular pore band, each duct with a broad inner duct and flower-shaped terminal gland; (ii) medium size tubular ducts present in transverse rows across all abdominal segments, and submedially between meso- and metathoracic coxae, and in transverse band in area between prothoracic and mesothoracic coxae, each duct rather similar to large size ducts but smaller and with a narrower inner duct; (iii) smaller tubular ducts present submarginally from area posterior to vulvar region near anal folds up to about abdominal segment II, intermixed with larger size tubular ducts. Ventral tubular ducts of all
<typeStatus id="48468803FFD2FFAB2D4DC89BBD368DCF" box="[1069,1138,1496,1522]" pageId="7" pageNumber="8">types</typeStatus>
absent marginally and submarginally from head and thorax and from the outer submarginal and marginal areas of abdomen. Ventral submarginal setae slender, straight or slightly bent, arranged in a row; other ventral setae present in transverse rows across all abdominal and thoracic segments; about 4 pairs anterior to tentorial bridge; a transverse group present between procoxae and mouthparts and sparsely distributed elsewhere, each seta 7.517.5 μm long, with longest setae (45113 μm long) present on last 3 abdominal segments. Spiracles well developed, anterior spiracular peritremes each 7585 μm wide, posterior peritremes each 90103 μm wide, without a sclerotization around each spiracle. Legs well developed, each coxa 265325 μm long (at widest point), trochanter + femur 400435 μm long; tibia + tarsus 395455 μm long, with tibio-tarsal scleroses; claw 47.5 52.5 μm long, without a denticle. Tarsal digitules both similar, knobbed, one slightly thicker than other; claw digutules similar and broad. Antennae each 410440 μm long, 7 segmented, with fleshy setae present on last 2 antennal segments; with a long slender seta present on antennal segment V. Preantennal pores present, small each 2.5 μm wide. With about 3 pairs of interantennal setae. Mouthparts relatively small; clypeolabral shield 193215 μm wide; labium 1 segmented, with 4 pairs of labial setae.
</paragraph>
</subSubSection>
<subSubSection id="DFE7652AFFDDFFA429A6CDD4BD918DB6" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="974236A1FFDDFFA429A6CDD4BA478A72" blockId="8.[151,1437,151,1569]" pageId="8" pageNumber="9">
<emphasis id="A589EAB3FFDDFFA429A6CDD4B802888C" bold="true" box="[198,326,151,177]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
The adult female of
<taxonomicName id="50FD4D22FFDDFFA42B26CDDBBA4C888C" box="[582,776,152,177]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="drimyswinteri">
<emphasis id="A589EAB3FFDDFFA42B26CDDBBA4C888C" box="[582,776,152,177]" italics="true" pageId="8" pageNumber="9">P. drimyswinteri</emphasis>
</taxonomicName>
can be diagnosed by the combination of the following features: (1) producing a long ovisac, up to 4 times its body length; (2) dorsal microducts, tubular ducts and tubercles absent; (3) dorsal setae sharply spinose, short; (4) antennae well developed, 7 segmented; (5) legs well developed, with tibio-tarsal scleroses; (6) ventral tubular ducts absent from head and body margins plus submargins, but elsewhere 3
<typeStatus id="48468803FFDDFFA42B6DCC6DBB148975" box="[525,592,302,328]" pageId="8" pageNumber="9">types</typeStatus>
present: (i) large size tubular ducts on each side of abdomen in a submarginal band from posterior of vulvar region near anal folds anteriorly to just posterior to metathoracic spiracular pore band, (ii) medium size tubular ducts in transverse rows across all abdominal segments, submedially between meso- and metathoracic coxae, and in transverse band in area between prothoracic and mesothoracic coxae, (iii) smaller tubular ducts present in the inner submarginal areas of abdomen from posterior of vulvar region near anal folds to about abdominal segment II, intermixed with larger size tubular ducts; and (7) perivulvar pores with 58 loculi, present on perivulvar region, across mid areas of all abdominal and thoracic segments, and around coxae.
</paragraph>
<paragraph id="974236A1FFDDFFA429A6CF18BD918DB6" blockId="8.[151,1437,151,1569]" pageId="8" pageNumber="9">
The genus
<taxonomicName id="50FD4D22FFDDFFA42828CF18B8828A49" box="[328,454,603,628]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A589EAB3FFDDFFA42828CF18B8828A49" box="[328,454,603,628]" italics="true" pageId="8" pageNumber="9">Pulvinaria</emphasis>
</taxonomicName>
is composed of more than 100 species (
<bibRefCitation id="F36C4B50FFDDFFA42AC1CF18BD1E8A48" author="Hodgson" box="[929,1114,603,629]" pageId="8" pageNumber="9" refString="Hodgson, C. J. (1994) The Scale Insect family Coccidae: An Identification Manual to Genera. CAB International, Wallingford, Oxon, UK. 639 pp." type="book" year="1994">Hodgson, 1994</bibRefCitation>
; Ben-Dov, 2009), of which 25 have been recorded from the New World (Ben-Dov
<emphasis id="A589EAB3FFDDFFA42A4CCFC2BA258AA7" box="[812,865,641,666]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
., 2009). Reviewing the genus
<taxonomicName id="50FD4D22FFDDFFA42DAFCFC2BC0A8AA7" box="[1231,1358,641,666]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A589EAB3FFDDFFA42DAFCFC2BC0A8AA7" box="[1231,1358,641,666]" italics="true" pageId="8" pageNumber="9">Pulvinaria</emphasis>
</taxonomicName>
of the New World was out of the scope of this paper. Five species in the genus
<taxonomicName id="50FD4D22FFDDFFA42D79CFE5BDD88A82" box="[1049,1180,678,703]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A589EAB3FFDDFFA42D79CFE5BDD88A82" box="[1049,1180,678,703]" italics="true" pageId="8" pageNumber="9">Pulvinaria</emphasis>
</taxonomicName>
are considered to be invasive species in South
<collectingCountry id="EFEA7631FFDDFFA428A5CF8FBB698ADB" box="[453,557,716,742]" name="United States of America" pageId="8" pageNumber="9">America</collectingCountry>
, namely
<taxonomicName id="50FD4D22FFDDFFA42BF6CF8FBABB8ADB" authority="Newstead" authorityName="Newstead" box="[662,1023,716,742]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="A589EAB3FFDDFFA42BF6CF8FBAC58AD8" box="[662,897,716,741]" italics="true" pageId="8" pageNumber="9">Pulvinaria elongata</emphasis>
Newstead
</taxonomicName>
,
<taxonomicName id="50FD4D22FFDDFFA42D6ACF8FBCDC8ADB" authority="Westwood" authorityName="Westwood" box="[1034,1432,716,742]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="floccifera">
<emphasis id="A589EAB3FFDDFFA42D6ACF8FBDBB8AD8" box="[1034,1279,716,741]" italics="true" pageId="8" pageNumber="9">Pulvinaria floccifera</emphasis>
(Westwood)
</taxonomicName>
,
<taxonomicName id="50FD4D22FFDDFFA429F7CFB2B8A78B36" authority="Signoret" authorityName="Signoret" box="[151,483,753,779]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iceryi">
<emphasis id="A589EAB3FFDDFFA429F7CFB2B8248B37" box="[151,352,753,778]" italics="true" pageId="8" pageNumber="9">Pulvinaria iceryi</emphasis>
(Signoret)
</taxonomicName>
,
<taxonomicName id="50FD4D22FFDDFFA42890CFB2BA668B36" authority="Maskell" authorityName="Maskell" box="[496,802,753,779]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="psidii">
<emphasis id="A589EAB3FFDDFFA42890CFB2BBFC8B37" box="[496,696,753,778]" italics="true" pageId="8" pageNumber="9">Pulvinaria psidii</emphasis>
Maskell
</taxonomicName>
, and
<taxonomicName id="50FD4D22FFDDFFA42A02CFB2BDE58B36" authority="Linnaeus" authorityName="Linnaeus" box="[866,1185,753,779]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vitis">
<emphasis id="A589EAB3FFDDFFA42A02CFB2BD5D8B37" box="[866,1049,753,778]" italics="true" pageId="8" pageNumber="9">Pulvinaria vitis</emphasis>
(Linnaeus)
</taxonomicName>
(Kondo unpublished data).
<taxonomicName id="50FD4D22FFDDFFA42981CE54BB488B0D" box="[225,524,791,816]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="drimyswinteri">
<emphasis id="A589EAB3FFDDFFA42981CE54BB488B0D" box="[225,524,791,816]" italics="true" pageId="8" pageNumber="9">Pulvinaria drimyswinteri</emphasis>
</taxonomicName>
does not appear to be an invasive species considering its native host
<taxonomicName id="50FD4D22FFDDFFA42C25CE54B8D28B6B" class="Magnoliopsida" family="Winteraceae" genus="Drimys" kingdom="Plantae" order="Canellales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="winteri">
<emphasis id="A589EAB3FFDDFFA42C25CE54B9AE8B6B" italics="true" pageId="8" pageNumber="9">Drimys winteri</emphasis>
(Winteraceae)
</taxonomicName>
and its geographical location in the Chilean
<taxonomicName id="50FD4D22FFDDFFA42AC0CE7EBD638B6B" box="[928,1063,829,854]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="A589EAB3FFDDFFA42AC0CE7EBD638B6B" box="[928,1063,829,854]" italics="true" pageId="8" pageNumber="9">Nothofagus</emphasis>
</taxonomicName>
forest.
<taxonomicName id="50FD4D22FFDDFFA42D1FCE7EBC2F8B6B" box="[1151,1387,829,854]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="A589EAB3FFDDFFA42D1FCE7EBC2F8B6B" box="[1151,1387,829,854]" italics="true" pageId="8" pageNumber="9">Pulvinaria elongata</emphasis>
</taxonomicName>
and
<taxonomicName id="50FD4D22FFDDFFA429F7CE21B9B28B46" box="[151,246,866,891]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iceryi">
<emphasis id="A589EAB3FFDDFFA429F7CE21B9B28B46" box="[151,246,866,891]" italics="true" pageId="8" pageNumber="9">P. iceryi</emphasis>
</taxonomicName>
occur commonly on grasses (Ben-Dov
<emphasis id="A589EAB3FFDDFFA42BA5CE21BBBC8B46" box="[709,760,866,891]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
., 2009), and the morphology of
<taxonomicName id="50FD4D22FFDDFFA42D13CE21BC778B46" box="[1139,1331,866,892]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="drimyswinteri">
<emphasis id="A589EAB3FFDDFFA42D13CE21BDC68B46" box="[1139,1154,866,891]" italics="true" pageId="8" pageNumber="9">P</emphasis>
.
<emphasis id="A589EAB3FFDDFFA42DF0CE21BC778B46" box="[1168,1331,866,891]" italics="true" pageId="8" pageNumber="9">drimyswinteri</emphasis>
</taxonomicName>
does not match the morphology of any of the other three species considered to be invasive (character states of
<taxonomicName id="50FD4D22FFDDFFA42CE5CECBB87E8BFA" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="drimyswinteri">
<emphasis id="A589EAB3FFDDFFA42CE5CECBBCD08B9C" box="[1413,1428,904,929]" italics="true" pageId="8" pageNumber="9">P</emphasis>
.
<emphasis id="A589EAB3FFDDFFA429F7CEEDB87E8BFA" box="[151,314,942,967]" italics="true" pageId="8" pageNumber="9">drimyswinteri</emphasis>
</taxonomicName>
in parenthesis), i.e.,
<taxonomicName id="50FD4D22FFDDFFA42B50CEEDBBCA8BFA" box="[560,654,942,967]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="psidii">
<emphasis id="A589EAB3FFDDFFA42B50CEEDBBCA8BFA" box="[560,654,942,967]" italics="true" pageId="8" pageNumber="9">P. psidii</emphasis>
</taxonomicName>
has fringed marginal setae (sharply spinose, with a pointed apex);
<taxonomicName id="50FD4D22FFDDFFA429F7CE90B8628BD1" box="[151,294,979,1004]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="floccifera">
<emphasis id="A589EAB3FFDDFFA429F7CE90B8628BD1" box="[151,294,979,1004]" italics="true" pageId="8" pageNumber="9">P. floccifera</emphasis>
</taxonomicName>
and
<emphasis id="A589EAB3FFDDFFA42801CE90B8E98BD1" box="[353,429,979,1004]" italics="true" pageId="8" pageNumber="9">P. v i t i s</emphasis>
have dorsal submarginal tubercles (absent) [character states taken from
<bibRefCitation id="F36C4B50FFDDFFA42C6FCE90BCD38BD0" author="Gill" box="[1295,1431,979,1005]" pageId="8" pageNumber="9" refString="Gill, R. J. (1988) The Scale Insects of California: Part 1. The Soft Scales (Homoptera: Coccoidea: Coccidae). California Dept. of Food &amp; Agriculture, Sacramento, California. 132 pp." type="book" year="1988">Gill (1988)</bibRefCitation>
, Hodgson (2002) and
<bibRefCitation id="F36C4B50FFDDFFA428F0CEBABBF88C2E" author="Tanaka" box="[400,700,1017,1043]" pageId="8" pageNumber="9" refString="Tanaka, H. &amp; Amano, H. (2007) Redescription of Pulvinaria floccifera (Insecta: Hemiptera: Coccidae). Species Diversity, 12, 211 - 215." type="journal article" year="2007">Tanaka &amp; Amano (2007)</bibRefCitation>
]. No other
<taxonomicName id="50FD4D22FFDDFFA42A5ECEBABAFF8C2F" box="[830,955,1017,1042]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A589EAB3FFDDFFA42A5ECEBABAFF8C2F" box="[830,955,1017,1042]" italics="true" pageId="8" pageNumber="9">Pulvinaria</emphasis>
</taxonomicName>
species have been described from
<collectingCountry id="EFEA7631FFDDFFA42C35CEBABCDD8C2E" box="[1365,1433,1017,1043]" name="Chile" pageId="8" pageNumber="9">Chile</collectingCountry>
. There are five species described from neighboring
<collectingCountry id="EFEA7631FFDDFFA42B8BC95DBA248C05" box="[747,864,1054,1080]" name="Argentina" pageId="8" pageNumber="9">Argentina</collectingCountry>
and none recorded from neighboring
<collectingCountry id="EFEA7631FFDDFFA42C7CC95DBC168C05" box="[1308,1362,1054,1080]" name="Peru" pageId="8" pageNumber="9">Peru</collectingCountry>
(Ben- Dov
<emphasis id="A589EAB3FFDDFFA429B2C907B8438C60" box="[210,263,1092,1117]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
., 2009). In Granara de Willink´s (1999) key to species recorded from
<collectingCountry id="EFEA7631FFDDFFA42D32C907BD8F8C63" box="[1106,1227,1092,1118]" name="Argentina" pageId="8" pageNumber="9">Argentina</collectingCountry>
,
<taxonomicName id="50FD4D22FFDDFFA42DB9C907BCD88C60" box="[1241,1436,1092,1117]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="drimyswinteri">
<emphasis id="A589EAB3FFDDFFA42DB9C907BCD88C60" box="[1241,1436,1092,1117]" italics="true" pageId="8" pageNumber="9">P. drimyswinteri</emphasis>
</taxonomicName>
comes closest to
<taxonomicName id="50FD4D22FFDDFFA42808C92ABB238CBE" authority="Brethes" authorityName="Brethes" box="[360,615,1129,1155]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="A589EAB3FFDDFFA42808C92ABB478CBF" box="[360,515,1129,1154]" italics="true" pageId="8" pageNumber="9">P. flavescens</emphasis>
Brethes
</taxonomicName>
and
<taxonomicName id="50FD4D22FFDDFFA42BC4C92ABAF38CBE" authority="Brethes. It" authorityName="Brethes. It" box="[676,951,1129,1155]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="platensis">
<emphasis id="A589EAB3FFDDFFA42BC4C92ABA6B8CBF" box="[676,815,1129,1154]" italics="true" pageId="8" pageNumber="9">P. platensis</emphasis>
Brethes. It
</taxonomicName>
differs from
<taxonomicName id="50FD4D22FFDDFFA42D3AC92ABDB28CBF" box="[1114,1270,1129,1154]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="A589EAB3FFDDFFA42D3AC92ABDB28CBF" box="[1114,1270,1129,1154]" italics="true" pageId="8" pageNumber="9">P. flavescens</emphasis>
</taxonomicName>
by having an elongate, oval-shaped body and abundant ventral tubular ducts across mid areas of the abdominal segments (body pyriform shaped and ventral tubular ducts absent across mid areas of abdominal segments in
<taxonomicName id="50FD4D22FFDDFFA42CE5C9F6B84B8CCE" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="A589EAB3FFDDFFA42CE5C9F6B84B8CCE" italics="true" pageId="8" pageNumber="9">P. flavescens</emphasis>
</taxonomicName>
), and from
<emphasis id="A589EAB3FFDDFFA428F8C999BB5D8CCE" box="[408,537,1242,1267]" italics="true" pageId="8" pageNumber="9">P. p l a t e n s i s</emphasis>
by the lack of dorsal tubular ducts, in having straight to slightly bent sharply spinose marginal setae, and by having smaller tubular ducts present submarginally from area posterior to vulvar region near anal folds anteriorly to about abdominal segment II (dorsal tubular ducts present, marginal setae curved and with a round apex, and with some smaller tubular ducts present on submarginal areas of thorax and abdomen in
<emphasis id="A589EAB3FFDDFFA428CCC832BB698DB7" box="[428,557,1393,1418]" italics="true" pageId="8" pageNumber="9">P. p l a t e n s i s</emphasis>
) (character states taken from Granara de Willink (1999)).
</paragraph>
</subSubSection>
<subSubSection id="DFE7652AFFDDFFA429A6C8D5B85D8DEB" pageId="8" pageNumber="9" type="etymology">
<paragraph id="974236A1FFDDFFA429A6C8D5B85D8DEB" blockId="8.[151,1437,151,1569]" pageId="8" pageNumber="9">
<emphasis id="A589EAB3FFDDFFA429A6C8D5B8178D8D" bold="true" box="[198,339,1430,1456]" pageId="8" pageNumber="9">Etymology.</emphasis>
The species is named after its host:
<taxonomicName id="50FD4D22FFDDFFA42A64C8D5BD268D8D" box="[772,1122,1430,1456]" class="Magnoliopsida" family="Winteraceae" genus="Drimys" kingdom="Plantae" order="Canellales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="winteri">
<emphasis id="A589EAB3FFDDFFA42A64C8D5BAF18D92" box="[772,949,1430,1455]" italics="true" pageId="8" pageNumber="9">Drimys winteri</emphasis>
(Winteraceae)
</taxonomicName>
and the name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DFE7652AFFDDFFA429A6C8A2B84D8E1C" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="974236A1FFDDFFA429A6C8A2B84D8E1C" blockId="8.[151,1437,151,1569]" pageId="8" pageNumber="9">
<emphasis id="A589EAB3FFDDFFA429A6C8A2B86E8DC6" bold="true" box="[198,298,1505,1531]" pageId="8" pageNumber="9">Biology.</emphasis>
Adult females were collected on the leaves of the host, locally known as canelo. No males were observed.
</paragraph>
</subSubSection>
</treatment>
</document>