plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/08/7E/16/087E167D2A59574098026EED663B2B18.xml
maintenance 835ba71d3d added 08
2024-06-21 12:28:49 +02:00

430 lines
49 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1063.69942" ID-GBIF-Dataset="28e9e42b-c1df-47e6-a327-6e4cbc06db46" ID-PMC="PMC8545786" ID-Pensoft-Pub="1313-2970-1063-1" ID-Pensoft-UUID="94FEB0FF5E515631AA6239A1BB296EEB" ID-PubMed="34720622" ID-ZooBank="3687D7229CD64D82ACF8D6701E399F4B" ModsDocID="1313-2970-1063-1" checkinTime="1634632146073" checkinUser="pensoft" docAuthor="Yin, Wen-Qi, Shi, Hong-Liang &amp; Liang, Hong-Bin" docDate="2021" docId="087E167D2A59574098026EED663B2B18" docLanguage="en" docName="ZooKeys 1063: 1-21" docOrigin="ZooKeys 1063" docPubDate="2021-10-18" docSource="http://dx.doi.org/10.3897/zookeys.1063.69942" docTitle="Pterostichus (Koreonialoe) syleus Kirschenhofer 1997" docType="treatment" docVersion="4" id="94FEB0FF5E515631AA6239A1BB296EEB" lastPageNumber="1" masterDocId="94FEB0FF5E515631AA6239A1BB296EEB" masterDocTitle="Revision of the Chinese species of subgenus Koreonialoe (Coleoptera: Carabidae: Pterostichus), with descriptions of two new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="1" updateTime="1668150977456" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Chinese species of subgenus Koreonialoe (Coleoptera: Carabidae: Pterostichus), with descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Wen-Qi</mods:namePart>
<mods:affiliation>College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hong-Liang</mods:namePart>
<mods:affiliation>College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hong-Bin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-10-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1063</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>21</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1063.69942</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1063.69942</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1063-1</mods:identifier>
<mods:identifier type="ZooBank">3687D7229CD64D82ACF8D6701E399F4B</mods:identifier>
<mods:identifier type="Pensoft-UUID">94FEB0FF5E515631AA6239A1BB296EEB</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="1" type="taxonomy">
<treatment ID-GBIF-Taxon="189468675" LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18" httpUri="http://treatment.plazi.org/id/087E167D2A59574098026EED663B2B18" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
3.
<taxonomicName LSID="087E167D-2A59-5740-9802-6EED663B2B18" authority="Kirschenhofer, 1997" authorityName="Kirschenhofer" authorityYear="1997" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe) syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus" subGenus="Koreonialoe">Pterostichus (Koreonialoe) syleus Kirschenhofer, 1997</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus of Pterostichus (Pterostichus Koreonialoe) spp. from China 1 P. micropoides sp. nov., Holotype. 2 P. micropoides sp. nov., male, Paratype from Hunchun city, Jilin 3 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 4 P. syleus Kirschenhofer, a female from Zhuanghe city, Liaoning 5 P. quadrilobatus sp. nov., Holotype 6 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1063.69942.figures1-6" httpUri="https://binary.pensoft.net/fig/599856" pageId="0" pageNumber="1">Figs 3</figureCitation>
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus of Pterostichus (Pterostichus Koreonialoe) spp. from China 1 P. micropoides sp. nov., Holotype. 2 P. micropoides sp. nov., male, Paratype from Hunchun city, Jilin 3 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 4 P. syleus Kirschenhofer, a female from Zhuanghe city, Liaoning 5 P. quadrilobatus sp. nov., Holotype 6 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1063.69942.figures1-6" httpUri="https://binary.pensoft.net/fig/599856" pageId="0" pageNumber="1">, 4</figureCitation>
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Male genitalia of Pterostichus (Koreonialoe) spp. from China, left lateral view of median lobe of aedeagus 7 P. micropoides sp. nov., Holotype 8 P. micropoides sp. nov., Paratype from Changbai county, Jilin 9 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 10 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 11 P. quadrilobatus sp. nov., Holotype 12 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures7-12" httpUri="https://binary.pensoft.net/fig/599857" pageId="0" pageNumber="1">, 9</figureCitation>
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Male genitalia of Pterostichus (Koreonialoe) spp. from China, left lateral view of median lobe of aedeagus 7 P. micropoides sp. nov., Holotype 8 P. micropoides sp. nov., Paratype from Changbai county, Jilin 9 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 10 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 11 P. quadrilobatus sp. nov., Holotype 12 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures7-12" httpUri="https://binary.pensoft.net/fig/599857" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus (Koreonialoe) spp. from China, dorsal view of median lobe of aedeagus 13 P. micropoides sp. nov., Holotype 14 P. micropoides sp. nov. Paratype from Changbai county 15 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 16 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 17 P. quadrilobatus sp. nov., Paratype from Laopingding, Liaoning 18 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures13-18" httpUri="https://binary.pensoft.net/fig/599858" pageId="0" pageNumber="1">, 15</figureCitation>
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus (Koreonialoe) spp. from China, dorsal view of median lobe of aedeagus 13 P. micropoides sp. nov., Holotype 14 P. micropoides sp. nov. Paratype from Changbai county 15 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 16 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 17 P. quadrilobatus sp. nov., Paratype from Laopingding, Liaoning 18 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures13-18" httpUri="https://binary.pensoft.net/fig/599858" pageId="0" pageNumber="1">, 16</figureCitation>
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">, 24</figureCitation>
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">, 25</figureCitation>
<figureCitation captionStart="Figures 28–35" captionStartId="F6" captionText="Figures 28 - 35. Female genitalia of Pterostichus (Koreonialoe) spp. from China, Figs 28 - 31 ventral view and inner lateral view of ovipositor. Figs 32 - 35 female reproductive system. 28, 32 P. tetralobatus, Paratype from Laopingding, Liaoning 29, 33 P. syleus, a female from Baishilazi, Liaoning 30, 34 P. micropoides, female, Paratype from Antu county, Jilin 31, 35 P. bellatrix, a female from Fusong county, Jilin. Scale bar: 0.5 mm. Abbreviations: g 1: gonocoxite I; g 2: gonocoxite II; es: ensiform setae; ns, nematiform setae; bc: bursa copulatrix; co: common oviduct; sc: seminal canal; rc: receptaculum; spc: spermathecal canal." figureDoi="10.3897/zookeys.1063.69942.figures28-35" httpUri="https://binary.pensoft.net/fig/599861" pageId="0" pageNumber="1">, 29</figureCitation>
<figureCitation captionStart="Figures 28–35" captionStartId="F6" captionText="Figures 28 - 35. Female genitalia of Pterostichus (Koreonialoe) spp. from China, Figs 28 - 31 ventral view and inner lateral view of ovipositor. Figs 32 - 35 female reproductive system. 28, 32 P. tetralobatus, Paratype from Laopingding, Liaoning 29, 33 P. syleus, a female from Baishilazi, Liaoning 30, 34 P. micropoides, female, Paratype from Antu county, Jilin 31, 35 P. bellatrix, a female from Fusong county, Jilin. Scale bar: 0.5 mm. Abbreviations: g 1: gonocoxite I; g 2: gonocoxite II; es: ensiform setae; ns, nematiform setae; bc: bursa copulatrix; co: common oviduct; sc: seminal canal; rc: receptaculum; spc: spermathecal canal." figureDoi="10.3897/zookeys.1063.69942.figures28-35" httpUri="https://binary.pensoft.net/fig/599861" pageId="0" pageNumber="1">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Kirschenhofer" authorityYear="1997" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe) syleus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="syleus" subGenus="Koreonialoe">Pterostichus (Koreonialoe) syleus</taxonomicName>
Kirschenhofer, 1997: 694 (Holotype deposited in Naturhistorisches Museum Wien; type locality: &quot;Shi-fang Ding&quot;, sg.
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Nialoe">Pterostichus Nialoe</taxonomicName>
);
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. 2013</bibRefCitation>
: 430 (based on the misidentification of an unknown species).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
(
<specimenCount type="male">8 males</specimenCount>
and
<specimenCount type="female">16 females</specimenCount>
):
<specimenCount type="male">4 males</specimenCount>
,
<materialsCitation collectingDate="2008-08-15" collectingMethod="pitfall trap" country="China" county="Dandong" latitude="41.0143" location="Mixed forest" longLatPrecision="6" longitude="124.302" municipality="Saima town" specimenCount="1" stateProvince="Liaoning">
&quot;
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Liaoning">Liaoning</collectingRegion>
,
<collectingCounty>Dandong</collectingCounty>
city,
<collectingMunicipality>Saima town</collectingMunicipality>
;
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:543B6859596EB2D5809DD02F5B590010" country="China" county="Dandong" latitude="41.0143" longLatPrecision="6" longitude="124.302" municipality="Saima town" name="Mixed forest" stateProvince="Liaoning">Mixed forest</location>
,
<geoCoordinate degrees="41.0143" direction="north" orientation="latitude" precision="5" value="41.0143">41.0143N</geoCoordinate>
,
<geoCoordinate degrees="124.3020" direction="east" orientation="longitude" precision="5" value="124.302">124.3020E</geoCoordinate>
&quot;; &quot;Day,
<collectingDate value="2008-08-15">2008.VIII.15</collectingDate>
,
<collectingMethod>pitfall trap</collectingMethod>
, Shan HC lgt.&quot;
</materialsCitation>
;
<specimenCount type="male">1 male</specimenCount>
,
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="China" location="Snake island" municipality="Dalian City" specimenCount="1" stateProvince="Liaoning province">
&quot;
<collectingRegion country="China" name="Liaoning">Liaoning province</collectingRegion>
,
<collectingMunicipality>Dalian City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:89A32FE9B1BC2DE8A6B1A51D93DBDF83" country="China" municipality="Dalian City" name="Snake island" stateProvince="Liaoning province">Snake island</location>
; 2012, Shi JS lgt.&quot;
</materialsCitation>
;
<specimenCount type="male">2 males</specimenCount>
,
<materialsCitation collectingDate="2011-08-20" country="China" elevation="530" latitude="41.5644" location="Guanmenshan mountain" longLatPrecision="6" longitude="123.5779" municipality="Benxi City" specimenCount="1" stateProvince="Liaoning">
&quot;
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Liaoning">Liaoning</collectingRegion>
,
<collectingMunicipality>Benxi City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:8D8C16E2D9068D2BDF30F6FF44EFA431" country="China" latitude="41.5644" longLatPrecision="6" longitude="123.5779" municipality="Benxi City" name="Guanmenshan mountain" stateProvince="Liaoning">Guanmenshan mountain</location>
;
<geoCoordinate degrees="41.5644" direction="north" orientation="latitude" precision="5" value="41.5644">41.5644N</geoCoordinate>
,
<geoCoordinate degrees="123.5779" direction="east" orientation="longitude" precision="5" value="123.5779">123.5779E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.3" unit="m" value="530.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.3" unit="m" value="530.0">530m</elevation>
</quantity>
,
<collectingDate value="2011-08-20">2011.VIII.20</collectingDate>
, night, Huang XL lgt.&quot;
</materialsCitation>
;
<specimenCount type="male">1 male</specimenCount>
,
<materialsCitation collectingDate="2011-08-23" country="China" county="Benxi City" elevation="530" latitude="41.5644" location="Guanmenshan mountain" longLatPrecision="6" longitude="123.5779" municipality="Benxi county" specimenCount="1" stateProvince="Liaoning province">
&quot;
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Liaoning">Liaoning province</collectingRegion>
,
<collectingCounty>Benxi City</collectingCounty>
,
<collectingMunicipality>Benxi county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:EDEDC958B7DC6733E069C94A8758F253" country="China" county="Benxi City" latitude="41.5644" longLatPrecision="6" longitude="123.5779" municipality="Benxi county" name="Guanmenshan mountain" stateProvince="Liaoning province">Guanmenshan mountain</location>
;
<geoCoordinate degrees="41.5644" direction="north" orientation="latitude" precision="5" value="41.5644">41.5644N</geoCoordinate>
,
<geoCoordinate degrees="123.5779" direction="east" orientation="longitude" precision="5" value="123.5779">123.5779E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.3" unit="m" value="530.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.3" unit="m" value="530.0">530m</elevation>
</quantity>
,
<collectingDate value="2011-08-23">2011.VIII.23</collectingDate>
, day, Huang XL lgt.&quot;
</materialsCitation>
;
<specimenCount type="male">2 males</specimenCount>
and
<specimenCount type="female">5 females</specimenCount>
,
<materialsCitation collectingDate="2019-08-15" collectingMethod="pitfall trap" country="China" elevation="1133" latitude="40.0862" location="Buyunshan mountain" longLatPrecision="6" longitude="122.7234" municipality="Zhuanghe City" specimenCount="1" stateProvince="Liaoning province">
&quot;
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Liaoning">Liaoning province</collectingRegion>
,
<collectingMunicipality>Zhuanghe City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:929DB29B0E2394981C1CF17097EC2338" country="China" latitude="40.0862" longLatPrecision="6" longitude="122.7234" municipality="Zhuanghe City" name="Buyunshan mountain" stateProvince="Liaoning province">Buyunshan mountain</location>
;
<geoCoordinate degrees="40.0862" direction="north" orientation="latitude" precision="5" value="40.0862">40.0862N</geoCoordinate>
,
<geoCoordinate degrees="122.7234" direction="east" orientation="longitude" precision="5" value="122.7234">122.7234E</geoCoordinate>
,
<collectingMethod>pitfall trap</collectingMethod>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.133" unit="m" value="1133.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.133" unit="m" value="1133.0">1133m</elevation>
</quantity>
&quot;; &quot;
<collectingDate value="2019-08-15">2019.VIII.15</collectingDate>
, Zhu PZ &amp; Wang C lgt.&quot;
</materialsCitation>
;
<specimenCount type="female">6 females</specimenCount>
,
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="China" county="Fushun" location="Nankouqian town" municipality="Qingyuan county" specimenCount="1" stateProvince="Liaoning">
&quot;
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Liaoning">Liaoning</collectingRegion>
,
<collectingCounty>Fushun</collectingCounty>
city,
<collectingMunicipality>Qingyuan county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:36E1BAAECE2E382542AD81AED12DE8ED" country="China" county="Fushun" municipality="Qingyuan county" name="Nankouqian town" stateProvince="Liaoning">Nankouqian town</location>
; mixed forest; 2020.VIII, local collector.&quot;
</materialsCitation>
;
<specimenCount type="female">2 females</specimenCount>
,
<materialsCitation collectingDate="2015-08-14" country="China" elevation="567" latitude="40.9394" location="Baishilazi" longLatPrecision="6" longitude="124.8025" specimenCount="1" stateProvince="Liaoning province">
&quot;
<collectingRegion country="China" name="Liaoning">Liaoning province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:EF1F497B77984F1DD52A6EAE328AB6EA" country="China" latitude="40.9394" longLatPrecision="6" longitude="124.8025" name="Baishilazi" stateProvince="Liaoning province">Baishilazi</location>
,
<geoCoordinate degrees="40.9394" direction="north" orientation="latitude" precision="5" value="40.9394">40.9394N</geoCoordinate>
,
<geoCoordinate degrees="124.8025" direction="east" orientation="longitude" precision="5" value="124.8025">124.8025E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.67" unit="m" value="567.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.67" unit="m" value="567.0">567m</elevation>
</quantity>
,
<collectingDate value="2015-08-14">2015.VIII.14</collectingDate>
, mixed forest&quot;
</materialsCitation>
;
<specimenCount type="female">1 female</specimenCount>
,
<materialsCitation collectingDate="2015-08-15" country="China" elevation="713" latitude="40.9385" location="Baishilazi" longLatPrecision="6" longitude="124.7891" specimenCount="1" stateProvince="Liaoning province">
&quot;
<collectingRegion country="China" name="Liaoning">Liaoning province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:2B0DD6CD678FE56EAC000977E86D19F1" country="China" latitude="40.9385" longLatPrecision="6" longitude="124.7891" name="Baishilazi" stateProvince="Liaoning province">Baishilazi</location>
,
<geoCoordinate degrees="40.9385" direction="north" orientation="latitude" precision="5" value="40.9385">40.9385N</geoCoordinate>
,
<geoCoordinate degrees="124.7891" direction="east" orientation="longitude" precision="5" value="124.7891">124.7891E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.13" unit="m" value="713.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.13" unit="m" value="713.0">713m</elevation>
</quantity>
,
<collectingDate value="2015-08-15">2015.VIII.15</collectingDate>
, broadleaved mixed forest&quot;
</materialsCitation>
;
<specimenCount type="female">2 females</specimenCount>
,
<materialsCitation collectingDate="2015-09-12" country="China" elevation="591" latitude="40.9412" location="Baishilazi" longLatPrecision="6" longitude="124.7986" specimenCount="1" stateProvince="Liaoning province">
&quot;
<collectingRegion country="China" name="Liaoning">Liaoning province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:087E167D2A59574098026EED663B2B18:71299F249904BBBF9A5EB833F9375452" country="China" latitude="40.9412" longLatPrecision="6" longitude="124.7986" name="Baishilazi" stateProvince="Liaoning province">Baishilazi</location>
,
<geoCoordinate degrees="40.9412" direction="north" orientation="latitude" precision="5" value="40.9412">40.9412N</geoCoordinate>
,
<geoCoordinate degrees="124.7986" direction="east" orientation="longitude" precision="5" value="124.7986">124.7986E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.91" unit="m" value="591.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.91" unit="m" value="591.0">591m</elevation>
</quantity>
,
<collectingDate value="2015-09-12">2015.IX.12</collectingDate>
, mixed forest&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsally dark reddish brown; fifth tarsomere without ventral seta. Median lobe of aedeagus stout, ventral surface slightly tumid forming an inconspicuous tubercle. Endophallus elongate, directed basal-ventrally, gonopore opened to the ventral-basal direction of aedeagus; three lobes present: la and lp both oblate, without hook at apex, similar in size and well separate.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comparisons">
<paragraph pageId="0" pageNumber="1">Comparison.</paragraph>
<paragraph pageId="0" pageNumber="1">
From the sclerotized features of male genitalia,
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="tetralobatus">
<emphasis italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
</taxonomicName>
sp. nov. for they both have an inconspicuous tubercle on the ventral margin of median lobe. But they are quite different in the endophallus.
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
is distinguishable from
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="tetralobatus">
<emphasis italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
</taxonomicName>
sp. nov. for the endophallus with la not divided and ra only very faintly defined, while in the latter species with la divided into two sub-lobes, and ra well defined.
</paragraph>
<paragraph pageId="0" pageNumber="1">
From the external features,
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. micropoides" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="micropoides">
<emphasis italics="true" pageId="0" pageNumber="1">P. micropoides</emphasis>
</taxonomicName>
by its slightly larger size (BL 16.8-19.7 mm versus 13.7-15.7 mm), and from
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. bellatrix" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="bellatrix">
<emphasis italics="true" pageId="0" pageNumber="1">P. bellatrix</emphasis>
</taxonomicName>
and
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. tetralobatus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="tetralobatus">
<emphasis italics="true" pageId="0" pageNumber="1">P. tetralobatus</emphasis>
</taxonomicName>
by the smaller size (BL 16.8-19.7 mm versus 19.7-21.6 mm) and lighter color (dark reddish brown versus nearly black).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
BL 16.8-19.7 mm, BW 6.7-7.2 mm. Body form robust, dorsally dark reddish brown, elytra opaque, without metallic luster. Head large, widest at temporae; frons smooth; frontal grooves shallow; temporae strongly swollen, a little shorter than eyes; eyes small and hemisphere; terminal segment of labial palpus fusiform. Pronotum strongly cordate, disc evenly and densely covered with fine punctures, area anterior to sub-anterior transversal sulci well punctate; PW/HW = 1.26-1.32, PW/PL = 1.46-1.51, widest near anterior third; anterior margin a little wider than posterior margin, PAW/PBW = 1.19-1.21. Lateral margins slightly arched from anterior angles to the middle, strongly sinuate and then nearly straight before posterior angles; posterior angles rectangular; mid-lateral setae present at anterior fifth of lateral margins; lateral expansions equal width anteriorly and posteriorly. Basal foveae with inner and outer grooves faintly defined and partly fused, forming deep depression between them, outer groove slightly shorter than inner one; middle area between two basal foveae longitudinal rugose. Elytra oblong, shoulders widely rounded; basal ridge and lateral margin forming an obtuse angle; elytra 1.59-1.64 times longer than wide. Usually three discal pores present on third interval, all adjoining the second stria; the first one before middle, position of the second one variable, the last one at apical sixth to eighth. Umbilicate series on ninth interval continuous, sparse at middle. Ventral side: metepisternum nearly smooth; sternite VII of males without secondary sexual modification. Fifth tarsomere without ventral seta. Male genitalia: median lobe of aedeagus stout, gradually curved at basal third, slightly dilated near middle, apical portion abruptly deflected ventrally; ventral surface shallowly tumid near middle, forming an inconspicuous tubercle (Figs
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Male genitalia of Pterostichus (Koreonialoe) spp. from China, left lateral view of median lobe of aedeagus 7 P. micropoides sp. nov., Holotype 8 P. micropoides sp. nov., Paratype from Changbai county, Jilin 9 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 10 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 11 P. quadrilobatus sp. nov., Holotype 12 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures7-12" httpUri="https://binary.pensoft.net/fig/599857" pageId="0" pageNumber="1">9</figureCitation>
,
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Male genitalia of Pterostichus (Koreonialoe) spp. from China, left lateral view of median lobe of aedeagus 7 P. micropoides sp. nov., Holotype 8 P. micropoides sp. nov., Paratype from Changbai county, Jilin 9 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 10 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 11 P. quadrilobatus sp. nov., Holotype 12 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures7-12" httpUri="https://binary.pensoft.net/fig/599857" pageId="0" pageNumber="1">10</figureCitation>
); apical lamella narrow, apex rounded-triangular, very faintly oblique to the left in dorsal view (Figs
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus (Koreonialoe) spp. from China, dorsal view of median lobe of aedeagus 13 P. micropoides sp. nov., Holotype 14 P. micropoides sp. nov. Paratype from Changbai county 15 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 16 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 17 P. quadrilobatus sp. nov., Paratype from Laopingding, Liaoning 18 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures13-18" httpUri="https://binary.pensoft.net/fig/599858" pageId="0" pageNumber="1">15</figureCitation>
,
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus (Koreonialoe) spp. from China, dorsal view of median lobe of aedeagus 13 P. micropoides sp. nov., Holotype 14 P. micropoides sp. nov. Paratype from Changbai county 15 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 16 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 17 P. quadrilobatus sp. nov., Paratype from Laopingding, Liaoning 18 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures13-18" httpUri="https://binary.pensoft.net/fig/599858" pageId="0" pageNumber="1">16</figureCitation>
). Endophallus (Figs
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">24</figureCitation>
,
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">25</figureCitation>
) long, directed basal-ventrally, gonopore opened to ventral-basal direction of aedeagus; three endophallic lobes recognized: right apical lobe (
<emphasis bold="true" pageId="0" pageNumber="1">ra</emphasis>
) faintly defined, close to gonopore; left apical lobe (
<emphasis bold="true" pageId="0" pageNumber="1">la</emphasis>
) oblate, apex not hooked; left preapical lobe (
<emphasis bold="true" pageId="0" pageNumber="1">lp</emphasis>
) similar shape and size as la, these two lobes well separated. Female genitalia typical in this subgenus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Widely distributed in the eastern part of Liaoning province, along the Qianshan mountain range. (Fig.
<figureCitation captionStart="Figure 36" captionStartId="F7" captionText="Figure 36. Confirmed distribution of Pterostichus (Koreonialoe) spp. in China: Green: P. micropoides sp. nov.; Red: P. tetralobatus sp. nov.; Yellow: P. bellatrix Tschitscherine; Blue: P. syleus Kirschenhofer." figureDoi="10.3897/zookeys.1063.69942.figure36" httpUri="https://binary.pensoft.net/fig/599862" pageId="0" pageNumber="1">36</figureCitation>
, blue)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. (2013)</bibRefCitation>
redescribed this species and illustrated its endophallus based on a specimen from the type locality (Shi-Fang-Ding, the highest peak of Baishilazi nature reserve). However, their description and illustrations do not accord with the original literature (
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="0" pageNumber="1" pagination="689 - 714" refId="B8" refString="Kirschenhofer, E, 1997. Neue Arten der Gattungen Pterostichus Bonelli 1810, Synuchus Gyllenhal 1810, Lesticus Dejean 1828 und Trigonotoma Dejean 1828 aus Ost- und Suedostasien (Coleoptera, Carabidae: Pterostichinae). Linzer Biologische Beitraege 29: 689 - 714" title="Neue Arten der Gattungen Pterostichus Bonelli 1810, Synuchus Gyllenhal 1810, Lesticus Dejean 1828 und Trigonotoma Dejean 1828 aus Ost- und Suedostasien (Coleoptera, Carabidae: Pterostichinae)." volume="29" year="1997">Kirschenhofer 1997</bibRefCitation>
) or our examined specimens from the type locality (Baishilazi).
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. (2013)</bibRefCitation>
indicated that
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
is the only species of
<taxonomicName authorityName="Park &amp; Kwon" authorityYear="1996" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Koreonialoe">
<emphasis italics="true" pageId="0" pageNumber="1">Pterostichus Koreonialoe</emphasis>
</taxonomicName>
with the fifth tarsomeres setose beneath, but in the original description
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="0" pageNumber="1" pagination="689 - 714" refId="B8" refString="Kirschenhofer, E, 1997. Neue Arten der Gattungen Pterostichus Bonelli 1810, Synuchus Gyllenhal 1810, Lesticus Dejean 1828 und Trigonotoma Dejean 1828 aus Ost- und Suedostasien (Coleoptera, Carabidae: Pterostichinae). Linzer Biologische Beitraege 29: 689 - 714" title="Neue Arten der Gattungen Pterostichus Bonelli 1810, Synuchus Gyllenhal 1810, Lesticus Dejean 1828 und Trigonotoma Dejean 1828 aus Ost- und Suedostasien (Coleoptera, Carabidae: Pterostichinae)." volume="29" year="1997">Kirschenhofer (1997)</bibRefCitation>
mentioned this species has: &quot;
<emphasis italics="true" pageId="0" pageNumber="1">Klauenglieder unterseits glatt</emphasis>
&quot;. Moreover, compared with the line drawing of the male genitalia of
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
(
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="0" pageNumber="1" pagination="689 - 714" refId="B8" refString="Kirschenhofer, E, 1997. Neue Arten der Gattungen Pterostichus Bonelli 1810, Synuchus Gyllenhal 1810, Lesticus Dejean 1828 und Trigonotoma Dejean 1828 aus Ost- und Suedostasien (Coleoptera, Carabidae: Pterostichinae). Linzer Biologische Beitraege 29: 689 - 714" title="Neue Arten der Gattungen Pterostichus Bonelli 1810, Synuchus Gyllenhal 1810, Lesticus Dejean 1828 und Trigonotoma Dejean 1828 aus Ost- und Suedostasien (Coleoptera, Carabidae: Pterostichinae)." volume="29" year="1997">Kirschenhofer 1997</bibRefCitation>
: Fig.
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">23</figureCitation>
), specimen described by Sasakawa et al. (
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. 2013</bibRefCitation>
: Fig.
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus of Pterostichus (Pterostichus Koreonialoe) spp. from China 1 P. micropoides sp. nov., Holotype. 2 P. micropoides sp. nov., male, Paratype from Hunchun city, Jilin 3 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 4 P. syleus Kirschenhofer, a female from Zhuanghe city, Liaoning 5 P. quadrilobatus sp. nov., Holotype 6 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1063.69942.figures1-6" httpUri="https://binary.pensoft.net/fig/599856" pageId="0" pageNumber="1">2D</figureCitation>
) is different in the sclerotized part of male genitalia in the ventral tubercle a little larger and the apical portion of median lobe less deflected ventrally in lateral view. Thus, although the specimen described by
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. (2013)</bibRefCitation>
is exactly from the type locality of
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
, we thought it should belong to an unknown new species different from
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
and all other known species of
<taxonomicName authorityName="Park &amp; Kwon" authorityYear="1996" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Koreonialoe)" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Koreonialoe">
<emphasis italics="true" pageId="0" pageNumber="1">Pterostichus Koreonialoe</emphasis>
</taxonomicName>
. Besides the differences mentioned above, these two species are also different in their endophallus:
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="P. syleus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="syleus">
<emphasis italics="true" pageId="0" pageNumber="1">P. syleus</emphasis>
</taxonomicName>
has the endophallus more strongly directed ventro-basally, and lp is oblate, with apex not hooked (Figs
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">24</figureCitation>
,
<figureCitation captionStart="Figures 22–27" captionStartId="F5" captionText="Figures 22 - 27. Endophallus of Pterostichus (Koreonialoe) spp. from China, left lateral view and right lateral view 22 P. micropoides sp. nov., Paratype from Antu city, Jilin 23 P. micropoides sp. nov., Paratype from Changbaishan mt., Jilin 24 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 25 P. syleus Kirschenhofer, a male from Shedao Island, Liaoning 26 P. quadrilobatus sp. nov., Holotype 27 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1063.69942.figures22-27" httpUri="https://binary.pensoft.net/fig/599860" pageId="0" pageNumber="1">25</figureCitation>
); but the species by
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. (2013)</bibRefCitation>
has endophallus less directed ventral-basally, and lp is coniform, with distinctly hooked apex (
<bibRefCitation DOI="https://doi.org/10.1016/j.aspen.2013.05.003" author="Sasakawa, K" journalOrPublisher="Zoological Science" pageId="0" pageNumber="1" refId="B18" refString="Sasakawa, K, Kim, JK, Kim, JK, Kubota, K, 2013. Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16(4): 429-431. https://doi.org/10.1016/j.aspen.2013.05.003" title="Notes on the ground beetle Pterostichus togyusanus (Coleoptera: Carabidae): Phylogenetic evidence for species status and new distribution record. Journal of Asia-Pacific Entomology, 2013, 16 (4): 429 - 431." url="https://doi.org/10.1016/j.aspen.2013.05.003" year="2013">Sasakawa et al. 2013</bibRefCitation>
: Fig.
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus of Pterostichus (Pterostichus Koreonialoe) spp. from China 1 P. micropoides sp. nov., Holotype. 2 P. micropoides sp. nov., male, Paratype from Hunchun city, Jilin 3 P. syleus Kirschenhofer, a male from Fengcheng city, Liaoning 4 P. syleus Kirschenhofer, a female from Zhuanghe city, Liaoning 5 P. quadrilobatus sp. nov., Holotype 6 P. bellatrix Tschitscherine, a male from Changbai county, Jilin. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1063.69942.figures1-6" httpUri="https://binary.pensoft.net/fig/599856" pageId="0" pageNumber="1">2D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
Reference in a new issue View git blame Copy permalink