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<document id="420EB66486BA5150F18B79E62C406835" ID-DOI="10.5852/ejt.2024.946.2615" ID-ISSN="2118-9773" ID-Zenodo-Dep="13177205" ID-ZooBank="EFC377D4-50BD-43BD-A2E0-F2256AC38523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722591843161" checkinUser="plazi" docAuthor="Mbia, Donald-L’or Nyame, Ngon, Eric Belmond Biram à, Legendre, Frédéric &amp; Menbohan, Samuel Foto" docDate="2024" docId="03E8277D7E50FF7BFDB5AC26FBBDF9E7" docLanguage="en" docName="EJT.2024.946.1-57.pdf" docOrigin="European Journal of Taxonomy 946 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2615/11981" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Africalolampra Roth 1995" docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="FFD15F057E57FF7CFFFAAE35FFA1FFA8" masterDocTitle="Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995" masterLastPageNumber="57" masterPageNumber="1" pageNumber="8" updateTime="1722630795436" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4C64F93558216DFB97A58EB3D80A53BA">Semi-aquatic Epilamprinae cockroaches (Blattodea: Blaberidae) in Cameroon: towards a revision of continental African species of Rhabdoblatta Kirby, 1903 and Africalolampra Roth, 1995</mods:title>
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<mods:namePart id="49D896EFC6E95B787615618AFB5D842E">Mbia, Donald-L’or Nyame</mods:namePart>
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<mods:affiliation id="FC83695CCCE6FD2DF90BBAB056998112">Laboratory of Hydrobiology and Environment, Faculty of Science, P. O Box 812, Yaounde, Cameroon.</mods:affiliation>
<mods:nameIdentifier id="ACF381245F294A65E30105CE4DB63C3B" type="email">nyamedonal@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1C93B8D1DF0265BBF71ADD6B35F10C16">Ngon, Eric Belmond Biram à</mods:namePart>
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<mods:affiliation id="3B661D3F9BFAABACE5C4A126D5DA2F17">Laboratory of Hydrobiology and Environment, Faculty of Science, P. O Box 812, Yaounde, Cameroon. &amp; Institute of Geological and Mining Research, P. O Box 4110, Yaounde, Cameroon.</mods:affiliation>
<mods:nameIdentifier id="B359A763153729C7FF27E56164D5A267" type="email">birame.eric@yahoo.fr</mods:nameIdentifier>
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<mods:namePart id="C1F6833A653B824C3430FBBB82884649">Legendre, Frédéric</mods:namePart>
<mods:nameIdentifier id="30570BF31C557694316A9BD181BBF3FF" type="LSID">73B1BA6E-E129-4B14-A604-44B2220673C5</mods:nameIdentifier>
<mods:affiliation id="E70686C22B437062AC97E18DEFC65AAB">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="C9FEFBD535F4CDAE612FC108001328CB" type="email">frederic.legendre@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="A99761E9750C4C727A84C48C33E16575">Menbohan, Samuel Foto</mods:namePart>
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<mods:affiliation id="2450449C703E59F2E4F4508A65F2E0B8">Laboratory of Hydrobiology and Environment, Faculty of Science, P. O Box 812, Yaounde, Cameroon.</mods:affiliation>
<mods:nameIdentifier id="4D5E86DFDEAB4C24B767F2BD6BF40798" type="email">sfotomenbohan@yahoo.com</mods:nameIdentifier>
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<mods:date id="3741099F3502675749ABB0955694B9D5">2024</mods:date>
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<subSubSection id="C35BC5E07E50FF7BFDB5AC26FC44FD85" box="[591,997,530,557]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BFE966B7E50FF7BFDB5AC26FC44FD85" blockId="7.[591,997,530,557]" box="[591,997,530,557]" pageId="7" pageNumber="8">
<heading id="D0B621077E50FF7BFDB5AC26FC44FD85" box="[591,997,530,557]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
Genus
<taxonomicName id="4C41EDE87E50FF7BFD5BAC27FC44FD85" ID-CoL="8MPXF" authority="Roth, 1995" authorityName="Roth" authorityYear="1995" box="[673,997,530,557]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFD5BAC27FCFAFD85" bold="true" box="[673,859,530,557]" italics="true" pageId="7" pageNumber="8">Africalolampra</emphasis>
<bibRefCitation id="EFD0EB9A7E50FF7BFC98AC27FC44FD85" author="Roth L. M." box="[866,997,530,557]" pageId="7" pageNumber="8" pagination="89 - 98" refId="ref20469" refString="Roth L. M. 1995. Africalolampra ehrmanni new genus and species, and the male of Paraplecta parva Princis (Blattaria: Blaberidae). Psyche 102: 89 - 98. https: // doi. org / 10.1155 / 1995 / 49658" type="journal article" year="1995">Roth, 1995</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35BC5E07E50FF7BFF47AC61FDDAFD35" pageId="7" pageNumber="8" type="type_taxon">
<paragraph id="8BFE966B7E50FF7BFF47AC61FEF9FDC6" blockId="7.[189,344,596,622]" box="[189,344,596,622]" pageId="7" pageNumber="8">
<heading id="D0B621077E50FF7BFF47AC61FEF9FDC6" bold="true" box="[189,344,596,622]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9354A797E50FF7BFF47AC61FEF9FDC6" bold="true" box="[189,344,596,622]" pageId="7" pageNumber="8">
<typeStatus id="54FA28C97E50FF7BFF47AC61FF5BFDC6" box="[189,250,596,622]" pageId="7" pageNumber="8">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE966B7E50FF7BFF47ACB7FDDAFD35" blockId="7.[189,635,642,669]" box="[189,635,642,669]" pageId="7" pageNumber="8">
<taxonomicName id="4C41EDE87E50FF7BFF47ACB7FDD6FD35" ID-CoL="65F3V" authority="Roth, 1995" authorityName="Roth" authorityYear="1995" box="[189,631,642,669]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ehrmanni">
<emphasis id="B9354A797E50FF7BFF47ACB7FE48FD34" box="[189,489,642,668]" italics="true" pageId="7" pageNumber="8">Africalolampra ehrmanni</emphasis>
<bibRefCitation id="EFD0EB9A7E50FF7BFE0BACB7FDD6FD35" author="Roth L. M." box="[497,631,642,669]" pageId="7" pageNumber="8" pagination="89 - 98" refId="ref20469" refString="Roth L. M. 1995. Africalolampra ehrmanni new genus and species, and the male of Paraplecta parva Princis (Blattaria: Blaberidae). Psyche 102: 89 - 98. https: // doi. org / 10.1155 / 1995 / 49658" type="journal article" year="1995">Roth, 1995</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35BC5E07E50FF7BFF47ACF1FBBDF9E7" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BFE966B7E50FF7BFF47ACF1FE6DFD76" blockId="7.[189,460,708,734]" box="[189,460,708,734]" pageId="7" pageNumber="8">
<heading id="D0B621077E50FF7BFF47ACF1FE6DFD76" bold="true" box="[189,460,708,734]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9354A797E50FF7BFF47ACF1FE6DFD76" bold="true" box="[189,460,708,734]" pageId="7" pageNumber="8">Diagnosis and history</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE966B7E50FF7BFF47ACC7FBE1FB51" blockId="7.[188,1399,754,1274]" pageId="7" pageNumber="8">
The genus
<taxonomicName id="4C41EDE87E50FF7BFEC0ACC7FE4EFCA4" authorityName="Roth" authorityYear="1995" box="[314,495,754,780]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFEC0ACC7FE4EFCA4" box="[314,495,754,780]" italics="true" pageId="7" pageNumber="8">Africalolampra</emphasis>
</taxonomicName>
was erected to accommodate the new species
<taxonomicName id="4C41EDE87E50FF7BFBFEACC7FF5BFC98" authority="Roth, 1995" authorityName="Roth" authorityYear="1995" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ehrmanni">
<emphasis id="B9354A797E50FF7BFBFEACC7FA8FFCA4" box="[1028,1326,754,780]" italics="true" pageId="7" pageNumber="8">Africalolampra ehrmanni</emphasis>
<bibRefCitation id="EFD0EB9A7E50FF7BFACCACC7FF5BFC98" author="Roth L. M." pageId="7" pageNumber="8" pagination="89 - 98" refId="ref20469" refString="Roth L. M. 1995. Africalolampra ehrmanni new genus and species, and the male of Paraplecta parva Princis (Blattaria: Blaberidae). Psyche 102: 89 - 98. https: // doi. org / 10.1155 / 1995 / 49658" type="journal article" year="1995">Roth, 1995</bibRefCitation>
</taxonomicName>
. Later,
<bibRefCitation id="EFD0EB9A7E50FF7BFEAEAD20FD8EFC87" author="Anisyutkin L. N." box="[340,559,789,815]" pageId="7" pageNumber="8" pagination="139 - 152" refId="ref17486" refString="Anisyutkin L. N. 2016. New and little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Museum d'histoire naturelle de Geneve and the Zoological Institute RAS, Saint Petersburg. Part 2. Revue suisse de Zoologie 123 (1): 139 - 152. https: // doi. org / 10.5281 / zenodo. 46293" type="journal article" year="2016">Anisyutkin (2016)</bibRefCitation>
transferred two species of
<taxonomicName id="4C41EDE87E50FF7BFC81AD20FBBAFC87" authorityName="Kirby" authorityYear="1903" box="[891,1051,789,815]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFC81AD20FBBAFC87" box="[891,1051,789,815]" italics="true" pageId="7" pageNumber="8">Rhabdoblatta</emphasis>
</taxonomicName>
to this genus:
<taxonomicName id="4C41EDE87E50FF7BFB2BAD23FAD7FC87" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" box="[1233,1398,789,816]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="erubescens">
<emphasis id="B9354A797E50FF7BFB2BAD23FAD7FC87" box="[1233,1398,789,816]" italics="true" pageId="7" pageNumber="8">A. erubescens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C41EDE87E50FF7BFF0BAD0CFE08FCFB" baseAuthorityName="Saussure" baseAuthorityYear="1895" box="[241,425,825,851]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B9354A797E50FF7BFF0BAD0CFE08FCFB" box="[241,425,825,851]" italics="true" pageId="7" pageNumber="8">A. punctipennis</emphasis>
</taxonomicName>
. According to
<bibRefCitation id="EFD0EB9A7E50FF7BFDA6AD0CFD4EFCFC" author="Roth L. M." box="[604,751,825,852]" pageId="7" pageNumber="8" pagination="89 - 98" refId="ref20469" refString="Roth L. M. 1995. Africalolampra ehrmanni new genus and species, and the male of Paraplecta parva Princis (Blattaria: Blaberidae). Psyche 102: 89 - 98. https: // doi. org / 10.1155 / 1995 / 49658" type="journal article" year="1995">Roth (1995)</bibRefCitation>
, the following combination of characters is diagnostic of the genus
<taxonomicName id="4C41EDE87E50FF7BFEA7AD69FDB3FCDE" authorityName="Roth" authorityYear="1995" box="[349,530,860,886]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFEA7AD69FDB3FCDE" box="[349,530,860,886]" italics="true" pageId="7" pageNumber="8">Africalolampra</emphasis>
</taxonomicName>
: weak sexual dimorphism (tegmina and wings completely developed in both sexes – although this is not true for
<taxonomicName id="4C41EDE87E50FF7BFD45ADB5FCDBFC31" baseAuthorityName="Saussure" baseAuthorityYear="1895" box="[703,890,896,922]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B9354A797E50FF7BFD45ADB5FCDBFC31" box="[703,890,896,922]" italics="true" pageId="7" pageNumber="8">A. punctipennis</emphasis>
</taxonomicName>
), metatarsus with 2 rows of spines along lower margin, tarsal claws distinctly serrated, first abdominal tergite of male with medial specialization (this is not true either for
<taxonomicName id="4C41EDE87E50FF7BFE00ADF3FD15FC77" baseAuthorityName="Saussure" baseAuthorityYear="1895" box="[506,692,966,992]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B9354A797E50FF7BFE00ADF3FD15FC77" box="[506,692,966,992]" italics="true" pageId="7" pageNumber="8">A. punctipennis</emphasis>
</taxonomicName>
), left stylus absent. This combination of characters is also found in two species from continental Africa belonging to the genus
<taxonomicName id="4C41EDE87E50FF7BFC20ADDCFBDBFBAB" authorityName="Kirby" authorityYear="1903" box="[986,1146,1001,1027]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFC20ADDCFBDBFBAB" box="[986,1146,1001,1027]" italics="true" pageId="7" pageNumber="8">Rhabdoblatta</emphasis>
</taxonomicName>
, and we thus propose the following new combinations:
<taxonomicName id="4C41EDE87E50FF7BFDAFAA39FC14FB8E" authorityName="Mbia &amp; Ngon &amp; Legendre &amp; Menbohan" authorityYear="2024" baseAuthorityName="Borg" baseAuthorityYear="1902" box="[597,949,1036,1062]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="camerunensis">
<emphasis id="B9354A797E50FF7BFDAFAA39FC14FB8E" box="[597,949,1036,1062]" italics="true" pageId="7" pageNumber="8">Africalolampra camerunensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C41EDE87E50FF7BFC0FAA39FAA4FB8E" authorityName="Mbia &amp; Ngon &amp; Legendre &amp; Menbohan" authorityYear="2024" baseAuthorityName="Walker" baseAuthorityYear="1868" box="[1013,1285,1036,1062]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="stipata">
<emphasis id="B9354A797E50FF7BFC0FAA39FAA4FB8E" box="[1013,1285,1036,1062]" italics="true" pageId="7" pageNumber="8">Africalolampra stipata</emphasis>
</taxonomicName>
. We also reinstate
<taxonomicName id="4C41EDE87E50FF7BFEDDAA1AFDC5FBE1" baseAuthorityName="Saussure" baseAuthorityYear="1895" box="[295,612,1071,1097]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B9354A797E50FF7BFEDDAA1AFDC5FBE1" box="[295,612,1071,1097]" italics="true" pageId="7" pageNumber="8">Rhabdoblatta punctipennis</emphasis>
</taxonomicName>
because it does not share the apomorphies of
<taxonomicName id="4C41EDE87E50FF7BFB80AA1AFA8EFBE1" authorityName="Roth" authorityYear="1995" box="[1146,1327,1071,1097]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFB80AA1AFA8EFBE1" box="[1146,1327,1071,1097]" italics="true" pageId="7" pageNumber="8">Africalolampra</emphasis>
</taxonomicName>
. Note that
<taxonomicName id="4C41EDE87E50FF7BFF08AA66FDF3FBC4" authority="(Gerstaecker, 1883)" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" box="[242,594,1106,1132]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lyncea">
<emphasis id="B9354A797E50FF7BFF08AA66FEC1FBC4" box="[242,352,1106,1132]" italics="true" pageId="7" pageNumber="8">R. lyncea</emphasis>
(
<bibRefCitation id="EFD0EB9A7E50FF7BFE8BAA67FDEBFBC4" author="Gerstaecker A." box="[369,586,1106,1132]" pageId="7" pageNumber="8" pagination="39 - 102" refId="ref18580" refString="Gerstaecker A. 1883. Beitrag zur Kenntniss der Orthopteren-Fauna Guinea's: nach den von R. Buchholz wahrend der Jahre 1872 bis 1875 daselbst gesammelten Arten. Mittheilungen aus dem naturwissenschaftlichen Verein fur Neu-Vorpommern und Rugen 14: 39 - 102. Available from https: // www. biodiversitylibrary. org / page / 55843440 [accessed 4 Jun. 2023]." type="journal article" year="1883">Gerstaecker, 1883</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C41EDE87E50FF7BFD77AA66FC03FBC4" authority="(Borg, 1902)" baseAuthorityName="Borg" baseAuthorityYear="1902" box="[653,930,1106,1132]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9354A797E50FF7BFD77AA66FCA3FBC4" box="[653,770,1106,1132]" italics="true" pageId="7" pageNumber="8">R. pallida</emphasis>
(
<bibRefCitation id="EFD0EB9A7E50FF7BFCE9AA67FC3BFBC4" author="Borg H." box="[787,922,1106,1132]" pageId="7" pageNumber="8" pagination="1 - 36" refId="ref17912" refString="Borg H. 1902. Blattodeen aus Kamerun. Bidrag till svenska Vetenskaps-Akademiens Handlingar 28: 1 - 36. Available from https: // www. biodiversitylibrary. org / page / 14210018 [accessed 4 Jun. 2023]." type="journal article" year="1902">Borg, 1902</bibRefCitation>
)
</taxonomicName>
also have a tergal specialization but on the 5
<superScript id="7C343B237E50FF7BFF03AA40FEA7FB2B" attach="left" box="[249,262,1141,1155]" fontSize="6" pageId="7" pageNumber="8">th</superScript>
and at the junction of the 8–9
<superScript id="7C343B237E50FF7BFD8AAA40FDDCFB2B" attach="left" box="[624,637,1141,1155]" fontSize="6" pageId="7" pageNumber="8">th</superScript>
abdominal tergites, respectively.
<taxonomicName id="4C41EDE87E50FF7BFBF5AA40FAA4FB27" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" box="[1039,1285,1141,1167]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lyncea">
<emphasis id="B9354A797E50FF7BFBF5AA40FAA4FB27" box="[1039,1285,1141,1167]" italics="true" pageId="7" pageNumber="8">Rhabdoblatta lyncea</emphasis>
</taxonomicName>
may also have slightly serrated claws (
<bibRefCitation id="EFD0EB9A7E50FF7BFDE9AAACFD08FB1C" author="Kumar R." box="[531,681,1177,1204]" pageId="7" pageNumber="8" pagination="27 - 121" refId="ref19321" refString="Kumar R. 1975. A review of the cockroaches of West Africa and the Congo Basin (Dictyoptera: Blattaria). Bulletin de Institut fondamental d'Afrique noire, serie A 37: 27 - 121." type="journal article" year="1975">Kumar 1975</bibRefCitation>
) but two styli, whereas
<taxonomicName id="4C41EDE87E50FF7BFC3AAAAFFB95FB1B" baseAuthorityName="Borg" baseAuthorityYear="1902" box="[960,1076,1177,1203]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9354A797E50FF7BFC3AAAAFFB95FB1B" box="[960,1076,1177,1203]" italics="true" pageId="7" pageNumber="8">R. pallida</emphasis>
</taxonomicName>
has a single stylus. Finally, Malagasy species are reported but males of some species have not been observed, so we do not dwell further on their generic affiliation, which would require further examination.
</paragraph>
<paragraph id="8BFE966B7E50FF7BFF47AB15FE3FFAF6" blockId="7.[189,1398,1312,1374]" pageId="7" pageNumber="8">
Below is the updated diagnosis of the genus
<taxonomicName id="4C41EDE87E50FF7BFD20AB15FC2CFA92" authorityName="Roth" authorityYear="1995" box="[730,909,1312,1338]" class="Insecta" family="Blaberidae" genus="Africalolampra" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9354A797E50FF7BFD20AB15FC2CFA92" box="[730,909,1312,1338]" italics="true" pageId="7" pageNumber="8">Africalolampra</emphasis>
</taxonomicName>
, slightly modified from
<bibRefCitation id="EFD0EB9A7E50FF7BFB48AB15FAE2FA93" author="Roth L. M." box="[1202,1347,1312,1339]" pageId="7" pageNumber="8" pagination="89 - 98" refId="ref20469" refString="Roth L. M. 1995. Africalolampra ehrmanni new genus and species, and the male of Paraplecta parva Princis (Blattaria: Blaberidae). Psyche 102: 89 - 98. https: // doi. org / 10.1155 / 1995 / 49658" type="journal article" year="1995">Roth (1995)</bibRefCitation>
and
<bibRefCitation id="EFD0EB9A7E50FF7BFF47AB71FE3BFAF6" author="Anisyutkin L. N." box="[189,410,1348,1374]" pageId="7" pageNumber="8" pagination="139 - 152" refId="ref17486" refString="Anisyutkin L. N. 2016. New and little known Epilamprinae (Dictyoptera: Blaberidae) from the collections of the Museum d'histoire naturelle de Geneve and the Zoological Institute RAS, Saint Petersburg. Part 2. Revue suisse de Zoologie 123 (1): 139 - 152. https: // doi. org / 10.5281 / zenodo. 46293" type="journal article" year="2016">Anisyutkin (2016)</bibRefCitation>
:
</paragraph>
<paragraph id="8BFE966B7E50FF7BFF47ABB0FBBDF9E7" blockId="7.[187,1399,1413,1616]" pageId="7" pageNumber="8">
Weak sexual dimorphism, with tegmina and wings fully developed in both sexes. Anteroventral margin of front femur
<typeStatus id="54FA28C97E50FF7BFE90AB9CFE3CFA6B" box="[362,413,1449,1475]" pageId="7" pageNumber="8">type</typeStatus>
B2; genicular spine absent from fore femur, present on mid and hind femora; spines present on antero- and posteroventral margins of all femora; pulvilli on four proximal tarsomeres; ventral margin of first tarsal segments with two rows of spines; tarsal claws symmetrical and serrated, arolia well-developed. Male with glandular specialization on first abdominal tergite; subgenital plate weakly asymmetrical, with only a single, small, right style (i.e., left style absent).
</paragraph>
</subSubSection>
</treatment>
</document>
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