505 lines
67 KiB
XML
505 lines
67 KiB
XML
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<mods:title id="25DE40F2FCA00571A07C3B7D80438CF3">Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts</mods:title>
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<mods:name id="DE4031D984F14CAB5F0C5D887C4130EF" type="personal">
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<mods:namePart id="4C337029B6A83A6B09DAAD9AC49F2602">Kato, Makoto</mods:namePart>
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<mods:nameIdentifier id="A2DA1482F063E538F118A2DF4E15147F" type="email">kato@zoo.zool.kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="A0A3DE852BA1CA5351D03BD60DB45C7C">Yamamori, Luna</mods:namePart>
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<mods:nameIdentifier id="E1FFC7BD51D8014CB9DA4A19A6040572" type="ORCID">https://orcid.org/0000-0002-5342-1277</mods:nameIdentifier>
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<mods:namePart id="2F9A7101F84568F421F0C8B8418910F4">Imada, Yume</mods:namePart>
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<mods:nameIdentifier id="8928AA9BDC98766F2F03C4C619EFE2A8" type="ORCID">https://orcid.org/0000-0003-2173-7389</mods:nameIdentifier>
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<mods:title id="C26F040A5E270BBAC6E9801E856F30D5">ZooKeys</mods:title>
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<mods:date id="7FEBEC1A831C561F3F17D5FF2959D83D">2022</mods:date>
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<mods:number id="6422B784CD7AC221516FE2742B8B8B39">2022-11-30</mods:number>
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<mods:number id="4BE2EA4CB969A8EEB6E39E4F0961DCDA">1133</mods:number>
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<mods:url id="CBF1C3C66556468EE72424EF67EB20C0">http://dx.doi.org/10.3897/zookeys.1133.94530</mods:url>
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<mods:identifier id="4963DE15C5016B53CFB68AD504C8A9B3" type="DOI">http://dx.doi.org/10.3897/zookeys.1133.94530</mods:identifier>
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<treatment id="190EAA651A7051A2B2CEABEBB1717315" ID-GBIF-Taxon="205344583" LSID="urn:lsid:zoobank.org:act:FD0892D0-011B-4A1A-A1FE-F1ECC92FF89E" httpUri="http://treatment.plazi.org/id/190EAA651A7051A2B2CEABEBB1717315" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="CBB58E35301C88DE73938A71F8DFAF92" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="1D4373E7606148D184E822E4F831D579" pageId="0" pageNumber="1">
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4.
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<taxonomicName id="1FB19A9D433623B8616A4F597EC37DAA" ID-CoL="BDJLT" LSID="https://zoobank.org/FD0892D0-011B-4A1A-A1FE-F1ECC92FF89E" authority="Kato" authorityName="Kato" authorityYear="2022" class="Insecta" family="Agromyzidae" genus="Phytoliriomyza" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Phytoliriomyza marchantiae" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marchantiae" status="sp. nov.">Phytoliriomyza marchantiae Kato</taxonomicName>
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<taxonomicNameLabel id="85C1977FD827578474EBDD3F650DEAA1" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5359AEE8595E8FD2B79EC91DFD7D1AD2" pageId="0" pageNumber="1" type="description">
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<paragraph id="4D14B8B6DAD16478DE9241DFF8E44617" pageId="0" pageNumber="1">
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<figureCitation id="E8F89B38CEFE264857D78F14EF33CC5D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">Figs 6</figureCitation>
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<figureCitation id="7B06FDE77D3079A70C8F46F5659636BA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B7CC3092D97266B7E4C23F18D33E7AC4" pageId="0" pageNumber="1" type="material examined">
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<paragraph id="DD2FB8C3C82B8D6452C89CE2C42233C5" pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph id="AD336908D35B302826F9638154448E38" pageId="0" pageNumber="1">
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<emphasis id="E3829054DAFB3229E3460128EC310CCA" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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<emphasis id="45A8111FB68DB6B055768DBC73369DD8" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Japan: 1♂ (MK-AG-a320), Namari-kawa, Yakumo, Futami, Hokkaido (
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<geoCoordinate id="BDF29F878E1465227C1737AF01821886" degrees="42.187765" direction="north" orientation="latitude" precision="1" value="42.187763">42.187765°N</geoCoordinate>
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,
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<geoCoordinate id="88B68070EDF76AA402774C13124FE8C6" degrees="140.122182" direction="east" orientation="longitude" precision="1" value="140.12218">140.122182°E</geoCoordinate>
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, 190 m asl), 2-VI-2021 (as larva on
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<taxonomicName id="53B25C69FA43060B399E57B74B62D949" authority="paleacea" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="paleacea">
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<emphasis id="359742EAE7B192DDB177BA8F3369D5A5" italics="true" pageId="0" pageNumber="1">Marchantia paleacea paleacea</emphasis>
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</taxonomicName>
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), emerged on 17-VI-2021, NSMT-I-Dip 31903.
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<emphasis id="668C81BC0FD6601CDD9F43740C2B8288" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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<emphasis id="A24245E541E7792FA98EB84FF5F7B355" bold="true" pageId="0" pageNumber="1">:</emphasis>
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Japan: 1♂2♀ (MK-AG-a484, a485, a442), same data as holotype, emerged on 18-VI-3-VII-2021, NSMT-I-Dip 31904-31906; 1♂ (MK-AG-a322), Renge-onsen, Itoigawa, Niigata Pref., 11-VII-2021 (as larva on
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<taxonomicName id="755F42EB0547338C9716AC5C5393B6E5" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. p. subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="paleacea">
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<emphasis id="D8EAC41F3B8DC8BE6190F144A7099BC5" italics="true" pageId="0" pageNumber="1">M. p. paleacea</emphasis>
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</taxonomicName>
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), emerged on 5-VIII-2021, NSMT-I-Dip 31907; 1♂ (MK-AG-a321), Nippara, Okutama, Tokyo Pref., 27-III-2021 (as larva on
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<taxonomicName id="34AC369A8D9344BC109D2594ABD0DC71" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. p. subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="paleacea">
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<emphasis id="29A2456896F42E643D78F3522511092B" italics="true" pageId="0" pageNumber="1">M. p. paleacea</emphasis>
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</taxonomicName>
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), emerged on 27-V-2021, NSMT-I-Dip 31908; 1♀ (MK-AG-302), Umegashima, Aoi-ku, Shizuoka Pref., 5-I-2014 (as larva on
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<taxonomicName id="06EBA999CA3B001E66A6227F20924AAD" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. p. subsp. diptera" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="diptera">
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<emphasis id="93E0B753A5F6C2BD775930E72191753C" italics="true" pageId="0" pageNumber="1">M. p. diptera</emphasis>
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</taxonomicName>
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), emerged on?-V-2014, NSMT-I-Dip 31909; 1♂ (MK-AG-a356), Inago, Shibakawa, Fujinomiya, Shizuoka Pref., 20-III-2000 (as larva on
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<taxonomicName id="609DF7A2AAC3066BBBC420D84E2F0F83" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. p. subsp. diptera" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="diptera">
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<emphasis id="F93EF759AC20910E06C253F805CCE387" italics="true" pageId="0" pageNumber="1">M. p. diptera</emphasis>
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</taxonomicName>
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), emerged on 17-IV-2000, NSMT-I-Dip 31910; 1♂ (MK-AG-a355), Muramatsu, Iwakura, Sakyo-ku, Kyoto Pref., 11-VI-2021 (as larva on
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<taxonomicName id="236360D3B0DF1632BD0B9A76D964658A" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. polymorpha" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="polymorpha">
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<emphasis id="F6B7868AF49E66B4C2A7D63B92300BFD" italics="true" pageId="0" pageNumber="1">M. polymorpha</emphasis>
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</taxonomicName>
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), emerged on 21-VII-2021, NSMT-I-Dip 31911; 1♂ (MK-AG-a358), Naiku, Oe, Fukuchiyama, Kyoto Pref., 19-V-2021 (as larva on
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<taxonomicName id="F245F4F59B0F8AB4CABBD7BFCE7CA9C4" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. papillata subsp. grossibarba" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="papillata" subSpecies="grossibarba">
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<emphasis id="45A3B2D1E1F82E505025662FC93E74AE" italics="true" pageId="0" pageNumber="1">M. papillata grossibarba</emphasis>
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</taxonomicName>
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), emerged on 3-VII-2021, NSMT-I-Dip 31912.
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</paragraph>
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</subSubSection>
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<subSubSection id="D0470B156716D01977595E9813CA9BE9" pageId="0" pageNumber="1" type="other material">
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<paragraph id="FB54518E66A3F5FA43935FDFC653313C" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="4657E9BB3F4C9CF0F955D90D31D4D2A6" pageId="0" pageNumber="1">
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Japan: On
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<taxonomicName id="DACA6D4D05FC946C66E243C8593EF89B" authority="paleacea" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="paleacea">
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<emphasis id="0B31B07454BEFB91A2C4C40FBBA8C0A4" italics="true" pageId="0" pageNumber="1">Marchantia paleacea paleacea</emphasis>
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</taxonomicName>
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: 22♂16♀, Namari-kawa, Yakumo, Futami, Hokkaido, 2-VI-2021 (as larva), emerged on 9-VI-7-VII-2021; 38♂44♀, Renge-onsen, Itoigawa, Niigata Pref., I-VII-2021 (as larva), emerged on 29-VII-9-VIII-2021; 2♂1♀, Narahara, Ueno, Tano, Gunnma Pref., 18-IV-2021 (as larva), emerged on 30-31-V-2021; 2♂4♀, Kanna-gawa, Nakatsugawa, Chichibu, Saitama Pref., 14-XI-2010 (as larva), emerged on 26-IV-2010; 2♂1♀, Nippara, Okutama, Tokyo Pref., 27-III-2021 (as larva), emerged on 27-IV-21-V-2021; 5♂5♀, Yashajin-toge, Minami-arupusu, Yamanashi Pref., 25-III-2021 (as larva), emerged on 2-V-1-VI-2021.
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</paragraph>
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<paragraph id="83DFC9CE70F9741CE64FCC8FEF24AA7D" pageId="0" pageNumber="1">
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On
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<taxonomicName id="CB70052859D9E3B5B3D67E4F78CA385C" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. diptera" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="diptera">
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<emphasis id="E26F027829FFB99F0DB19061B2FB36C0" italics="true" pageId="0" pageNumber="1">Marchantia paleacea diptera</emphasis>
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</taxonomicName>
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: 2♂3♀, Inago, Shibakawa, Fujinomiya, Shizuoka Pref., 26-V-2021 (as larva), emerged on 17-27-VI-2021; 1♂5♀, Abe-toge, Aoi-ku, Shizuoka Pref., 30-XI-2014 (as larva), emerged on 30-IV-8-V-2014; 3♂5♀, Kuchisakamoto, Aoi-ku, Shizuoka Pref., 26-V-2021 (as larva), emerged on 1-26-VI-2021; 6♂8♀, Tsudono, Aoi-ku, Shizuoka Pref., 30-XI-2014 (as larva), emerged on 30-IV-6-V-2014; 2♂4♀, Tosayama, Kochi, Kochi Pref., 27-II-2011 (as larva), emerged on 23-28-IV-2011; 1♂3♀, Mt. Nabejiri, Taga, Shiga Pref., 4-V-2021 (as larva), emerged on 27-V-15-VI-2021; 6♂3♀, Naiku, Oe, Fukuchiyama, Kyoto Pref., 19-V-2021 (as larva), emerged on 2-10-VII-2021; 2♂3♀, Iya-kei, Ikeda, Miyoshi, Tokushima Pref., 1-II-2014 (as larva), emerged on 4-4-V-2014; 2♂5♀, Okujisso, Isa, Kagoshima Pref., 17-XII-2017 (as larva), emerged on 22-30-III-2012.
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</paragraph>
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<paragraph id="9935BEEDE4E9317F113499A31FE5CE68" pageId="0" pageNumber="1">
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On
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<taxonomicName id="31FBD945A0F7CD9013FEC5765B08E578" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia polymorpha" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="polymorpha">
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<emphasis id="4DAC49D9C916D55258DC58A68820886E" italics="true" pageId="0" pageNumber="1">Marchantia polymorpha</emphasis>
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</taxonomicName>
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: 1♀, Matsubara-ko, Koumi, Nagano Pref., 18-IV-2021 (as larva), emerged on 18-VI-2021; 7♂2♀, Odarumi, Makioka, Yamanashi Pref., 30-VI-2021 (as larva), emerged on 28-VII-4-VII-2021; 5♂15♀, Muramatsu, Iwakura, Sakyo-ku, Kyoto Pref., 31-XII-2013 (as larva), emerged on?-V-2013.
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</paragraph>
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<paragraph id="134AEA9F711F1EB3E4FF698AA50541BA" pageId="0" pageNumber="1">
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On
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<taxonomicName id="925C0C0835609ADF24A5F6C672CC6A42" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia papillata subsp. grossibarba" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="papillata" subSpecies="grossibarba">
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<emphasis id="04F2E02DE2C64519682405C1B28CE3F5" italics="true" pageId="0" pageNumber="1">Marchantia papillata grossibarba</emphasis>
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</taxonomicName>
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: 2♂1♀, Inago, Shibakawa, Fujinomiya, Shizuoka Pref., 26-V-2021 (as larva), emerged on 18-VII-2021; 3♂5♀, Naiku, Oe, Fukuchiyama, Kyoto Pref., 19-V-2021 (as larva), emerged on 2-10-VII-2021; 2♂1♀, Seikandoro, Kumanogawa, Shingu, Wakayama Pref., 7-VII-2021 (as larva), emerged on 27-VII-4-VIII-2021; 1♀, Nagabuchi, Ume, Saeki, Oita Pref., 29-XI-2011 (as larva), emerged on 3-V-2011.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78B1760CF6D92D37576428A8F514FBC2" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="73A99B1D72FAE7BE0C22CF0FABCEACAD" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="D141F409776E0D7FDC4ABFBF17FA2258" pageId="0" pageNumber="1">
|
||
A medium-sized species (wing length 1.6-1.8 mm) having dark gray scutum, dark-cornered yellow scutellum, black 1st flagellomere, black maxillary palpus, gray halteres, and brown legs. Male epandrium with a comb comprising seven or eight fused long tubercle-like setae. Larva mines the thallus of
|
||
<taxonomicName id="767D7D6640BA70B414EA431A3A1C9F7D" authorityName="C.Linnaeus" authorityYear="1753" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="23454B203714F2D00517E35350C235B7" italics="true" pageId="0" pageNumber="1">Marchantia</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC9DB051F5E0B9E1CDCDC343DB3A08A7" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="9EE3980A52E47EF2B5CB506CBF7A5BA5" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="265F96DD2568B6DFAD8224320BD1ACDF" pageId="0" pageNumber="1">
|
||
<emphasis id="CBF297803716369BFFB95E46374E1573" bold="true" pageId="0" pageNumber="1">Adult male</emphasis>
|
||
(Fig.
|
||
<figureCitation id="5D1ECE2E8D371BFB7FB508303CCD913D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6A-E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="331036EB5022F897A5B66BB7CF3B1C26" pageId="0" pageNumber="1">
|
||
<emphasis id="69C8FFD97D706572BA606D35A97BCD90" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
<emphasis id="3BA0C698FF4C5FDDEA08A5CD509E9A24" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
Head light yellow, with ocellar tubercle brown, frons light yellow, back of head dark brown (Fig.
|
||
<figureCitation id="40F4432E2F5E2319A5D28B530FA8FA38" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6C</figureCitation>
|
||
). Antenna porrect, first flagellomere black, pedicel and scape brown. Arista subbasal, black, pubescent. Clypeus, face, gena, parafacial and postgena yellow. Proboscis normal, yellow; palpus brown, cylindrical.
|
||
<emphasis id="49D65734A4EDD87AF2199684B1CB9FB0" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
|
||
<emphasis id="1CF786C1271B365D4E4A60D5C3E65A65" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
Front orbitals three pairs; one ori directed inward; two ors directed upward (Fig.
|
||
<figureCitation id="8BD0AE799DD254256A21E81BE2D657F0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6B</figureCitation>
|
||
). Orbital setulae minute and erect, in a single row.
|
||
</paragraph>
|
||
<caption id="2DD76830DE2EB114401A57AAD14C9A12" doi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
|
||
<paragraph id="4D2C070E35F8FD7EC21EE497D9E4A11F" pageId="0" pageNumber="1">
|
||
<emphasis id="D24FF26921AF7CB82376C0F36AB5A1C4" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
|
||
<taxonomicName id="C2D0FF9D400D279C2D36BDE3F166E3B9" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza marchantiae" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marchantiae">
|
||
<emphasis id="AA94FB3A9188ADA23AE22F5B72202C75" italics="true" pageId="0" pageNumber="1">Phytoliriomyza marchantiae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="4E80A83C4287F092CACFD57DCFE9E30A" bold="true" pageId="0" pageNumber="1">A-E</emphasis>
|
||
holotype male
|
||
<emphasis id="820143017C769CAC6EF8E1D8B3750A9D" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
habitus
|
||
<emphasis id="70F5812EB8AC501F3ACEDCAB18BE2A8E" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
lateral
|
||
<emphasis id="E05EB0F0EE2968C8B2BDC54D11943E36" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
frontal
|
||
<emphasis id="DEB7CD3B00FD68F0B254A9BFC66DE30A" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
dorsal
|
||
<emphasis id="9F13C214FFB75B63E44500C8E4C6106B" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
posterior
|
||
<emphasis id="40B01E9B83CD19A969BDAB5892655177" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
paratype female (MK-AG-a442), dorsal
|
||
<emphasis id="32341EF80AAF29B1F7F6D66E2BD0EA5C" bold="true" pageId="0" pageNumber="1">G-I</emphasis>
|
||
male genitalia
|
||
<emphasis id="E8DF6871686872990FCC2AE5D38340EF" bold="true" pageId="0" pageNumber="1">G-I</emphasis>
|
||
emerged from
|
||
<taxonomicName id="588A55EF48CB793B55B5B0DA3B641F72" authority="paleacea" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="paleacea">
|
||
<emphasis id="95480A3EE7F79A76B8C034E23EA8568A" italics="true" pageId="0" pageNumber="1">Marchantia paleacea paleacea</emphasis>
|
||
</taxonomicName>
|
||
at type locality
|
||
<emphasis id="43FB86C8267BD555A2517B370090DD35" bold="true" pageId="0" pageNumber="1">J-M</emphasis>
|
||
emerged from
|
||
<taxonomicName id="D8AD235BD6A044D807FADFA9CCA80EE2" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. papillata subsp. grossibarba" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="papillata" subSpecies="grossibarba">
|
||
<emphasis id="06C2BDC2966691D40DC2C5E6A27CE0D9" italics="true" pageId="0" pageNumber="1">M. papillata grossibarba</emphasis>
|
||
</taxonomicName>
|
||
at Mt. Oe.
|
||
<emphasis id="358728286F741E6D251106A686A18081" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
whole genitalia, ventral
|
||
<emphasis id="A22F69D0B7666E910A31118A859FBAB7" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||
phallus complex, lateral
|
||
<emphasis id="28F22E95C26950C21F6E94FDE9DDAA7A" bold="true" pageId="0" pageNumber="1">I, L</emphasis>
|
||
epandrium, ventral
|
||
<emphasis id="84222DC6C4C2ECBF22A2A71A69A16170" bold="true" pageId="0" pageNumber="1">J, K</emphasis>
|
||
phallic complex, ventral and lateral
|
||
<emphasis id="AEEBDF84C69CE40E004FB1A92595D42A" bold="true" pageId="0" pageNumber="1">M</emphasis>
|
||
ejaculatory apodeme, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="28FB131D52530BA406CDA624D53EE57B" pageId="0" pageNumber="1">
|
||
<emphasis id="D2302C0EF0776EDC24E5B86707A8A579" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
|
||
<emphasis id="4B0586BAA69FF52AC818267B91517A01" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
Thorax pruinose. Scutum pruinose gray, with a dark gray medial stripe on anterior 2/3, one pair of dark gray lateral stripes, and with narrow yellow patch along posterior margin (Fig.
|
||
<figureCitation id="B0005E8180047A28EFF5159E715C491C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6D</figureCitation>
|
||
). Scutellum light yellow with lateral corners widely brownish. Subscutellum brown with anterior margin light yellow. Mediotergite, anatergite and katatergite dark brown. Pleuron largely yellow; postpronotal lobe with anterior brown spot; notopleuron with brown narrow spot along anterior lower margin; anepisternum and anepimeron with brown venters; katepisternum and meron with dark brown patches on venter (Fig.
|
||
<figureCitation id="81A8D53DBF2AAAD2875D9C108E84833D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6B</figureCitation>
|
||
). Haltere yellowish gray, with yellow stalk. Calypter margin and hairs gray. Leg segments brown; tibia and tarsus darker (Fig.
|
||
<figureCitation id="26E67DF532A836EC0DC1E416AE334154" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6A</figureCitation>
|
||
).
|
||
<emphasis id="531B9802F94D6B5CFB172B95FCFD1254" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
|
||
<emphasis id="D5394075787808376F8E801E98043A98" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
Scutum with 1+3 dorsocentrals, shortened anteriorly (Fig.
|
||
<figureCitation id="6144DA1B3BBD80937D7FD4D9C2CEAB5E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6D</figureCitation>
|
||
). Acrostichal setulae five or six pairs largely in two rows.
|
||
<emphasis id="972AEF1285EDCE4A0B7CA3D21DE73D9A" bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
|
||
: Wing length 1.6-1.7 mm, costa reaching M1 (Fig.
|
||
<figureCitation id="1B2312341DC3E3FBF12D6F50242471E9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6A</figureCitation>
|
||
). Length of ultimate section of vein M4 divided by penultimate section 1.6-1.8.
|
||
</paragraph>
|
||
<paragraph id="1B7ECC44199B966EF277FDACA03556B5" pageId="0" pageNumber="1">
|
||
<emphasis id="867F9EB00C6660882733EB248F4B9576" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
||
<emphasis id="C527DB2FA1833DD055AC2C360393BC99" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
Abdomen dorsally subshiny brown (Fig.
|
||
<figureCitation id="F21527D3B55809F2E77BFDED0895F520" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6E</figureCitation>
|
||
).
|
||
<emphasis id="9D52F7473D6B857AA5207443E3214EBB" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
|
||
<emphasis id="7099E7673DED5A864BF09F5BAF02C1A3" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
(Fig.
|
||
<figureCitation id="D96FDFE77CF91F354F8A89B033F74100" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6G-M</figureCitation>
|
||
) Epandrium dark brown, rounded apically, inner-lateral surface medially with one tubercle-like setae; inner-anterior surface ventrally with a comb comprising eight fused long tubercle-like setae; ventral margin with a row of several minute tubercle-like setae (Fig.
|
||
<figureCitation id="A75D3E740FA0C0379758275EDFE1046A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6I, L</figureCitation>
|
||
). Surstylus subrectangular, setose apically. Cercus narrow, setose. Subepandrial sclerite with one pair of spines and one pair of plate-like ventral processes; tips hook-like (Fig.
|
||
<figureCitation id="FF384402D16828969BE3AC8DDF9027A7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6I</figureCitation>
|
||
). Hypandrium slightly sclerotized along outer margin. Postgonite bare and goose barnacle-shaped (Fig.
|
||
<figureCitation id="0381FE3B68A13F717D0C46818B68CB11" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H</figureCitation>
|
||
). Phallophorus with shallow incision below (Fig.
|
||
<figureCitation id="0432F7A56262929247C6CD2E93C73B10" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6J</figureCitation>
|
||
), articulated with phallapodeme, fused to epiphallus (Fig.
|
||
<figureCitation id="577A979BF1C9CB6A025D01D26B552BB9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H, K</figureCitation>
|
||
). Basiphallus with subtriangular dorsal sclerite; 2/3
|
||
<normalizedToken id="9B0E1F47155068AEE9F0A8081260B160" originalValue="×">x</normalizedToken>
|
||
as long as mesophallus (Fig.
|
||
<figureCitation id="2C08DF8E65B5D368FF920F3CCB18198A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H, K</figureCitation>
|
||
). Hypophallus subtriangular membranous, lined with numerous microtrichia ventrally, medially with a pair of fused linear pale sclerites (Fig.
|
||
<figureCitation id="5EB1937AE35286CB05FD24CA1B4D3E83" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H, K</figureCitation>
|
||
). Paraphalli consisting of a pair of incurved narrow pigmented sclerites and a pair of triangular plate-like pigmented sclerites (Fig.
|
||
<figureCitation id="FE9B0392973AD7494E6F713B235B7124" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H, K</figureCitation>
|
||
). Mesophallus dark, cylindrical, widest subbasally, slightly longer than distiphallus (Fig.
|
||
<figureCitation id="B43A9D7959F44A32395476D6A259B111" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H</figureCitation>
|
||
). Distiphallus comprising one pair of stout tubules; basal half composed of ventral dark scalpel-shaped sclerite and weaker medial region; distal half cylindrical, dorsally and laterally pigmented, with truncated, unpigmented apex (Fig.
|
||
<figureCitation id="B0C2239BB816CFF3E048AE8459031B59" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6H, K</figureCitation>
|
||
). Ejaculatory apodeme fan-shaped, blade pale with apical half clear; sclerite of sperm pump with lateral extension; sperm pump clear (Fig.
|
||
<figureCitation id="24E0E97369710D155662B3EF9D1A7EAC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7EDEBBDE85DA4F227EE0C0B2C585CDF0" pageId="0" pageNumber="1">
|
||
<emphasis id="D9812CB6319D920ADF4111309DB03E10" bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||
(Fig.
|
||
<figureCitation id="1644D1600EA42CEFB6F8E9A29600F1F1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6I-K</figureCitation>
|
||
). Similar to male, but scutellum darker than male (Fig.
|
||
<figureCitation id="16C4A40A7AD3F778F4345671965AB5D9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phytoliriomyza marchantiae sp. nov. A-E holotype male A habitus B lateral C frontal D dorsal E posterior F paratype female (MK-AG-a 442), dorsal G-I male genitalia G-I emerged from Marchantia paleacea paleacea at type locality J-M emerged from M. papillata grossibarba at Mt. Oe. G whole genitalia, ventral H phallus complex, lateral I, L epandrium, ventral J, K phallic complex, ventral and lateral M ejaculatory apodeme, lateral." figureDoi="10.3897/zookeys.1133.94530.figure6" httpUri="https://binary.pensoft.net/fig/775379" pageId="0" pageNumber="1">6F</figureCitation>
|
||
). Wing length 1.6 mm.
|
||
<emphasis id="6A7A1FB98733FB0D52866EEC2F1271C9" bold="true" italics="true" pageId="0" pageNumber="1">Postabdomen</emphasis>
|
||
<emphasis id="9347811E07E505EF15056FE872E28F8E" bold="true" pageId="0" pageNumber="1">:</emphasis>
|
||
(Fig.
|
||
<figureCitation id="842D4CC8C3629329A485CACBCC0B096D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7A, B</figureCitation>
|
||
) Oviscape dark brown, setigerous (Fig.
|
||
<figureCitation id="9098A9A9D5B924C966D97F30DE5E1464" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7A</figureCitation>
|
||
). Tergite 10 cruciform, laterally uniting narrow pleural sclerites (Fig.
|
||
<figureCitation id="3467B9E9AA9DB8CF1DD1E7F987760AA4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7B</figureCitation>
|
||
). Cercus with two stout, apical, trichoid sensilla,
|
||
<normalizedToken id="06E5DC50F1BD094039E8028CBE1A4795" originalValue="½">1/2</normalizedToken>
|
||
length of cercus. Spermathecae semi-orbicular (Fig.
|
||
<figureCitation id="11388EEF1C37D982DCBFE8AEB2AF6A0E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="996014D23D51817FC85CCD259FDA6F9A" doi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
|
||
<paragraph id="521C47ED9FC18DCA79361FA427E4B66A" pageId="0" pageNumber="1">
|
||
<emphasis id="80D087E543A535599CE3ED02F3E80398" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
|
||
Female morphology and larval ecology of
|
||
<taxonomicName id="15E2BA833EF906CC43B2A8FF0AA196ED" authorityName="Kato" authorityYear="2022" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza marchantiae" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marchantiae">
|
||
<emphasis id="A613D89B1C3B98573ED67C46A5A31A36" italics="true" pageId="0" pageNumber="1">Phytoliriomyza marchantiae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="033BCC692C40ED5CFCDBBDED3141C3F2" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
female postabdomen (at Renge-onsen)
|
||
<emphasis id="C535E16FE122F9D2ECAB76708B4E90B3" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
oviscape and spermatheca
|
||
<emphasis id="3FB75591A8DFE07634EC644B07C2017D" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
tergite 10
|
||
<emphasis id="FC1B1C4FF4891401B2B996D80E857699" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
|
||
habitat and mined thalli on
|
||
<taxonomicName id="7B4C5985B30C821130CDCA4D238D6C9A" authority="paleacea" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="paleacea">
|
||
<emphasis id="595948F1BCAC3EE470B376BFA559C795" italics="true" pageId="0" pageNumber="1">Marchantia paleacea paleacea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D2FAD897C6EE76B4428111DF48E49725" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
habitat at type locality
|
||
<emphasis id="2546C12B254310E384E0D908436930D9" bold="true" pageId="0" pageNumber="1">D, E</emphasis>
|
||
mined thalli at Yashajin-toge and Renge-onsen
|
||
<emphasis id="A109067F7A11706D5166A6B04491220F" bold="true" pageId="0" pageNumber="1">F-H</emphasis>
|
||
habitat and mined thalli on
|
||
<taxonomicName id="55C7D789AA07E03CD09DDE96730449FB" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. p. subsp. diptera" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="diptera">
|
||
<emphasis id="B9777E0F5A362F776B12684832F9B278" italics="true" pageId="0" pageNumber="1">M. p. diptera</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8009AD631751C6E5F22960390F169E6A" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
habitat at Kuchisakamoto
|
||
<emphasis id="406ABD97273FD3139BCC1A8CBB3AC1D0" bold="true" pageId="0" pageNumber="1">G, H</emphasis>
|
||
mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="8026B6CDCE4911D2EAE2EE99257BA2BA" pageId="0" pageNumber="1" type="variation">
|
||
<paragraph id="DA33DB358C83F25D7B254DEC2D95DC7D" pageId="0" pageNumber="1">Variation.</paragraph>
|
||
<paragraph id="DA029B614726DA037710E018E00308EB" pageId="0" pageNumber="1">The yellow patch on posterior scutum slightly varied from distinct to obscure ones among localities but not among host liverwort species/subspecies (Fig, 6F-K).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="12B58AB5A7136C2A6CF399B9264F01F9" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="BE322901447EB130D7847BDE9D4031E9" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="30977AC50EED2CFD960FBC8A88A4EEF2" pageId="0" pageNumber="1">
|
||
The specific name refers to host plant genus,
|
||
<taxonomicName id="115C91C82A2D32EB237320C9E4B4BB85" authorityName="C.Linnaeus" authorityYear="1753" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="A5F24A085BEB706B1DDCBD3F4BBEBDA0" italics="true" pageId="0" pageNumber="1">Marchantia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="81FFE533CF27768E9A1DE707E3A14650" pageId="0" pageNumber="1" type="japanese name">
|
||
<paragraph id="A33C37366BBA61CE2E7669AA15907DDB" pageId="0" pageNumber="1">Japanese name.</paragraph>
|
||
<paragraph id="41697824D66D36D719FB67D02ED7CCAB" pageId="0" pageNumber="1">Uzumibi-zenigoke-hamoguribae.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="937626F05A6DE2C6C8A3E1BD9FCBDB62" pageId="0" pageNumber="1" type="host plants">
|
||
<paragraph id="C9BC1A4781CE28CE80F6160407964102" pageId="0" pageNumber="1">Host plants.</paragraph>
|
||
<paragraph id="66759535A70D3921F9D7E872055D4E9C" pageId="0" pageNumber="1">
|
||
The main host plants are
|
||
<taxonomicName id="B523D94202034BBE0E444E57D570258B" authority="paleacea" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="paleacea">
|
||
<emphasis id="3D6BFFF7DD454C2E9B06EAB603A13058" italics="true" pageId="0" pageNumber="1">Marchantia paleacea paleacea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="04B1BAB21F3B1AA6FAE0CF0ED9B190A7" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. p. subsp. diptera" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="diptera">
|
||
<emphasis id="A151DAAB6FB85AE81AA022154F42A9C6" italics="true" pageId="0" pageNumber="1">M. p. diptera</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="C14FF84516E3B97CC0EE9244902828D4" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="family">Marchantiaceae</taxonomicName>
|
||
), with
|
||
<taxonomicName id="BFAC151DDF0E4E99443137F9C68E28C3" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. polymorpha" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="polymorpha">
|
||
<emphasis id="2DFD245B518E9BE09F698A79CD536690" italics="true" pageId="0" pageNumber="1">M. polymorpha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A50E5E8325CF949052B6DC065AA0C40A" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. papillata subsp. grossibarba" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="papillata" subSpecies="grossibarba">
|
||
<emphasis id="592778E2E39BC2B39D529E1380107E91" italics="true" pageId="0" pageNumber="1">M. papillata grossibarba</emphasis>
|
||
</taxonomicName>
|
||
also recorded as host in some localities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DD8F5FDACC41425F9C06D16890AA816F" pageId="0" pageNumber="1" type="mine">
|
||
<paragraph id="5B5041F8816AF6D91CA42A240C83E63E" pageId="0" pageNumber="1">Mine.</paragraph>
|
||
<paragraph id="483E5B0997A4F7754066454F250D1C74" pageId="0" pageNumber="1">
|
||
Larvae construct linear mines in the thallus, particularly in the midrib, and pupate in the mined thalli (Fig.
|
||
<figureCitation id="42345C8CAFCF0CFB5ACC18C7B7F63F2F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7D, E, G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E477448D3E8921C553C596998B148D16" pageId="0" pageNumber="1" type="biological notes">
|
||
<paragraph id="CBA10289B3F236C47C4FF1012F13A5FD" pageId="0" pageNumber="1">Biological notes.</paragraph>
|
||
<paragraph id="D6F46CD505920EB409E4CCEC70A05714" pageId="0" pageNumber="1">
|
||
<taxonomicName id="24667F5C1985F6827000B58CB1EFEA83" authority="paleacea" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia paleacea subsp. paleacea" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="paleacea" subSpecies="paleacea">
|
||
<emphasis id="8B3601B42D0ABDC9047B3D482B7C6821" italics="true" pageId="0" pageNumber="1">Marchantia paleacea paleacea</emphasis>
|
||
</taxonomicName>
|
||
grows on rocky substrates of cliffs, slopes and riverbanks in cool temperate deciduous forests (Fig.
|
||
<figureCitation id="31F219E737F8D538B3633A5C28BF1799" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7C</figureCitation>
|
||
) and
|
||
<taxonomicName id="5B427BABAF4F2D969C3B1C658AB2E0BF" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia p. subsp. diptera" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="p." subSpecies="diptera">
|
||
<emphasis id="77E95922EEB194A40EB0B8FA187F5954" italics="true" pageId="0" pageNumber="1">Marchantia p. diptera</emphasis>
|
||
</taxonomicName>
|
||
in warm temperate evergreen forests (Fig.
|
||
<figureCitation id="CC46554BF88A203DFD35FC5A06A8F396" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female morphology and larval ecology of Phytoliriomyza marchantiae sp. nov. A, B female postabdomen (at Renge-onsen) A oviscape and spermatheca B tergite 10 C-E habitat and mined thalli on Marchantia paleacea paleacea C habitat at type locality D, E mined thalli at Yashajin-toge and Renge-onsen F-H habitat and mined thalli on M. p. diptera F habitat at Kuchisakamoto G, H mined thalli at Iwaya-kei and Tosayama. Arrows indicate internal puparia." figureDoi="10.3897/zookeys.1133.94530.figure7" httpUri="https://binary.pensoft.net/fig/775380" pageId="0" pageNumber="1">7F</figureCitation>
|
||
). While these subspecies are frequently utilized by
|
||
<taxonomicName id="0C933B9EB9BCCFCDF9DB3FF43B512D41" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. marchantiae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="marchantiae">
|
||
<emphasis id="18F6D91C973FE025C0A6BE1F4C0954EB" italics="true" pageId="0" pageNumber="1">P. marchantiae</emphasis>
|
||
</taxonomicName>
|
||
, they are never utilized by the other three
|
||
<taxonomicName id="8A0242A89482154CE37A1DFF61776E8D" authorityName="C.Linnaeus" authorityYear="1753" class="Marchantiopsida" family="Marchantiaceae" genus="Marchantia" kingdom="Plantae" lsidName="Marchantia" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="3C834CA7C9C4D69FA21A873F995778E7" italics="true" pageId="0" pageNumber="1">Marchantia</emphasis>
|
||
</taxonomicName>
|
||
-associated species,
|
||
<taxonomicName id="7E64B3306856A7E87816B4E1FA363376" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. igniculus" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="igniculus">
|
||
<emphasis id="756FD9C6E85BFC6C0683B6AF88A7450D" italics="true" pageId="0" pageNumber="1">P. igniculus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3EEE0F0403FA22AC0A412A39EFBF64D5" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. tsukuyomi" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="tsukuyomi">
|
||
<emphasis id="014DCD95506BFAE9A494455FB8D090A0" italics="true" pageId="0" pageNumber="1">P. tsukuyomi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="43E90EC2B9158C3772B6198CF2127D91" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. nubatama" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="nubatama">
|
||
<emphasis id="DBD675B58B92A0BD8757B140306A5E0B" italics="true" pageId="0" pageNumber="1">P. nubatama</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="54E77BAFD4521410E603C5A339DD4A4F" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. marchantiae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="marchantiae">
|
||
<emphasis id="2C0B4F2A7D8DFF9CAF284DB9EF531E87" italics="true" pageId="0" pageNumber="1">P. marchantiae</emphasis>
|
||
</taxonomicName>
|
||
is also recorded from
|
||
<taxonomicName id="0CA13909348EC0A4001FE43916E5C88D" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. polymorpha" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="polymorpha">
|
||
<emphasis id="97C68488F71A4A9DC9176A6F56AAB719" italics="true" pageId="0" pageNumber="1">M. polymorpha</emphasis>
|
||
</taxonomicName>
|
||
in cool temperate forests and
|
||
<taxonomicName id="5F4E5458E9C354E2166821715395845F" class="Marchantiopsida" family="Marchantiaceae" kingdom="Plantae" lsidName="M. papillata subsp. grossibarba" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="subSpecies" species="papillata" subSpecies="grossibarba">
|
||
<emphasis id="00CECDFD499E719763A2E27B0D133019" italics="true" pageId="0" pageNumber="1">M. papillata grossibarba</emphasis>
|
||
</taxonomicName>
|
||
in warm temperate evergreen forests in some localities. Our rearing records suggest that this species is multivoltine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55FFA6BE90EEAEDD0E9944462582FF3A" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="5C272D46EDFB07E6ECEEC4BDFAF2CD89" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="1583745AF0F1DD7A2726D76A648FF277" pageId="0" pageNumber="1">
|
||
Japan: Hokkaido, Honshu, Shikoku, Kyushu (Fig.
|
||
<figureCitation id="7F41196B037055F8BB086C59B21E9166" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Locality records of three Phytoliriomyza species associated with Marchantia spp.: P. dorsata, P. igniculus, P. tsukuyomi, P. marchantiae and P. nubatama." figureDoi="10.3897/zookeys.1133.94530.figure2" httpUri="https://binary.pensoft.net/fig/775375" pageId="0" pageNumber="1">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7EFA874DECFF1F723AC773FB2AFC9FAD" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="27B260ECE5DE7F5FAFB84B7CE6646926" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="4740B7D1D39B0AAB2FB6325D15CAB3EB" pageId="0" pageNumber="1">
|
||
This species resembles
|
||
<taxonomicName id="93110E583CBD3AE8BA581E93F58FAA28" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. rebouliae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="rebouliae">
|
||
<emphasis id="4FDF5E7757D1F7525C920456464F248B" italics="true" pageId="0" pageNumber="1">P. rebouliae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="CFFF69C1CDB4A28FE002003DA195C2BE" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. lanternaria" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="lanternaria">
|
||
<emphasis id="A16394E8D1A662507487296338472BD4" italics="true" pageId="0" pageNumber="1">P. lanternaria</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="87BEC2514C1D47661CBFCBC8FABD77E4" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. conocephali" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="conocephali">
|
||
<emphasis id="74D24D0D6399FC5ADE75F6BED1AD03CE" italics="true" pageId="0" pageNumber="1">P. conocephali</emphasis>
|
||
</taxonomicName>
|
||
in the narrow yellow posterior margin of the scutum and medial yellow stripe of the scutellum; it is distinguished from
|
||
<taxonomicName id="64E732054FCAA5BDEFCBF416AD955636" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. rebouliae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="rebouliae">
|
||
<emphasis id="2EC0AA42D7F858E4DCB08345EC6BE662" italics="true" pageId="0" pageNumber="1">P. rebouliae</emphasis>
|
||
</taxonomicName>
|
||
by number of tubercle-like setae in a comb of the male epandrium (8 in
|
||
<taxonomicName id="E4064B62A60D79D35B9C168D06110538" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. marchantiae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="marchantiae">
|
||
<emphasis id="E765D9FCAF64C3D6101477A3D1FA7959" italics="true" pageId="0" pageNumber="1">P. marchantiae</emphasis>
|
||
</taxonomicName>
|
||
; 7 in
|
||
<taxonomicName id="81F7B1C8D8A56D7D5FDE868BBB1CED18" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. rebouliae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="rebouliae">
|
||
<emphasis id="CFBA7BDF9C73A4E9D4B9A58787ACC5E0" italics="true" pageId="0" pageNumber="1">P. rebouliae</emphasis>
|
||
</taxonomicName>
|
||
), from
|
||
<taxonomicName id="1F4142C371A73EC5F86B79D2471CE56F" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. lanternaria" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="lanternaria">
|
||
<emphasis id="2B967ACD1C603C0D83B11FD207F41905" italics="true" pageId="0" pageNumber="1">P. lanternaria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="186D505D7C65DF8B214756034D7D6D04" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. conocephali" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="conocephali">
|
||
<emphasis id="60C1D6FCACB336C719BF53CDE8B44D3A" italics="true" pageId="0" pageNumber="1">P. conocephali</emphasis>
|
||
</taxonomicName>
|
||
by the number of tubercle-like setae on the surstylus of the male epandrium (0 in
|
||
<taxonomicName id="93626C6FC35F0E2C5F979F098D3FEA90" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. marchantiae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="marchantiae">
|
||
<emphasis id="9B17E1031DEC9E97C312A357EE82FCDC" italics="true" pageId="0" pageNumber="1">P. marchantiae</emphasis>
|
||
</taxonomicName>
|
||
; 1-2 in
|
||
<taxonomicName id="81BF44F44B24E71C4F9E068DE079767A" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. lanternaria" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="lanternaria">
|
||
<emphasis id="859865F0832BA058CC0200180CF51DAF" italics="true" pageId="0" pageNumber="1">P. lanternaria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D835115E8A45EF91B7B61C69C8EAADA1" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. conocephali" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="conocephali">
|
||
<emphasis id="988C4C573EDCACA276C307B753F08562" italics="true" pageId="0" pageNumber="1">P. conocephali</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7C932268AC60712A3628210649E8D714" pageId="0" pageNumber="1">
|
||
This species also resembles
|
||
<taxonomicName id="915BA9765BED11E8C45038ACCAD3B3C4" class="Insecta" family="Dendrocerotaceae" genus="Phytoliriomyza" kingdom="Animalia" lsidName="Phytoliriomyza miki" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="miki">
|
||
<emphasis id="7ADF10E1DE38191EA0492018097F0D16" italics="true" pageId="0" pageNumber="1">Phytoliriomyza miki</emphasis>
|
||
</taxonomicName>
|
||
(Strobl, 1898) in color pattern of the scutum and morphology of the male genitalia; it is distinguished from the latter by the number of tubercle-like setae in a comb on the male epandrium (8 in
|
||
<taxonomicName id="8C985BA27C8467BEB9116E50F2EDDDC3" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. marchantiae" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="marchantiae">
|
||
<emphasis id="5E9E99CCBD463E5AF0CC732CCCDE56E6" italics="true" pageId="0" pageNumber="1">P. marchantiae</emphasis>
|
||
</taxonomicName>
|
||
; 5 in
|
||
<taxonomicName id="036F20A7E82BB1AD406560F9508AAFE4" class="Marchantiopsida" family="Aytoniaceae" kingdom="Plantae" lsidName="P. miki" order="Marchantiales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="miki">
|
||
<emphasis id="5F7539D52572987C11BA2CBBD8EF7FD4" italics="true" pageId="0" pageNumber="1">P. miki</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |