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<document id="CC8282BA337A1D1CCF28F52FD2DC0E88" ID-CLB-Dataset="24776" ID-DOI="10.11646/zootaxa.4701.4.2" ID-GBIF-Dataset="3f56c3ca-c520-4e2c-9310-95178c282c9d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3558175" ID-ZooBank="6417DB61-737F-443E-B91B-CF1723A891E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575044514958" checkinUser="plazi" docAuthor="Schmidt, Joachim, Scholz, Stefan &amp; Kavanaugh, David H." docDate="2019" docId="E86787E1FFC0FFFDFF39D3E1FCFBFBBA" docLanguage="en" docName="zootaxa.4701.4.2.pdf" docOrigin="Zootaxa 4701 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaeonebria inexspectata Schmidt &amp; Scholz &amp; Kavanaugh 2019, sp. n." docType="treatment" docVersion="9" lastPageNumber="365" masterDocId="145EFF99FFC3FFF2FFAED24FFFF1FFBC" masterDocTitle="Unexpected findings in the Eocene Baltic amber forests: Ground beetle fossils of the tribe Nebriini (Coleoptera: Carabidae)" masterLastPageNumber="370" masterPageNumber="350" pageNumber="353" updateTime="1698769972962" updateUser="ExternalLinkService">
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<mods:title id="1B1825B802619BFA8AD4C2787D856785">Unexpected findings in the Eocene Baltic amber forests: Ground beetle fossils of the tribe Nebriini (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="26F391437F61EF24B8E7B7AC90633286">Schmidt, Joachim</mods:namePart>
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<mods:namePart id="86D3E08667936388EA12E09214781CC9">Scholz, Stefan</mods:namePart>
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<mods:namePart id="34666E1B6FCCCD307B40D42C40FB94B6">Kavanaugh, David H.</mods:namePart>
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<mods:typeOfResource id="DF7E7A1BD11D345FF0304AC43736E462">text</mods:typeOfResource>
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<mods:title id="59CAFCA9FC06AFAA79A196D9BF3EDEFB">Zootaxa</mods:title>
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<mods:date id="309305907613B222D1C8F9AFEACB3B54">2019</mods:date>
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<mods:number id="306575B40AC6B61F55233440A2B9AECB">2019-11-29</mods:number>
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<treatment id="E86787E1FFC0FFFDFF39D3E1FCFBFBBA" ID-DOI="http://doi.org/10.5281/zenodo.5617574" ID-GBIF-Taxon="160605707" ID-Zenodo-Dep="5617574" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA" httpUri="http://treatment.plazi.org/id/E86787E1FFC0FFFDFF39D3E1FCFBFBBA" lastPageId="15" lastPageNumber="365" pageId="3" pageNumber="353">
<subSubSection id="28D4657CFFC0FFF1FF39D3E1FC93FE75" box="[151,866,430,457]" pageId="3" pageNumber="353" type="nomenclature">
<paragraph id="607136F7FFC0FFF1FF39D3E1FC93FE75" blockId="3.[151,866,430,457]" box="[151,866,430,457]" pageId="3" pageNumber="353">
<heading id="3B39819BFFC0FFF1FF39D3E1FC93FE75" bold="true" box="[151,866,430,457]" fontSize="11" level="1" pageId="3" pageNumber="353" reason="1">
<emphasis id="52BAEAE5FFC0FFF1FF39D3E1FC93FE75" bold="true" box="[151,866,430,457]" pageId="3" pageNumber="353">
<taxonomicName id="A7CE4D74FFC0FFF1FF39D3E1FCE2FE75" ID-CoL="8MFJR" authority="Schmidt &amp; Kavanaugh" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[151,787,430,457]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="species" species="inexspectata" status="sp. nov.">
<emphasis id="52BAEAE5FFC0FFF1FF39D3E1FE18FE74" bold="true" box="[151,489,430,457]" italics="true" pageId="3" pageNumber="353">Archaeonebria inexspectata</emphasis>
Schmidt &amp; Kavanaugh
</taxonomicName>
,
<taxonomicNameLabel id="4989579EFFC0FFF1FC8ED3E0FC93FE75" box="[800,866,431,457]" pageId="3" pageNumber="353" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28D4657CFFC0FFF1FF39D3B9FF2BF868" pageId="3" pageNumber="353" type="materials_examined">
<paragraph id="607136F7FFC0FFF1FF39D3B9FCB8FD79" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
<emphasis id="52BAEAE5FFC0FFF1FF39D3B9FEB5FDAC" bold="true" box="[151,324,501,528]" pageId="3" pageNumber="353">Type material.</emphasis>
<materialsCitation id="D0A63CAAFFC0FFF1FEE4D3B9FCB8FD79" ID-GBIF-Occurrence="2460219638" collectingDate="1935-10-20" collectionCode="GPIH" collectorName="Coll. Carsten Grohn &amp; CeNak &amp; Grohn" county="Male in Baltic" location="University" municipality="Geological-Palaeontological Institute" pageId="3" pageNumber="353" specimenCount="1" stateProvince="Hamburg" typeStatus="holotype">
<emphasis id="52BAEAE5FFC0FFF1FEE4D3B9FE31FDAC" bold="true" box="[330,448,502,528]" pageId="3" pageNumber="353">
<typeStatus id="BF758855FFC0FFF1FEE4D3B9FE4AFDAC" box="[330,443,502,528]" pageId="3" pageNumber="353" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCounty id="89104E7BFFC0FFF1FE68D3B9FD95FDAC" box="[454,612,502,528]" pageId="3" pageNumber="353">Male in Baltic</collectingCounty>
amber;
<collectorName id="CD3B5321FFC0FFF1FD13D3B9FC6FFDAC" box="[701,926,502,528]" pageId="3" pageNumber="353">Coll. Carsten Gröhn</collectorName>
in the collection of the
<collectingMunicipality id="8015AC8DFFC0FFF1FB0ED3B9FEB6FD88" pageId="3" pageNumber="353">Geological-Palaeontological Institute</collectingMunicipality>
of the
<location id="6511602CFFC0FFF1FE3BD055FDFAFD88" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FE3BD055FDFAFD88" box="[405,523,538,564]" county="Male in Baltic" municipality="Geological-Palaeontological Institute" name="University" pageId="3" pageNumber="353" stateProvince="Hamburg">University</location>
of
<collectingRegion id="A20AF815FFC0FFF1FD9FD055FD6BFD88" box="[561,666,538,564]" country="Germany" name="Hamburg" pageId="3" pageNumber="353">Hamburg</collectingRegion>
(
<collectionCode id="06DFAE32FFC0FFF1FD07D055FD01FD88" box="[681,752,538,564]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="353">GPIH</collectionCode>
), now in the
<collectingRegion id="A20AF815FFC0FFF1FC2FD055FC12FD88" box="[897,995,538,564]" country="France" name="Centre" pageId="3" pageNumber="353">Centrum</collectingRegion>
of
<location id="6511602CFFC0FFF1FBA7D055FB48FD88" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FBA7D055FB48FD88" box="[1033,1209,538,564]" county="Male in Baltic" municipality="Geological-Palaeontological Institute" name="Natural History" pageId="3" pageNumber="353" stateProvince="Hamburg">Natural History</location>
,
<collectingRegion id="A20AF815FFC0FFF1FB6AD055FADCFD88" box="[1220,1325,538,564]" country="Germany" name="Hamburg" pageId="3" pageNumber="353">Hamburg</collectingRegion>
(
<collectorName id="CD3B5321FFC0FFF1FA93D055FA60FD89" box="[1341,1425,538,565]" pageId="3" pageNumber="353">CeNak</collectorName>
), with collection number „
<collectionCode id="06DFAE32FFC0FFF1FE05D071FE1EFDE4" box="[427,495,574,600]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="353">GPIH</collectionCode>
no. 4980, coll.
<collectorName id="CD3B5321FFC0FFF1FD30D071FD17FDE4" box="[670,742,574,600]" pageId="3" pageNumber="353">Gröhn</collectorName>
no. 7919”.
<location id="6511602CFFC0FFF1FCC6D070FC7EFDE5" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FCC6D070FC7EFDE5" box="[872,911,575,601]" county="Male in Baltic" municipality="Geological-Palaeontological Institute" name="For" pageId="3" pageNumber="353" stateProvince="Hamburg">For</location>
better investigation of the fossil, the stone was cut in two pieces, with one piece bearing the
<taxonomicName id="A7CE4D74FFC0FFF1FD34D02DFD06FDC0" authorityName="Laporte de Castelnau" authorityYear="1834" box="[666,759,610,636]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Nebriini">Nebriini</taxonomicName>
fossil, and has a size of approx.
<collectingDate id="0434E9DFFFC0FFF1FBC3D02DFB10FDC1" box="[1133,1249,610,637]" isEnumeration="true" pageId="3" pageNumber="353">
<date id="14701037FFC0FFF1FBC3D02DFB44FDC0" box="[1133,1205,610,637]" pageId="3" pageNumber="353" value="1924-10-06">24 x 6</date>
<date id="14701037FFC0FFF1FB13D02CFB10FDC1" box="[1213,1249,611,637]" pageId="3" pageNumber="353">
x
<quantity id="A7369B12FFC0FFF1FB7DD02CFB10FDC1" box="[1235,1249,611,637]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="353" unit="mm" value="7.0">7</quantity>
</date>
</collectingDate>
mm (irregularly cut) (= „
<collectionCode id="06DFAE32FFC0FFF1FF5DD0C9FEC6FD1C" box="[243,311,646,672]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="353">GPIH</collectionCode>
no. 4980, coll.
<collectorName id="CD3B5321FFC0FFF1FE4AD0C9FDDDFD1C" box="[484,556,646,672]" pageId="3" pageNumber="353">Gröhn</collectorName>
no. 7919a”,
<figureCitation id="F8F52A72FFC0FFF1FD14D0C9FD0BFD1C" box="[698,762,646,672]" captionStart="FIGURES 13" captionStartId="4.[151,264,1924,1949]" captionTargetBox="[151,1284,174,1905]" captionTargetId="figure@4.[151,1284,166,1905]" captionTargetPageId="4" captionText="FIGURES 13. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the holotype. Fig. 1, general view with contours of the amber piece. Fig. 2, right lateral aspect of the embedded specimen. Fig. 3, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.3558177" httpUri="https://zenodo.org/record/3558177/files/figure.png" pageId="3" pageNumber="353">Fig. 1</figureCitation>
); the remaining piece has a size of approx.
<date id="14701037FFC0FFF1FB4ED0CAFAC2FD1C" box="[1248,1331,645,673]" pageId="3" pageNumber="353" value="1935-10-20">
<collectingDate id="0434E9DFFFC0FFF1FB4ED0CAFAC2FD1C" box="[1248,1331,645,673]" pageId="3" pageNumber="353" value="1935-10-20">35 x 20</collectingDate>
</date>
x
<quantity id="A7369B12FFC0FFF1FAE0D0C9FA6CFD1D" box="[1358,1437,646,673]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="353" unit="mm" value="16.0">16 mm</quantity>
(irregularly cut) (= „
<collectionCode id="06DFAE32FFC0FFF1FED4D0E5FE4FFD78" box="[378,446,682,708]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="353">GPIH</collectionCode>
no. 4980, coll.
<collectorName id="CD3B5321FFC0FFF1FDC3D0E5FD44FD78" box="[621,693,682,708]" pageId="3" pageNumber="353">Gröhn</collectorName>
no. 7919b”).
</materialsCitation>
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D081FB19FCC4" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
Preservation status: The amber is clear but pervaded by several flowlines and partly covered by very fine surface cracks; the embedded
<taxonomicName id="A7CE4D74FFC0FFF1FE6DD0BDFDD1FCB0" authorityName="Laporte de Castelnau" authorityYear="1834" box="[451,544,754,780]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Nebriini">Nebriini</taxonomicName>
fossil is extensively covered by flowlines and bubbles adjoining the beetle body (
<figureCitation id="F8F52A72FFC0FFF1FF31D159FEF3FC8C" box="[159,258,790,816]" captionStart="FIGURES 13" captionStartId="4.[151,264,1924,1949]" captionTargetBox="[151,1284,174,1905]" captionTargetId="figure@4.[151,1284,166,1905]" captionTargetPageId="4" captionText="FIGURES 13. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the holotype. Fig. 1, general view with contours of the amber piece. Fig. 2, right lateral aspect of the embedded specimen. Fig. 3, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.3558177" httpUri="https://zenodo.org/record/3558177/files/figure.png" pageId="3" pageNumber="353">Figs 23</figureCitation>
) and thus, hardly visible using light microscope. The exoskeleton of the specimen and its negative imprint on the inclusion wall is well preserved and could therefore be imaged in detail using micro-CT (
<figureCitation id="F8F52A72FFC0FFF1FB78D175FAB2FCE8" box="[1238,1347,826,852]" captionStart="FIGURES 46" captionStartId="5.[151,264,1924,1949]" captionTargetBox="[152,1234,167,1903]" captionTargetId="figure@5.[151,1234,166,1904]" captionTargetPageId="5" captionText="FIGURES 46. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the holotype. Fig. 4, dorsal aspect. Fig. 5, right lateral aspect. Fig. 6, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.3558179" httpUri="https://zenodo.org/record/3558179/files/figure.png" pageId="3" pageNumber="353">Figs. 46</figureCitation>
,
<figureCitation id="F8F52A72FFC0FFF1FAE1D175FF47FCC4" captionStart="FIGURES 3136" captionStartId="13.[151,264,1667,1692]" captionTargetBox="[151,1436,166,1602]" captionTargetId="figure@13.[151,1436,166,1603]" captionTargetPageId="13" captionText="FIGURES 3136. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the holotype (Figs 31, 32, 34) and paratype #2 (Fig. 33, 35, 36). Fig. 31, head, dorsal aspect; the arrows point to the insertions of the labral setae, clypeal seta, and supraorbital seta. Remark: Note the deformation of the labral apical margin with respect to its negative imprint on the inclusion wall, which represents its original state. Fig. 32, head, ventral aspect; the arrows point to the insertions of the setae on mentum and submentum. Figs 33, 34, mouthparts, apicoventral aspect of different specimens using different greyscale thresholds. Figs 35, 36, separated images of the ligula with the completely fused paraglossae (Fig. 35, ventral aspect; Fig. 36, apicoventral aspect). Abbreviations: ant = antenna; dp = dust particles; la = lacinia; lig = ligula; lig-a = apex of ligula; lp-l = left labial palpus; lp-r = right labial palpus; m = mentum; md-l = left mandible; md-r = right mandible; mp-l = left maxillary palpus; mp-r = right maxillary palpus; mx = maxilla; n = negative imprint of the labral margin; p = shrunken remains of the labral margin; pgl = paraglossa." figureDoi="http://doi.org/10.5281/zenodo.3558195" httpUri="https://zenodo.org/record/3558195/files/figure.png" pageId="3" pageNumber="353">31, 32, 34</figureCitation>
), as well as the aedeagal median lobe, except for the weekly sclerotized structures (
<figureCitation id="F8F52A72FFC0FFF1FBF3D111FB2DFCC4" box="[1117,1244,862,888]" captionStart="FIGURES 4253" captionStartId="16.[151,264,1641,1666]" captionTargetBox="[290,1297,168,1620]" captionTargetId="figure@16.[289,1298,166,1621]" captionTargetPageId="16" captionText="FIGURES 4253. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the aedeagus of the holotype using different grey scale thresholds (Figs 4249), and of the female ovipositor of paratype #2 (Figs 5053; the ovipositor was separated by the segmentation function of Amira software in Figs 5153). Figs 42, 45, left lateral view. Figs 43, 46, view slightly diagonal from behind, frontally to the apical lamella. Figs 44, 47, right lateral view. Fig. 48, frontal view towards median lobe base. Fig. 49, view from posterad towards median lobe apex. Fig. 50, section of the female abdomen in ventral view. Fig. 51, ovipositor in ventral view. Fig. 52, postero-left lateroventral view. Fig. 53, dorsal view.Abbreviations: al = apical lamella of aedeagal median lobe; ap = lanceolate apophysis of left paramere; ds = dorsal row of spiniform setae on the edge of the fused apical and basal gonocoxites; g = gut; gc 1 = basal gonocoxite; gc 2 = apical gonocoxite; ltg = laterotergite; mtg = mediotergite; o-a = extreme apex of apical ostium of median lobe; o-b = extreme base of apical ostium; pm-l = left paramere; pm-r = right paramere; se = sagittal aileron; sp = sensory pit; vs = ventral row of spiniform setae (only shown on right basal gonocoxite);?vs = the pore marks the insertion of a probable additional spiniform seta of the ventral series." figureDoi="http://doi.org/10.5281/zenodo.3558199" httpUri="https://zenodo.org/record/3558199/files/figure.png" pageId="3" pageNumber="353">Figs 42-49</figureCitation>
).
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D1CDFBEBFC7C" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
Syninclusions: One Nematocera fly and a dirt particle in the „
<collectionCode id="06DFAE32FFC0FFF1FCD9D1CDFC4AFC20" box="[887,955,898,924]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="353">GPIH</collectionCode>
no. 4980, coll. Gröhn no. 7919a” piece, an additional Nematocera fly in the „
<collectionCode id="06DFAE32FFC0FFF1FDBCD1E9FDA7FC7C" box="[530,598,934,960]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="353">GPIH</collectionCode>
no. 4980, coll. Gröhn no. 7919b” piece.
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D185FE5AFB91" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
<materialsCitation id="D0A63CAAFFC0FFF1FF69D185FE56FB91" ID-GBIF-Occurrence="2460219635" collectingDate="1920-10-31" collectionCode="CA" county="Female in Baltic" location="Museum of Palaeontology" municipality="University" pageId="3" pageNumber="353" specimenCode="UCMP 403984.01" specimenCount="1" stateProvince="California" typeStatus="paratype">
<emphasis id="52BAEAE5FFC0FFF1FF69D185FE92FC58" bold="true" box="[199,355,969,996]" pageId="3" pageNumber="353">
<typeStatus id="BF758855FFC0FFF1FF69D185FEC5FC58" box="[199,308,970,996]" pageId="3" pageNumber="353" type="paratype">Paratype</typeStatus>
#1:
</emphasis>
<collectingCounty id="89104E7BFFC0FFF1FEC2D185FDDBFC58" box="[364,554,970,996]" pageId="3" pageNumber="353">Female in Baltic</collectingCounty>
amber; deposited in the
<collectingMunicipality id="8015AC8DFFC0FFF1FCE7D185FC4EFC58" box="[841,959,970,996]" pageId="3" pageNumber="353">University</collectingMunicipality>
of
<collectingRegion id="A20AF815FFC0FFF1FC47D185FBACFC58" box="[1001,1117,970,996]" country="United States of America" name="California" pageId="3" pageNumber="353">California</collectingRegion>
,
<location id="6511602CFFC0FFF1FBC5D184FA69FC58" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FBC5D184FA69FC58" box="[1131,1432,970,997]" county="Female in Baltic" municipality="University" name="Museum of Palaeontology" pageId="3" pageNumber="353" stateProvince="California">Museum of Palaeontology</location>
,
<location id="6511602CFFC0FFF1FF39D1A1FF0CFBB4" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FF39D1A1FF0CFBB4" box="[151,253,1006,1032]" county="Female in Baltic" municipality="University" name="Berkeley" pageId="3" pageNumber="353" stateProvince="California">Berkeley</location>
,
<collectionCode id="06DFAE32FFC0FFF1FEA7D1A1FEC2FBB4" box="[265,307,1006,1032]" country="USA" name="Chicago Academy of Sciences" pageId="3" pageNumber="353" type="Museum">CA</collectionCode>
, with specimen number
<specimenCode id="30689E8CFFC0FFF1FDE6D1A1FCE9FBB4" box="[584,792,1006,1032]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="353" type="University or college">UCMP 403984.01</specimenCode>
, and locality number IP 15208 (
<figureCitation id="F8F52A72FFC0FFF1FB2AD1A1FB38FBB4" box="[1156,1225,1006,1032]" captionStart="FIGURES 710" captionStartId="6.[151,264,1876,1901]" captionTargetBox="[252,1329,166,1856]" captionTargetId="figure@6.[252,1329,166,1856]" captionTargetPageId="6" captionText="FIGURES 710. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #1. Fig. 7, dorsal aspect of the embedded specimen. Fig. 8, ventral aspect. Fig. 9, general view with contours of the amber piece, showing several embedded insects (see text for details); the arrow points to the position of the embedded Archaeonebria specimen. Fig. 10, basal part of the left hindleg; the arrows point to the positions of the anterior and posterior metacoxal setae. Abbreviations: mcx = metacoxa; mtr = metatrochanter; w-l = left hindwing; w-r = right hindwing." figureDoi="http://doi.org/10.5281/zenodo.3558181" httpUri="https://zenodo.org/record/3558181/files/figure.png" pageId="3" pageNumber="353">Fig. 9</figureCitation>
). Size of the piece approx.
<date id="14701037FFC0FFF1FF5DD65DFEB6FB90" box="[243,327,1042,1069]" pageId="3" pageNumber="353" value="1920-10-31">
<collectingDate id="0434E9DFFFC0FFF1FF5DD65DFEB6FB90" box="[243,327,1042,1069]" pageId="3" pageNumber="353" value="1920-10-31">31 x 20</collectingDate>
</date>
x
<quantity id="A7369B12FFC0FFF1FECDD65DFE56FB91" box="[355,423,1042,1069]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="353" unit="mm" value="8.0">8 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D679FB20FB5C" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
Preservation status: The amber is clear, pervaded by very few flowlines, partly covered by very fine surface cracks; the embedded
<taxonomicName id="A7CE4D74FFC0FFF1FE39D615FE05FBC8" authorityName="Laporte de Castelnau" authorityYear="1834" box="[407,500,1114,1140]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Nebriini">Nebriini</taxonomicName>
fossil is partly covered by milky coating and a small bubble adjoining the dorsal surface of the beetle body (
<figureCitation id="F8F52A72FFC0FFF1FE6AD631FDF9FB25" box="[452,520,1150,1177]" captionStart="FIGURES 710" captionStartId="6.[151,264,1876,1901]" captionTargetBox="[252,1329,166,1856]" captionTargetId="figure@6.[252,1329,166,1856]" captionTargetPageId="6" captionText="FIGURES 710. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #1. Fig. 7, dorsal aspect of the embedded specimen. Fig. 8, ventral aspect. Fig. 9, general view with contours of the amber piece, showing several embedded insects (see text for details); the arrow points to the position of the embedded Archaeonebria specimen. Fig. 10, basal part of the left hindleg; the arrows point to the positions of the anterior and posterior metacoxal setae. Abbreviations: mcx = metacoxa; mtr = metatrochanter; w-l = left hindwing; w-r = right hindwing." figureDoi="http://doi.org/10.5281/zenodo.3558181" httpUri="https://zenodo.org/record/3558181/files/figure.png" pageId="3" pageNumber="353">Fig. 7</figureCitation>
). Parts of the metatarsi are cut and thus lost. The exoskeleton of the
<taxonomicName id="A7CE4D74FFC0FFF1FB52D631FAA8FB24" authorityName="Laporte de Castelnau" authorityYear="1834" box="[1276,1369,1150,1176]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Nebriini">Nebriini</taxonomicName>
fossil and its negative imprint on the inclusion wall are moderately well preserved and could therefore be imaged in most details using micro-CT (
<figureCitation id="F8F52A72FFC0FFF1FE06D689FDC1FB5C" box="[424,560,1222,1248]" captionStart="FIGURES 1113" captionStartId="7.[151,264,1809,1834]" captionTargetBox="[151,1436,195,1728]" captionTargetId="figure@7.[151,1436,195,1728]" captionTargetPageId="7" captionText="FIGURES 1113. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the paratype #1. Fig. 11, dorsal aspect. Fig. 12, left lateral aspect. Fig. 13, ventral aspect. Remark: The wave-like lines covering some parts of the beetle body represent shading artifacts." figureDoi="http://doi.org/10.5281/zenodo.3558183" httpUri="https://zenodo.org/record/3558183/files/figure.png" pageId="3" pageNumber="353">Figs. 1113</figureCitation>
). Traces of the fossilized gonocoxites could also be imaged.
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D6A5FD33FA94" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
Syninclusions: One Trichoptera, three Nematocera flies (two of which are preserved in mating position), one
<taxonomicName id="A7CE4D74FFC0FFF1FF39D741FEE7FA94" authorityName="Linnaeus" authorityYear="1758" box="[151,278,1294,1320]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="3" pageNumber="353" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="A7CE4D74FFC0FFF1FE8FD741FE76FA94" authorityName="Fleming" authorityYear="1821" box="[289,391,1294,1320]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="family">Scirtidae</taxonomicName>
, stellate hairs, dirt particles.
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D77DFC6AFA29" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
<materialsCitation id="D0A63CAAFFC0FFF1FF69D77DFC66FA29" ID-GBIF-Occurrence="2460219636" collectingDate="2013-10" collectionCode="CCHH" country="Germany" county="Size" location="Female in Baltic" municipality="Muncheberg" pageId="3" pageNumber="353" specimenCount="1" stateProvince="Hamburg" typeStatus="paratype">
<emphasis id="52BAEAE5FFC0FFF1FF69D77DFE93FAF0" bold="true" box="[199,354,1329,1356]" pageId="3" pageNumber="353">
<typeStatus id="BF758855FFC0FFF1FF69D77DFEC5FAF0" box="[199,308,1330,1356]" pageId="3" pageNumber="353" type="paratype">Paratype</typeStatus>
#2:
</emphasis>
<location id="6511602CFFC0FFF1FEC4D77DFDD4FAF0" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FEC4D77DFDD4FAF0" box="[362,549,1330,1356]" country="Germany" county="Size" municipality="Muncheberg" name="Female in Baltic" pageId="3" pageNumber="353" stateProvince="Hamburg">Female in Baltic</location>
amber; no.
<collectionCode id="06DFAE32FFC0FFF1FD01D77DFD0CFAF0" box="[687,765,1330,1356]" pageId="3" pageNumber="353">CCHH</collectionCode>
#952-
<quantity id="A7369B12FFC0FFF1FCE5D77EFC83FAF0" box="[843,882,1329,1356]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="353" unit="in" value="1.0">1 in</quantity>
the collection of
<location id="6511602CFFC0FFF1FB95D77DFB65FAF0" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FB95D77DFB65FAF0" box="[1083,1172,1330,1356]" country="Germany" county="Size" municipality="Muncheberg" name="Christel" pageId="3" pageNumber="353" stateProvince="Hamburg">Christel</location>
and
<location id="6511602CFFC0FFF1FB62D77CFF21FACC" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FB62D77CFF21FACC" country="Germany" county="Size" municipality="Muncheberg" name="Hans Werner Hoffeins" pageId="3" pageNumber="353" stateProvince="Hamburg">Hans Werner Hoffeins</location>
,
<collectingRegion id="A20AF815FFC0FFF1FF72D719FEB9FACC" box="[220,328,1366,1392]" country="Germany" name="Hamburg" pageId="3" pageNumber="353">Hamburg</collectingRegion>
,
<collectingCountry id="18D97667FFC0FFF1FEFAD719FE4DFACC" box="[340,444,1366,1392]" name="Germany" pageId="3" pageNumber="353">Germany</collectingCountry>
(later to be deposited at the
<location id="6511602CFFC0FFF1FD50D719FADAFACC" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FD50D719FADAFACC" box="[766,1323,1366,1392]" country="Germany" county="Size" municipality="Muncheberg" name="Senckenberg Deutsches Entomologisches Institut" pageId="3" pageNumber="353" stateProvince="Hamburg">Senckenberg Deutsches Entomologisches Institut</location>
,
<collectingMunicipality id="8015AC8DFFC0FFF1FA99D719FF3AFA28" pageId="3" pageNumber="353">Müncheberg</collectingMunicipality>
,
<collectingCountry id="18D97667FFC0FFF1FF78D735FEB1FA28" box="[214,320,1402,1428]" name="Germany" pageId="3" pageNumber="353">Germany</collectingCountry>
) (
<figureCitation id="F8F52A72FFC0FFF1FEF9D735FE5AFA29" box="[343,427,1402,1429]" captionStart="FIGURES 1417" captionStartId="8.[151,264,1920,1945]" captionTargetBox="[151,1436,166,1901]" captionTargetId="figure@8.[151,1436,166,1901]" captionTargetPageId="8" captionText="FIGURES 1417. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #2. Fig. 14, dorsal aspect of the embedded specimen. Fig. 15, right lateral aspect. Fig. 16, general view with contours of the amber piece. Fig. 17, right antennomeres IIVI." figureDoi="http://doi.org/10.5281/zenodo.3558185" httpUri="https://zenodo.org/record/3558185/files/figure.png" pageId="3" pageNumber="353">Fig. 16</figureCitation>
).
<collectingCounty id="89104E7BFFC0FFF1FE11D735FE1EFA28" box="[447,495,1402,1428]" pageId="3" pageNumber="353">Size</collectingCounty>
of the piece approx.
<date id="14701037FFC0FFF1FD72D735FCFCFA29" box="[732,781,1402,1429]" pageId="3" pageNumber="353" value="2013-10">
<collectingDate id="0434E9DFFFC0FFF1FD72D735FCFCFA29" box="[732,781,1402,1429]" pageId="3" pageNumber="353" value="2013-10">13 x</collectingDate>
</date>
7.5
<date id="14701037FFC0FFF1FC90D734FC90FA29" box="[830,865,1403,1429]" pageId="3" pageNumber="353">
<collectingDate id="0434E9DFFFC0FFF1FC90D734FC90FA29" box="[830,865,1403,1429]" pageId="3" pageNumber="353">
x
<quantity id="A7369B12FFC0FFF1FCFDD734FC90FA29" box="[851,865,1403,1429]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="353" unit="mm" value="7.0">7</quantity>
</collectingDate>
</date>
mm
</materialsCitation>
.
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D7D1FB39F92C" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
Preservation status: The amber is clear, partly pervaded by very few flowlines; the embedded
<taxonomicName id="A7CE4D74FFC0FFF1FB4CD7D1FACEFA04" authorityName="Laporte de Castelnau" authorityYear="1834" box="[1250,1343,1438,1464]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Nebriini">Nebriini</taxonomicName>
fossil is partly covered by milky coating adjoining head and elytra (
<figureCitation id="F8F52A72FFC0FFF1FC9ED78DFC72FA60" box="[816,899,1474,1500]" captionStart="FIGURES 1417" captionStartId="8.[151,264,1920,1945]" captionTargetBox="[151,1436,166,1901]" captionTargetId="figure@8.[151,1436,166,1901]" captionTargetPageId="8" captionText="FIGURES 1417. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #2. Fig. 14, dorsal aspect of the embedded specimen. Fig. 15, right lateral aspect. Fig. 16, general view with contours of the amber piece. Fig. 17, right antennomeres IIVI." figureDoi="http://doi.org/10.5281/zenodo.3558185" httpUri="https://zenodo.org/record/3558185/files/figure.png" pageId="3" pageNumber="353">Fig. 14</figureCitation>
), and extensively covered by milky coating and small bubbles adjoining the ventral surface of the beetle body (
<figureCitation id="F8F52A72FFC0FFF1FCF9D7A9FC5BF9BC" box="[855,938,1509,1536]" captionStart="FIGURES 1417" captionStartId="8.[151,264,1920,1945]" captionTargetBox="[151,1436,166,1901]" captionTargetId="figure@8.[151,1436,166,1901]" captionTargetPageId="8" captionText="FIGURES 1417. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #2. Fig. 14, dorsal aspect of the embedded specimen. Fig. 15, right lateral aspect. Fig. 16, general view with contours of the amber piece. Fig. 17, right antennomeres IIVI." figureDoi="http://doi.org/10.5281/zenodo.3558185" httpUri="https://zenodo.org/record/3558185/files/figure.png" pageId="3" pageNumber="353">Fig. 15</figureCitation>
). Left antennomeres 911 and right metatarsomere 5 are cut and thus lost. The fossilized exoskeleton of the specimen is in parts shrunken and thus dissociated from the inclusion wall (
<figureCitation id="F8F52A72FFC0FFF1FE06D461FE0AF9F4" box="[424,507,1582,1608]" captionStart="FIGURES 1820" captionStartId="9.[151,264,1800,1825]" captionTargetBox="[153,1434,195,1731]" captionTargetId="figure@9.[151,1436,194,1731]" captionTargetPageId="9" captionText="FIGURES 1820. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the paratype #2. Figs 18, 19, dorsal aspect using different grey scale thresholds (Fig. 18 shows the negative imprint of the beetle body on the inclusion wall; Fig. 19 shows the fossilized exoskeleton which is partly dissociated from the inclusion wall). Fig. 20, left lateral aspect. Remark: The partly darkened areas near caudal apex of the beetle body represent shading artifacts." figureDoi="http://doi.org/10.5281/zenodo.3558187" httpUri="https://zenodo.org/record/3558187/files/figure.png" pageId="3" pageNumber="353">Fig. 19</figureCitation>
). However, most details of diagnostic importance of the external morphology could be reconstructed based on both, the remains of the fossilized exoskeleton and the negative imprint of the beetle body on the inclusion wall, using micro-CT (
<figureCitation id="F8F52A72FFC0FFF1FDFED439FD20F92C" box="[592,721,1654,1680]" captionStart="FIGURES 1820" captionStartId="9.[151,264,1800,1825]" captionTargetBox="[153,1434,195,1731]" captionTargetId="figure@9.[151,1436,194,1731]" captionTargetPageId="9" captionText="FIGURES 1820. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the paratype #2. Figs 18, 19, dorsal aspect using different grey scale thresholds (Fig. 18 shows the negative imprint of the beetle body on the inclusion wall; Fig. 19 shows the fossilized exoskeleton which is partly dissociated from the inclusion wall). Fig. 20, left lateral aspect. Remark: The partly darkened areas near caudal apex of the beetle body represent shading artifacts." figureDoi="http://doi.org/10.5281/zenodo.3558187" httpUri="https://zenodo.org/record/3558187/files/figure.png" pageId="3" pageNumber="353">Figs 1820</figureCitation>
,
<figureCitation id="F8F52A72FFC0FFF1FD72D439FD08F92C" box="[732,761,1654,1680]" captionStart="FIGURES 2730" captionStartId="12.[151,264,1779,1804]" captionTargetBox="[257,1330,166,1759]" captionTargetId="figure@12.[257,1330,166,1759]" captionTargetPageId="12" captionText="FIGURES 2730. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of paratypes #1 (Figs 27, 28, 30) and #2 (Fig. 29). Fig. 27, prothorax and part of the head, ventral aspect. Fig. 28, left labial palpus; the arrows point to the three setae inserted on the penultimate palpomere. Fig. 29, left metatarsomeres IIV. Fig. 30, abdomen and left metafemur, ventral view; the arrows point to the insertions of the two dorsoapical setae of the femur, the setae near apical margins of the sternites IV and V (if visible; the setae on sternite VI are shadowed by the right metatibia), and the two pairs of setae on sternite VII. Abbreviations: ce = compound eye; gu = gula; lp-2, 3 = labial palpomeres 2, 3; m = mentum; mf = metafemur; pcx = procoxa; pep = proepisternum; pst = prosternum; st-IVVII = abdominal sternites IVVII." figureDoi="http://doi.org/10.5281/zenodo.3558193" httpUri="https://zenodo.org/record/3558193/files/figure.png" pageId="3" pageNumber="353">29</figureCitation>
) including the gonocoxites (
<figureCitation id="F8F52A72FFC0FFF1FB97D439FB4CF92C" box="[1081,1213,1653,1680]" captionStart="FIGURES 4253" captionStartId="16.[151,264,1641,1666]" captionTargetBox="[290,1297,168,1620]" captionTargetId="figure@16.[289,1298,166,1621]" captionTargetPageId="16" captionText="FIGURES 4253. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the aedeagus of the holotype using different grey scale thresholds (Figs 4249), and of the female ovipositor of paratype #2 (Figs 5053; the ovipositor was separated by the segmentation function of Amira software in Figs 5153). Figs 42, 45, left lateral view. Figs 43, 46, view slightly diagonal from behind, frontally to the apical lamella. Figs 44, 47, right lateral view. Fig. 48, frontal view towards median lobe base. Fig. 49, view from posterad towards median lobe apex. Fig. 50, section of the female abdomen in ventral view. Fig. 51, ovipositor in ventral view. Fig. 52, postero-left lateroventral view. Fig. 53, dorsal view.Abbreviations: al = apical lamella of aedeagal median lobe; ap = lanceolate apophysis of left paramere; ds = dorsal row of spiniform setae on the edge of the fused apical and basal gonocoxites; g = gut; gc 1 = basal gonocoxite; gc 2 = apical gonocoxite; ltg = laterotergite; mtg = mediotergite; o-a = extreme apex of apical ostium of median lobe; o-b = extreme base of apical ostium; pm-l = left paramere; pm-r = right paramere; se = sagittal aileron; sp = sensory pit; vs = ventral row of spiniform setae (only shown on right basal gonocoxite);?vs = the pore marks the insertion of a probable additional spiniform seta of the ventral series." figureDoi="http://doi.org/10.5281/zenodo.3558199" httpUri="https://zenodo.org/record/3558199/files/figure.png" pageId="3" pageNumber="353">Figs 5053</figureCitation>
).
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D4D5FC7DF908" blockId="3.[151,1437,501,2004]" box="[199,908,1690,1717]" pageId="3" pageNumber="353">Syninclusions: No syninclusions except for a few dirt particles.</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D4F1FADFF941" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
<materialsCitation id="D0A63CAAFFC0FFF1FF69D4F1FADBF941" ID-GBIF-Occurrence="2460219637" collectingDate="2014-10-24" collectionCode="SDEI" country="Germany" county="Size" location="Female in Baltic" municipality="Muncheberg" pageId="3" pageNumber="353" specimenCount="1" typeStatus="paratype">
<emphasis id="52BAEAE5FFC0FFF1FF69D4F1FE93F964" bold="true" box="[199,354,1725,1752]" pageId="3" pageNumber="353">
<typeStatus id="BF758855FFC0FFF1FF69D4F1FEC5F964" box="[199,308,1726,1752]" pageId="3" pageNumber="353" type="paratype">Paratype</typeStatus>
#3:
</emphasis>
<location id="6511602CFFC0FFF1FEC4D4F1FDD4F964" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FEC4D4F1FDD4F964" box="[362,549,1726,1752]" country="Germany" county="Size" municipality="Muncheberg" name="Female in Baltic" pageId="3" pageNumber="353">Female in Baltic</location>
amber, deposited in the amber collection of the
<location id="6511602CFFC0FFF1FBE5D4F1FE8DF940" LSID="urn:lsid:plazi:treatment:E86787E1FFC0FFFDFF39D3E1FCFBFBBA:6511602CFFC0FFF1FBE5D4F1FE8DF940" country="Germany" county="Size" municipality="Muncheberg" name="Senckenberg Deutsches Entomologisches Institut" pageId="3" pageNumber="353">Senckenberg Deutsches Entomologisches Institut</location>
,
<collectingMunicipality id="8015AC8DFFC0FFF1FE26D4ADFDE9F940" box="[392,536,1762,1788]" pageId="3" pageNumber="353">Müncheberg</collectingMunicipality>
,
<collectingCountry id="18D97667FFC0FFF1FD8DD4ADFD7AF940" box="[547,651,1762,1788]" name="Germany" pageId="3" pageNumber="353">Germany</collectingCountry>
(
<collectionCode id="06DFAE32FFC0FFF1FD34D4ADFD2AF940" box="[666,731,1762,1788]" pageId="3" pageNumber="353">SDEI</collectionCode>
) (
<figureCitation id="F8F52A72FFC0FFF1FD5CD4ADFCB7F940" box="[754,838,1762,1788]" captionStart="FIGURES 2123" captionStartId="10.[151,264,1924,1949]" captionTargetBox="[151,1412,166,1904]" captionTargetId="figure@10.[151,1412,166,1904]" captionTargetPageId="10" captionText="FIGURES 2123. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #3. Fig. 21, dorsal aspect of the embedded specimen. Fig. 22, ventral aspect. Fig. 23, general view with contours of the amber piece." figureDoi="http://doi.org/10.5281/zenodo.3558189" httpUri="https://zenodo.org/record/3558189/files/figure.png" pageId="3" pageNumber="353">Fig. 23</figureCitation>
).
<collectingCounty id="89104E7BFFC0FFF1FCF4D4ADFC7BF940" box="[858,906,1762,1788]" pageId="3" pageNumber="353">Size</collectingCounty>
of the piece approx.
<date id="14701037FFC0FFF1FBD8D4ADFB3BF940" box="[1142,1226,1762,1789]" pageId="3" pageNumber="353" value="2014-10-24">
<collectingDate id="0434E9DFFFC0FFF1FBD8D4ADFB3BF940" box="[1142,1226,1762,1789]" pageId="3" pageNumber="353" value="2014-10-24">24 x 14</collectingDate>
</date>
x
<quantity id="A7369B12FFC0FFF1FB48D4AEFADBF941" box="[1254,1322,1761,1789]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="353" unit="mm" value="5.0">5 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D549FEA9F830" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">
Preservation status: The amber attached to the embedded
<taxonomicName id="A7CE4D74FFC0FFF1FCE0D549FC5AF89C" authorityName="Laporte de Castelnau" authorityYear="1834" box="[846,939,1798,1824]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Nebriini">Nebriini</taxonomicName>
fossil is pervaded by several flowlines, dorsally and ventrally of the fossil; its ventral surface is almost fully covered by milky coating (
<figureCitation id="F8F52A72FFC0FFF1FB14D565FAE1F8F8" box="[1210,1296,1834,1860]" captionStart="FIGURES 2123" captionStartId="10.[151,264,1924,1949]" captionTargetBox="[151,1412,166,1904]" captionTargetId="figure@10.[151,1412,166,1904]" captionTargetPageId="10" captionText="FIGURES 2123. Archaeonebria inexspectata gen. n., sp. n., light microscopic images of the paratype #3. Fig. 21, dorsal aspect of the embedded specimen. Fig. 22, ventral aspect. Fig. 23, general view with contours of the amber piece." figureDoi="http://doi.org/10.5281/zenodo.3558189" httpUri="https://zenodo.org/record/3558189/files/figure.png" pageId="3" pageNumber="353">Fig. 22</figureCitation>
). Due to the extensive flowlines, micro-CT scans resulted in low contrasts between amber and fossil particularly on its dorsal surface (
<figureCitation id="F8F52A72FFC0FFF1FF59D53DFEBDF830" box="[247,332,1906,1932]" captionStart="FIGURES 2426" captionStartId="11.[151,264,1924,1949]" captionTargetBox="[230,1340,168,1904]" captionTargetId="figure@11.[229,1342,166,1904]" captionTargetPageId="11" captionText="FIGURES 2426. Archaeonebria inexspectata gen. n., sp. n., volume rendering of the paratype #3. Fig. 24, dorsal aspect. Fig. 25, left lateral aspect. Fig. 26, ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.3558191" httpUri="https://zenodo.org/record/3558191/files/figure.png" pageId="3" pageNumber="353">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="607136F7FFC0FFF1FF69D5D9FF2BF868" blockId="3.[151,1437,501,2004]" pageId="3" pageNumber="353">Syninclusions: Two Nematocera flies, wing remains of an additional Nematocera fly, stellate hairs, dirt particles.</paragraph>
</subSubSection>
<caption id="34B1667FFFC7FFF6FF39D5CBFB44F87C" ID-DOI="http://doi.org/10.5281/zenodo.3558177" ID-Zenodo-Dep="3558177" httpUri="https://zenodo.org/record/3558177/files/figure.png" pageId="4" pageNumber="354" startId="4.[151,264,1924,1949]" targetBox="[151,1284,174,1905]" targetPageId="4">
<paragraph id="607136F7FFC7FFF6FF39D5CBFB44F87C" blockId="4.[151,1436,1924,1985]" pageId="4" pageNumber="354">
<emphasis id="52BAEAE5FFC7FFF6FF39D5CBFECBF821" bold="true" box="[151,314,1924,1949]" pageId="4" pageNumber="354">FIGURES 13.</emphasis>
<taxonomicName id="A7CE4D74FFC7FFF6FEEED5CAFDADF821" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[320,604,1925,1949]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="354" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFC7FFF6FEEED5CAFDADF821" box="[320,604,1925,1949]" italics="true" pageId="4" pageNumber="354">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFC7FFF6FDCFD5CAFD1DF821" box="[609,748,1925,1949]" pageId="4" pageNumber="354" rank="species">
<emphasis id="52BAEAE5FFC7FFF6FDCFD5CAFD59F821" bold="true" box="[609,680,1925,1949]" pageId="4" pageNumber="354">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFC7FFF6FD1AD5CAFD1DF821" bold="true" box="[692,748,1925,1949]" pageId="4" pageNumber="354">sp. n.</emphasis>
</taxonomicNameLabel>
, light microscopic images of the holotype. Fig. 1, general view with contours of the amber piece. Fig. 2, right lateral aspect of the embedded specimen. Fig. 3, dorsal aspect.
</paragraph>
</caption>
<caption id="34B1667FFFC6FFF7FF39D5CBFDF5F87C" ID-DOI="http://doi.org/10.5281/zenodo.3558179" ID-Zenodo-Dep="3558179" httpUri="https://zenodo.org/record/3558179/files/figure.png" pageId="5" pageNumber="355" startId="5.[151,264,1924,1949]" targetBox="[152,1234,167,1903]" targetPageId="5">
<paragraph id="607136F7FFC6FFF7FF39D5CBFDF5F87C" blockId="5.[151,1436,1924,1985]" pageId="5" pageNumber="355">
<emphasis id="52BAEAE5FFC6FFF7FF39D5CBFECBF821" bold="true" box="[151,314,1924,1949]" pageId="5" pageNumber="355">FIGURES 46.</emphasis>
<taxonomicName id="A7CE4D74FFC6FFF7FE91D5CAFDAAF821" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[319,603,1925,1949]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFC6FFF7FE91D5CAFDAAF821" box="[319,603,1925,1949]" italics="true" pageId="5" pageNumber="355">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFC6FFF7FDCED5CAFD1BF821" box="[608,746,1925,1949]" pageId="5" pageNumber="355" rank="species">
<emphasis id="52BAEAE5FFC6FFF7FDCED5CAFD57F821" bold="true" box="[608,678,1925,1949]" pageId="5" pageNumber="355">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFC6FFF7FD1CD5CAFD1BF821" bold="true" box="[690,746,1925,1949]" pageId="5" pageNumber="355">sp. n.</emphasis>
</taxonomicNameLabel>
, volume rendering of the holotype. Fig. 4, dorsal aspect. Fig. 5, right lateral aspect. Fig. 6, ventral aspect.
</paragraph>
</caption>
<caption id="34B1667FFFC5FFF4FF39D51BFCD9F841" ID-DOI="http://doi.org/10.5281/zenodo.3558181" ID-Zenodo-Dep="3558181" httpUri="https://zenodo.org/record/3558181/files/figure.png" pageId="6" pageNumber="356" startId="6.[151,264,1876,1901]" targetBox="[252,1329,166,1856]" targetPageId="6">
<paragraph id="607136F7FFC5FFF4FF39D51BFCD9F841" blockId="6.[151,1437,1876,2045]" pageId="6" pageNumber="356">
<emphasis id="52BAEAE5FFC5FFF4FF39D51BFEB6F8D1" bold="true" box="[151,327,1876,1901]" pageId="6" pageNumber="356">FIGURES 710.</emphasis>
<taxonomicName id="A7CE4D74FFC5FFF4FEE3D51AFD9BF8D0" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[333,618,1877,1901]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="356" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFC5FFF4FEE3D51AFD9BF8D0" box="[333,618,1877,1901]" italics="true" pageId="6" pageNumber="356">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFC5FFF4FDC1D51AFD0DF8D1" box="[623,764,1877,1901]" pageId="6" pageNumber="356" rank="species">
<emphasis id="52BAEAE5FFC5FFF4FDC1D51AFD47F8D1" bold="true" box="[623,694,1877,1901]" pageId="6" pageNumber="356">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFC5FFF4FD6DD51AFD0DF8D1" bold="true" box="[707,764,1877,1901]" pageId="6" pageNumber="356">sp. n.</emphasis>
</taxonomicNameLabel>
, light microscopic images of the paratype #1. Fig. 7, dorsal aspect of the embedded specimen. Fig. 8, ventral aspect. Fig. 9, general view with contours of the amber piece, showing several embedded insects (see text for details); the arrow points to the position of the embedded
<emphasis id="52BAEAE5FFC5FFF4FC69D5D2FBAFF809" box="[967,1118,1949,1973]" italics="true" pageId="6" pageNumber="356">Archaeonebria</emphasis>
specimen. Fig. 10, basal part of the left hindleg; the arrows point to the positions of the anterior and posterior metacoxal setae. Abbreviations: mcx = metacoxa; mtr = metatrochanter; w-l = left hindwing; w-r = right hindwing.
</paragraph>
</caption>
<caption id="34B1667FFFC4FFF5FF39D55EFEB4F8CE" ID-DOI="http://doi.org/10.5281/zenodo.3558183" ID-Zenodo-Dep="3558183" httpUri="https://zenodo.org/record/3558183/files/figure.png" pageId="7" pageNumber="357" startId="7.[151,264,1809,1834]" targetBox="[151,1436,195,1728]" targetPageId="7">
<paragraph id="607136F7FFC4FFF5FF39D55EFEB4F8CE" blockId="7.[151,1437,1809,1906]" pageId="7" pageNumber="357">
<emphasis id="52BAEAE5FFC4FFF5FF39D55EFEA2F896" bold="true" box="[151,339,1809,1834]" pageId="7" pageNumber="357">FIGURES 1113.</emphasis>
<taxonomicName id="A7CE4D74FFC4FFF5FEF6D55DFD84F896" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[344,629,1810,1834]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="357" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFC4FFF5FEF6D55DFD84F896" box="[344,629,1810,1834]" italics="true" pageId="7" pageNumber="357">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFC4FFF5FDD5D55DFCF6F896" box="[635,775,1810,1834]" pageId="7" pageNumber="357" rank="species">
<emphasis id="52BAEAE5FFC4FFF5FDD5D55DFD33F896" bold="true" box="[635,706,1810,1834]" pageId="7" pageNumber="357">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFC4FFF5FD60D55DFCF6F896" bold="true" box="[718,775,1810,1834]" pageId="7" pageNumber="357">sp. n.</emphasis>
</taxonomicNameLabel>
, volume rendering of the paratype #1. Fig. 11, dorsal aspect. Fig. 12, left lateral aspect. Fig. 13, ventral aspect. Remark: The wave-like lines covering some parts of the beetle body represent shading artifacts.
</paragraph>
</caption>
<caption id="34B1667FFFCBFFFAFF39D5CFFE51F85C" ID-DOI="http://doi.org/10.5281/zenodo.3558185" ID-Zenodo-Dep="3558185" httpUri="https://zenodo.org/record/3558185/files/figure.png" pageId="8" pageNumber="358" startId="8.[151,264,1920,1945]" targetBox="[151,1436,166,1901]" targetPageId="8">
<paragraph id="607136F7FFCBFFFAFF39D5CFFE51F85C" blockId="8.[151,1437,1920,2017]" pageId="8" pageNumber="358">
<emphasis id="52BAEAE5FFCBFFFAFF39D5CFFEA6F825" bold="true" box="[151,343,1920,1945]" pageId="8" pageNumber="358">FIGURES 1417.</emphasis>
<taxonomicName id="A7CE4D74FFCBFFFAFEF1D5CEFD8FF824" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[351,638,1921,1945]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="358" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFCBFFFAFEF1D5CEFD8FF824" box="[351,638,1921,1945]" italics="true" pageId="8" pageNumber="358">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFCBFFFAFD28D5CEFCEAF825" box="[646,795,1921,1945]" pageId="8" pageNumber="358" rank="species">
<emphasis id="52BAEAE5FFCBFFFAFD28D5CEFD21F825" bold="true" box="[646,720,1921,1945]" pageId="8" pageNumber="358">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFCBFFFAFD71D5CEFCEAF825" bold="true" box="[735,795,1921,1945]" pageId="8" pageNumber="358">sp. n.</emphasis>
</taxonomicNameLabel>
, light microscopic images of the paratype #2. Fig. 14, dorsal aspect of the embedded specimen. Fig. 15, right lateral aspect. Fig. 16, general view with contours of the amber piece. Fig. 17, right antennomeres IIVI.
</paragraph>
</caption>
<caption id="34B1667FFFCAFFFBFF39D547FC42F831" ID-DOI="http://doi.org/10.5281/zenodo.3558187" ID-Zenodo-Dep="3558187" httpUri="https://zenodo.org/record/3558187/files/figure.png" pageId="9" pageNumber="359" startId="9.[151,264,1800,1825]" targetBox="[153,1434,195,1731]" targetPageId="9">
<paragraph id="607136F7FFCAFFFBFF39D547FC42F831" blockId="9.[151,1436,1800,1933]" pageId="9" pageNumber="359">
<emphasis id="52BAEAE5FFCAFFFBFF39D547FEA4F89D" bold="true" box="[151,341,1800,1825]" pageId="9" pageNumber="359">FIGURES 1820.</emphasis>
<taxonomicName id="A7CE4D74FFCAFFFBFEF5D546FD89F89D" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[347,632,1801,1825]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="359" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFCAFFFBFEF5D546FD89F89D" box="[347,632,1801,1825]" italics="true" pageId="9" pageNumber="359">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFCAFFFBFDD0D546FCFDF89D" box="[638,780,1801,1825]" pageId="9" pageNumber="359" rank="species">
<emphasis id="52BAEAE5FFCAFFFBFDD0D546FD34F89D" bold="true" box="[638,709,1801,1825]" pageId="9" pageNumber="359">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFCAFFFBFD7CD546FCFDF89D" bold="true" box="[722,780,1801,1825]" pageId="9" pageNumber="359">sp. n.</emphasis>
</taxonomicNameLabel>
, volume rendering of the paratype #2. Figs 18, 19, dorsal aspect using different grey scale thresholds (Fig. 18 shows the negative imprint of the beetle body on the inclusion wall; Fig. 19 shows the fossilized exoskeleton which is partly dissociated from the inclusion wall). Fig. 20, left lateral aspect. Remark: The partly darkened areas near caudal apex of the beetle body represent shading artifacts.
</paragraph>
</caption>
<caption id="34B1667FFFC9FFF8FF39D5CBFAE7F87D" ID-DOI="http://doi.org/10.5281/zenodo.3558189" ID-Zenodo-Dep="3558189" httpUri="https://zenodo.org/record/3558189/files/figure.png" pageId="10" pageNumber="360" startId="10.[151,264,1924,1949]" targetBox="[151,1412,166,1904]" targetPageId="10">
<paragraph id="607136F7FFC9FFF8FF39D5CBFAE7F87D" blockId="10.[151,1436,1924,1985]" pageId="10" pageNumber="360">
<emphasis id="52BAEAE5FFC9FFF8FF39D5CBFEA6F821" bold="true" box="[151,343,1924,1949]" pageId="10" pageNumber="360">FIGURES 2123.</emphasis>
<taxonomicName id="A7CE4D74FFC9FFF8FEF1D5CBFD8FF820" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[351,638,1924,1948]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="360" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFC9FFF8FEF1D5CBFD8FF820" box="[351,638,1924,1948]" italics="true" pageId="10" pageNumber="360">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFC9FFF8FD28D5CBFCEAF820" box="[646,795,1924,1948]" pageId="10" pageNumber="360" rank="species">
<emphasis id="52BAEAE5FFC9FFF8FD28D5CBFD21F820" bold="true" box="[646,720,1924,1948]" pageId="10" pageNumber="360">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFC9FFF8FD71D5CBFCEAF820" bold="true" box="[735,795,1924,1948]" pageId="10" pageNumber="360">sp. n.</emphasis>
</taxonomicNameLabel>
, light microscopic images of the paratype #3. Fig. 21, dorsal aspect of the embedded specimen. Fig. 22, ventral aspect. Fig. 23, general view with contours of the amber piece.
</paragraph>
</caption>
<caption id="34B1667FFFC8FFF9FF39D5CBFDAEF87C" ID-DOI="http://doi.org/10.5281/zenodo.3558191" ID-Zenodo-Dep="3558191" httpUri="https://zenodo.org/record/3558191/files/figure.png" pageId="11" pageNumber="361" startId="11.[151,264,1924,1949]" targetBox="[230,1340,168,1904]" targetPageId="11">
<paragraph id="607136F7FFC8FFF9FF39D5CBFDAEF87C" blockId="11.[151,1437,1924,1985]" pageId="11" pageNumber="361">
<emphasis id="52BAEAE5FFC8FFF9FF39D5CBFEA5F821" bold="true" box="[151,340,1924,1949]" pageId="11" pageNumber="361">FIGURES 2426.</emphasis>
<taxonomicName id="A7CE4D74FFC8FFF9FEF4D5CAFD87F821" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[346,630,1925,1949]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="361" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFC8FFF9FEF4D5CAFD87F821" box="[346,630,1925,1949]" italics="true" pageId="11" pageNumber="361">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFC8FFF9FDD5D5CAFCF6F821" box="[635,775,1925,1949]" pageId="11" pageNumber="361" rank="species">
<emphasis id="52BAEAE5FFC8FFF9FDD5D5CAFD33F821" bold="true" box="[635,706,1925,1949]" pageId="11" pageNumber="361">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFC8FFF9FD61D5CAFCF6F821" bold="true" box="[719,775,1925,1949]" pageId="11" pageNumber="361">sp. n.</emphasis>
</taxonomicNameLabel>
, volume rendering of the paratype #3. Fig. 24, dorsal aspect. Fig. 25, left lateral aspect. Fig. 26, ventral aspect.
</paragraph>
</caption>
<caption id="34B1667FFFCFFFFEFF39D4BCFC7CF85F" ID-DOI="http://doi.org/10.5281/zenodo.3558193" ID-Zenodo-Dep="3558193" httpUri="https://zenodo.org/record/3558193/files/figure.png" pageId="12" pageNumber="362" startId="12.[151,264,1779,1804]" targetBox="[257,1330,166,1759]" targetPageId="12">
<paragraph id="607136F7FFCFFFFEFF39D4BCFC7CF85F" blockId="12.[151,1437,1779,2020]" pageId="12" pageNumber="362">
<emphasis id="52BAEAE5FFCFFFFEFF39D4BCFEA4F8B0" bold="true" box="[151,341,1779,1804]" pageId="12" pageNumber="362">FIGURES 2730.</emphasis>
<taxonomicName id="A7CE4D74FFCFFFFEFEF5D4BCFD89F8B7" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[347,632,1779,1803]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="362" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFCFFFFEFEF5D4BCFD89F8B7" box="[347,632,1779,1803]" italics="true" pageId="12" pageNumber="362">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFCFFFFEFDD1D4BBFCFFF8B0" box="[639,782,1780,1804]" pageId="12" pageNumber="362" rank="species">
<emphasis id="52BAEAE5FFCFFFFEFDD1D4BBFD36F8B0" bold="true" box="[639,711,1780,1804]" pageId="12" pageNumber="362">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFCFFFFEFD7AD4BBFCFFF8B0" bold="true" box="[724,782,1780,1804]" pageId="12" pageNumber="362">sp. n.</emphasis>
</taxonomicNameLabel>
, light microscopic images of paratypes #1 (Figs 27, 28, 30) and #2 (Fig. 29). Fig. 27, prothorax and part of the head, ventral aspect. Fig. 28, left labial palpus; the arrows point to the three setae inserted on the penultimate palpomere. Fig. 29, left metatarsomeres IIV. Fig. 30, abdomen and left metafemur, ventral view; the arrows point to the insertions of the two dorsoapical setae of the femur, the setae near apical margins of the sternites IV and V (if visible; the setae on sternite VI are shadowed by the right metatibia), and the two pairs of setae on sternite VII. Abbreviations: ce = compound eye; gu = gula; lp-2, 3 = labial palpomeres 2, 3; m = mentum; mf = metafemur; pcx = procoxa; pep = proepisternum; pst = prosternum; st-IVVII = abdominal sternites IVVII.
</paragraph>
</caption>
<caption id="34B1667FFFCEFFFFFF39D4CCFEF8F85C" ID-DOI="http://doi.org/10.5281/zenodo.3558195" ID-Zenodo-Dep="3558195" httpUri="https://zenodo.org/record/3558195/files/figure.png" pageId="13" pageNumber="363" startId="13.[151,264,1667,1692]" targetBox="[151,1436,166,1602]" targetPageId="13">
<paragraph id="607136F7FFCEFFFFFF39D4CCFEF8F85C" blockId="13.[151,1437,1667,2016]" pageId="13" pageNumber="363">
<emphasis id="52BAEAE5FFCEFFFFFF39D4CCFEA5F920" bold="true" box="[151,340,1667,1692]" pageId="13" pageNumber="363">FIGURES 3136.</emphasis>
<taxonomicName id="A7CE4D74FFCEFFFFFEF4D4CBFD87F920" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[346,630,1668,1692]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="363" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFCEFFFFFEF4D4CBFD87F920" box="[346,630,1668,1692]" italics="true" pageId="13" pageNumber="363">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFCEFFFFFDD2D4CBFCF9F920" box="[636,776,1668,1692]" pageId="13" pageNumber="363" rank="species">
<emphasis id="52BAEAE5FFCEFFFFFDD2D4CBFD33F920" bold="true" box="[636,706,1668,1692]" pageId="13" pageNumber="363">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFCEFFFFFD61D4CBFCF9F920" bold="true" box="[719,776,1668,1692]" pageId="13" pageNumber="363">sp. n.</emphasis>
</taxonomicNameLabel>
, volume rendering of the holotype (Figs 31, 32, 34) and paratype #2 (Fig. 33, 35, 36). Fig. 31, head, dorsal aspect; the arrows point to the insertions of the labral setae, clypeal seta, and supraorbital seta. Remark: Note the deformation of the labral apical margin with respect to its negative imprint on the inclusion wall, which represents its original state. Fig. 32, head, ventral aspect; the arrows point to the insertions of the setae on mentum and submentum. Figs 33, 34, mouthparts, apicoventral aspect of different specimens using different greyscale thresholds. Figs 35, 36, separated images of the ligula with the completely fused paraglossae (Fig. 35, ventral aspect; Fig. 36, apicoventral aspect). Abbreviations: ant = antenna; dp = dust particles; la = lacinia; lig = ligula; lig-a = apex of ligula; lp-l = left labial palpus; lp-r = right labial palpus; m = mentum; md-l = left mandible; md-r = right mandible; mp-l = left maxillary palpus; mp-r = right maxillary palpus; mx = maxilla; n = negative imprint of the labral margin; p = shrunken remains of the labral margin; pgl = paraglossa.
</paragraph>
</caption>
<caption id="34B1667FFFCDFFFCFF39D722FA95FA4D" ID-DOI="http://doi.org/10.5281/zenodo.3558197" ID-Zenodo-Dep="3558197" httpUri="https://zenodo.org/record/3558197/files/figure.png" pageId="14" pageNumber="364" startId="14.[151,264,1389,1414]" targetBox="[153,1433,166,1335]" targetPageId="14">
<paragraph id="607136F7FFCDFFFCFF39D722FA95FA4D" blockId="14.[151,1437,1389,1522]" pageId="14" pageNumber="364">
<emphasis id="52BAEAE5FFCDFFFCFF39D722FEA4FA3A" bold="true" box="[151,341,1389,1414]" pageId="14" pageNumber="364">FIGURES 3741.</emphasis>
<taxonomicName id="A7CE4D74FFCDFFFCFEF3D722FD8BFA39" authority="Schmidt &amp; Scholz &amp; Kavanaugh, 2019" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[349,634,1389,1413]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="364" phylum="Arthropoda" rank="species" species="inexspectata" status="gen. n., sp. n.">
<emphasis id="52BAEAE5FFCDFFFCFEF3D722FD8BFA39" box="[349,634,1389,1413]" italics="true" pageId="14" pageNumber="364">Archaeonebria inexspectata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4989579EFFCDFFFCFD2FD721FCE3FA3A" box="[641,786,1390,1414]" pageId="14" pageNumber="364" rank="species">
<emphasis id="52BAEAE5FFCDFFFCFD2FD721FD3BFA3A" bold="true" box="[641,714,1390,1414]" pageId="14" pageNumber="364">gen. n.</emphasis>
,
<emphasis id="52BAEAE5FFCDFFFCFD76D721FCE3FA3A" bold="true" box="[728,786,1390,1414]" pageId="14" pageNumber="364">sp. n.</emphasis>
</taxonomicNameLabel>
, volume rendering. Fig. 37, pronotum of the holotype. Fig. 38, pronotum of the paratype #1. Fig. 39, pronotum of the paratype #2; the arrows point to the lateral seta (pls) and basolateral seta (pbs). Figs 40, 41, right elytron of the holotype (Fig. 40, lateral aspect, Fig. 41, dorsal aspect); the arrows point to the insertions of the parascutellar seta (pss), discal setae (ds1
<emphasis id="52BAEAE5FFCDFFFCFDDFD795FD8FFA4E" bold="true" box="[625,638,1498,1522]" pageId="14" pageNumber="364"></emphasis>
3), praeapical seta (pas), and to the setae of the umbilicate series (us1
<emphasis id="52BAEAE5FFCDFFFCFA92D795FAB8FA4E" bold="true" box="[1340,1353,1498,1522]" pageId="14" pageNumber="364"></emphasis>
9).
</paragraph>
</caption>
<subSubSection id="28D4657CFFCDFFFDFF69D450FD86FC02" lastPageId="15" lastPageNumber="365" pageId="14" pageNumber="364" type="description">
<paragraph id="607136F7FFCDFFFCFF69D450FDA0F9E2" blockId="14.[151,1437,1567,2026]" pageId="14" pageNumber="364">
<emphasis id="52BAEAE5FFCDFFFCFF69D450FEA8F986" bold="true" box="[199,345,1567,1594]" pageId="14" pageNumber="364">Description.</emphasis>
Color: All specimens appear unicolored dark throughout, with pronotum and elytra markedly shiny. No metallic reflection is evident.
</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D427FE75F977" blockId="14.[151,1437,1567,2026]" pageId="14" pageNumber="364">
Microsculpture: Not visible on any part of the male
<typeStatus id="BF758855FFCDFFFCFCB0D427FC72F93E" box="[798,899,1640,1666]" pageId="14" pageNumber="364" type="holotype">holotype</typeStatus>
, and on the head surfaces of any of the female
<typeStatus id="BF758855FFCDFFFCFF39D4C3FEF7F91A" box="[151,262,1676,1702]" pageId="14" pageNumber="364" type="paratype">paratypes</typeStatus>
; on pronotum and elytra of females with very finely impressed small transverse meshes (visible at magnifications of&gt;
<date id="14701037FFCDFFFCFEE4D4FFFE89F977" box="[330,376,1712,1739]" pageId="14" pageNumber="364" value="1960-10">60x</date>
).
</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D49BFBD9F8AE" blockId="14.[151,1437,1567,2026]" pageId="14" pageNumber="364">
Measurements: Standardized body length 6.4 (
<typeStatus id="BF758855FFCDFFFCFD78D49BFCCDF952" box="[726,828,1748,1774]" pageId="14" pageNumber="364" type="holotype">holotype</typeStatus>
,
<typeStatus id="BF758855FFCDFFFCFCE7D49BFC58F952" box="[841,937,1748,1774]" pageId="14" pageNumber="364" type="paratype">paratype</typeStatus>
#1) to
<quantity id="A7369B12FFCDFFFCFC53D49BFBA4F952" box="[1021,1109,1748,1774]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="14" pageNumber="364" unit="mm" value="7.2">7.2 mm</quantity>
(
<typeStatus id="BF758855FFCDFFFCFBCAD49BFB37F952" box="[1124,1222,1748,1774]" pageId="14" pageNumber="364" type="paratype">paratype</typeStatus>
#3). Length of aedeagal median lobe
<quantity id="A7369B12FFCDFFFCFED6D4B7FE2DF8AE" box="[376,476,1784,1810]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="14" pageNumber="364" unit="mm" value="0.94">0.94 mm</quantity>
(
<typeStatus id="BF758855FFCDFFFCFE44D4B7FDA2F8AF" box="[490,595,1784,1811]" pageId="14" pageNumber="364" type="holotype">holotype</typeStatus>
). For additional measurements see
<tableCitation id="2D4C034CFFCDFFFCFC75D4B7FBD5F8AE" box="[987,1060,1784,1810]" captionStart="TABLE 2" captionStartId="15.[151,239,334,358]" captionTargetBox="[166,1407,370,811]" captionTargetPageId="15" captionText="TABLE 2. Measurements [µm]." httpUri="http://table.plazi.org/id/34B1667FFFCCFFFDFF39D301FE01FEDB" pageId="14" pageNumber="364" tableUuid="34B1667FFFCCFFFDFF39D301FE01FEDB">Tab. 2</tableCitation>
.
</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D553FC14F88A" blockId="14.[151,1437,1567,2026]" box="[199,997,1820,1846]" pageId="14" pageNumber="364">
Proportions, given for
<typeStatus id="BF758855FFCDFFFCFE6BD553FDDAF88A" box="[453,555,1820,1846]" pageId="14" pageNumber="364" type="holotype">holotype</typeStatus>
;
<typeStatus id="BF758855FFCDFFFCFD98D553FD67F88A" box="[566,662,1820,1846]" pageId="14" pageNumber="364" type="paratype">paratype</typeStatus>
#1;
<typeStatus id="BF758855FFCDFFFCFD66D553FCD9F88A" box="[712,808,1820,1846]" pageId="14" pageNumber="364" type="paratype">paratype</typeStatus>
#2;
<typeStatus id="BF758855FFCDFFFCFCF4D553FC4BF88A" box="[858,954,1820,1846]" pageId="14" pageNumber="364" type="paratype">paratype</typeStatus>
#3:
</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D50FFDCCF8E6" blockId="14.[151,1437,1567,2026]" box="[199,573,1856,1882]" pageId="14" pageNumber="364">PW/HW = 1.36; 1.34; 1.41; 1.43.</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D52BFDDFF8C2" blockId="14.[151,1437,1567,2026]" box="[199,558,1891,1918]" pageId="14" pageNumber="364">PW/PL = 1.59; 1.65; 1.62; 1.65.</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D5C7FDB7F81E" blockId="14.[151,1437,1567,2026]" box="[199,582,1928,1954]" pageId="14" pageNumber="364">PW/PWb = 1.03; 1.07; 1.08; 1.13.</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D5E3FDA0F87A" blockId="14.[151,1437,1567,2026]" box="[199,593,1964,1990]" pageId="14" pageNumber="364">PWb/PWa = 1.36; 1.27; 1.28; 1.22.</paragraph>
<paragraph id="607136F7FFCDFFFCFF69D59FFDC8F856" blockId="14.[151,1437,1567,2026]" box="[199,569,2000,2026]" pageId="14" pageNumber="364">EW/PW = 1.06; 1.22; 1.18; 1.18.</paragraph>
<paragraph id="607136F7FFCCFFFDFF69D2D6FDC3FF0F" blockId="15.[199,1221,152,288]" box="[199,562,152,179]" pageId="15" pageNumber="365">EL/EW = 1.50; 1.50; 1.46; 1.47.</paragraph>
<paragraph id="607136F7FFCCFFFDFF69D2F2FDD7FF6B" blockId="15.[199,1221,152,288]" box="[199,550,188,215]" pageId="15" pageNumber="365">EL/FL = 2.00; 2.10; 2.05; 2.16.</paragraph>
<paragraph id="607136F7FFCCFFFDFF69D2AEFB34FF47" blockId="15.[199,1221,152,288]" box="[199,1221,225,252]" pageId="15" pageNumber="365">
LPL3/LPL2 (minmax) = 1.211.23; 1.191.24; not measurable in
<typeStatus id="BF758855FFCCFFFDFC15D2AEFBEAFF47" box="[955,1051,225,251]" pageId="15" pageNumber="365" type="paratype">paratype</typeStatus>
#2; 1.261.28.
</paragraph>
<paragraph id="607136F7FFCCFFFDFF69D34AFDF3FE9C" blockId="15.[199,1221,152,288]" box="[199,514,260,288]" pageId="15" pageNumber="365">
EL/AedL = 4.15 (
<typeStatus id="BF758855FFCCFFFDFE20D34AFE06FE9C" box="[398,503,261,288]" pageId="15" pageNumber="365" type="holotype">holotype</typeStatus>
).
</paragraph>
<caption id="34B1667FFFCCFFFDFF39D301FE01FEDB" ID-Table-UUID="34B1667FFFCCFFFDFF39D301FE01FEDB" box="[151,496,334,359]" httpUri="http://table.plazi.org/id/34B1667FFFCCFFFDFF39D301FE01FEDB" pageId="15" pageNumber="365" targetBox="[166,1407,370,811]" targetIsTable="true" targetPageId="15">
<paragraph id="607136F7FFCCFFFDFF39D301FE01FEDB" blockId="15.[151,496,334,359]" box="[151,496,334,359]" pageId="15" pageNumber="365">
<emphasis id="52BAEAE5FFCCFFFDFF39D301FEFBFEDA" bold="true" box="[151,266,334,358]" pageId="15" pageNumber="365">TABLE 2.</emphasis>
Measurements [µm].
</paragraph>
</caption>
<paragraph id="607136F7FFCCFFFDFD26D33DFA9AFC97" pageId="15" pageNumber="365">
<table id="12CEC457FFCC000DFF08D33DFA8EFC97" box="[166,1407,370,811]" gridcols="5" gridrows="12" pageId="15" pageNumber="365">
<tr id="DEFE34B5FFCC000DFF08D33DFA8EFE37" box="[166,1407,370,395]" gridrow="0" pageId="15" pageNumber="365" rowspan-0="1">
<th id="9D2F5DC9FFCC000DFD26D33DFCF3FE37" box="[648,770,370,395]" gridcol="1" gridrow="0" pageId="15" pageNumber="365">holotype</th>
<th id="9D2F5DC9FFCC000DFCF6D33DFC14FE37" box="[856,997,370,395]" gridcol="2" gridrow="0" pageId="15" pageNumber="365">paratype “#1”</th>
<th id="9D2F5DC9FFCC000DFB89D33DFB4DFE37" box="[1063,1212,370,395]" gridcol="3" gridrow="0" pageId="15" pageNumber="365">paratype “#2”</th>
<th id="9D2F5DC9FFCC000DFB5CD33DFA8EFE37" box="[1266,1407,370,395]" gridcol="4" gridrow="0" pageId="15" pageNumber="365">paratype “#3”</th>
</tr>
<tr id="DEFE34B5FFCC000DFF08D3D7FA8EFE0D" box="[166,1407,408,433]" gridrow="1" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D3D7FDDEFE0D" box="[166,559,408,433]" gridcol="0" gridrow="1" pageId="15" pageNumber="365">length of head</th>
<td id="9D2F5DC9FFCC000DFD26D3D7FCF3FE0D" box="[648,770,408,433]" gridcol="1" gridrow="1" pageId="15" pageNumber="365">939</td>
<td id="9D2F5DC9FFCC000DFCF6D3D7FC14FE0D" box="[856,997,408,433]" gridcol="2" gridrow="1" pageId="15" pageNumber="365">896</td>
<td id="9D2F5DC9FFCC000DFB89D3D7FB4DFE0D" box="[1063,1212,408,433]" gridcol="3" gridrow="1" pageId="15" pageNumber="365">922</td>
<td id="9D2F5DC9FFCC000DFB5CD3D7FA8EFE0D" box="[1266,1407,408,433]" gridcol="4" gridrow="1" pageId="15" pageNumber="365">1013</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D3F1FA8EFE6B" box="[166,1407,446,471]" gridrow="2" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D3F1FDDEFE6B" box="[166,559,446,471]" gridcol="0" gridrow="2" pageId="15" pageNumber="365">width of head</th>
<td id="9D2F5DC9FFCC000DFD26D3F1FCF3FE6B" box="[648,770,446,471]" gridcol="1" gridrow="2" pageId="15" pageNumber="365">1810</td>
<td id="9D2F5DC9FFCC000DFCF6D3F1FC14FE6B" box="[856,997,446,471]" gridcol="2" gridrow="2" pageId="15" pageNumber="365">1674</td>
<td id="9D2F5DC9FFCC000DFB89D3F1FB4DFE6B" box="[1063,1212,446,471]" gridcol="3" gridrow="2" pageId="15" pageNumber="365">1859</td>
<td id="9D2F5DC9FFCC000DFB5CD3F1FA8EFE6B" box="[1266,1407,446,471]" gridcol="4" gridrow="2" pageId="15" pageNumber="365">1846</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D3ABFA8EFE41" box="[166,1407,484,509]" gridrow="3" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D3ABFDDEFE41" box="[166,559,484,509]" gridcol="0" gridrow="3" pageId="15" pageNumber="365">length of pronotum</th>
<td id="9D2F5DC9FFCC000DFD26D3ABFCF3FE41" box="[648,770,484,509]" gridcol="1" gridrow="3" pageId="15" pageNumber="365">1547</td>
<td id="9D2F5DC9FFCC000DFCF6D3ABFC14FE41" box="[856,997,484,509]" gridcol="2" gridrow="3" pageId="15" pageNumber="365">1363</td>
<td id="9D2F5DC9FFCC000DFB89D3ABFB4DFE41" box="[1063,1212,484,509]" gridcol="3" gridrow="3" pageId="15" pageNumber="365">1624</td>
<td id="9D2F5DC9FFCC000DFB5CD3ABFA8EFE41" box="[1266,1407,484,509]" gridcol="4" gridrow="3" pageId="15" pageNumber="365">1603</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D046FA8EFD9E" box="[166,1407,521,546]" gridrow="4" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D046FDDEFD9E" box="[166,559,521,546]" gridcol="0" gridrow="4" pageId="15" pageNumber="365">width of pronotum</th>
<td id="9D2F5DC9FFCC000DFD26D046FCF3FD9E" box="[648,770,521,546]" gridcol="1" gridrow="4" pageId="15" pageNumber="365">2461</td>
<td id="9D2F5DC9FFCC000DFCF6D046FC14FD9E" box="[856,997,521,546]" gridcol="2" gridrow="4" pageId="15" pageNumber="365">2248</td>
<td id="9D2F5DC9FFCC000DFB89D046FB4DFD9E" box="[1063,1212,521,546]" gridcol="3" gridrow="4" pageId="15" pageNumber="365">2624</td>
<td id="9D2F5DC9FFCC000DFB5CD046FA8EFD9E" box="[1266,1407,521,546]" gridcol="4" gridrow="4" pageId="15" pageNumber="365">2644</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D060FA8EFDF4" box="[166,1407,559,584]" gridrow="5" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D060FDDEFDF4" box="[166,559,559,584]" gridcol="0" gridrow="5" pageId="15" pageNumber="365">pronotal apical width</th>
<td id="9D2F5DC9FFCC000DFD26D060FCF3FDF4" box="[648,770,559,584]" gridcol="1" gridrow="5" pageId="15" pageNumber="365">1762</td>
<td id="9D2F5DC9FFCC000DFCF6D060FC14FDF4" box="[856,997,559,584]" gridcol="2" gridrow="5" pageId="15" pageNumber="365">1694</td>
<td id="9D2F5DC9FFCC000DFB89D060FB4DFDF4" box="[1063,1212,559,584]" gridcol="3" gridrow="5" pageId="15" pageNumber="365">1899</td>
<td id="9D2F5DC9FFCC000DFB5CD060FA8EFDF4" box="[1266,1407,559,584]" gridcol="4" gridrow="5" pageId="15" pageNumber="365">1922</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D01AFA8EFDD2" box="[166,1407,597,622]" gridrow="6" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D01AFDDEFDD2" box="[166,559,597,622]" gridcol="0" gridrow="6" pageId="15" pageNumber="365">pronotal basal width</th>
<td id="9D2F5DC9FFCC000DFD26D01AFCF3FDD2" box="[648,770,597,622]" gridcol="1" gridrow="6" pageId="15" pageNumber="365">2390</td>
<td id="9D2F5DC9FFCC000DFCF6D01AFC14FDD2" box="[856,997,597,622]" gridcol="2" gridrow="6" pageId="15" pageNumber="365">2096</td>
<td id="9D2F5DC9FFCC000DFB89D01AFB4DFDD2" box="[1063,1212,597,622]" gridcol="3" gridrow="6" pageId="15" pageNumber="365">2429</td>
<td id="9D2F5DC9FFCC000DFB5CD01AFA8EFDD2" box="[1266,1407,597,622]" gridcol="4" gridrow="6" pageId="15" pageNumber="365">2338</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D034FA8EFD28" box="[166,1407,635,660]" gridrow="7" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D034FDDEFD28" box="[166,559,635,660]" gridcol="0" gridrow="7" pageId="15" pageNumber="365">length of left / right elytron</th>
<td id="9D2F5DC9FFCC000DFD26D034FCF3FD28" box="[648,770,635,660]" gridcol="1" gridrow="7" pageId="15" pageNumber="365">3904 / 3892</td>
<td id="9D2F5DC9FFCC000DFCF6D034FC14FD28" box="[856,997,635,660]" gridcol="2" gridrow="7" pageId="15" pageNumber="365">4070 / 4117</td>
<td id="9D2F5DC9FFCC000DFB89D034FB4DFD28" box="[1063,1212,635,660]" gridcol="3" gridrow="7" pageId="15" pageNumber="365">4483 / 4500</td>
<td id="9D2F5DC9FFCC000DFB5CD034FA8EFD28" box="[1266,1407,635,660]" gridcol="4" gridrow="7" pageId="15" pageNumber="365">4574 / 4574</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D0EEFA8EFD06" box="[166,1407,673,698]" gridrow="8" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D0EEFDDEFD06" box="[166,559,673,698]" gridcol="0" gridrow="8" pageId="15" pageNumber="365">width of left / right elytron</th>
<td id="9D2F5DC9FFCC000DFD26D0EEFCF3FD06" box="[648,770,673,698]" gridcol="1" gridrow="8" pageId="15" pageNumber="365">1252 / 1345</td>
<td id="9D2F5DC9FFCC000DFCF6D0EEFC14FD06" box="[856,997,673,698]" gridcol="2" gridrow="8" pageId="15" pageNumber="365">1381 / 1365</td>
<td id="9D2F5DC9FFCC000DFB89D0EEFB4DFD06" box="[1063,1212,673,698]" gridcol="3" gridrow="8" pageId="15" pageNumber="365">1522 / 1564</td>
<td id="9D2F5DC9FFCC000DFB5CD0EEFA8EFD06" box="[1266,1407,673,698]" gridcol="4" gridrow="8" pageId="15" pageNumber="365">1577 / 1544</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D089FA8EFD63" box="[166,1407,710,735]" gridrow="9" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D089FDDEFD63" box="[166,559,710,735]" gridcol="0" gridrow="9" pageId="15" pageNumber="365">length of left / right labial palpomere 2</th>
<td id="9D2F5DC9FFCC000DFD26D089FCF3FD63" box="[648,770,710,735]" gridcol="1" gridrow="9" pageId="15" pageNumber="365">390 / 385</td>
<td id="9D2F5DC9FFCC000DFCF6D089FC14FD63" box="[856,997,710,735]" gridcol="2" gridrow="9" pageId="15" pageNumber="365">373 / 370</td>
<td id="9D2F5DC9FFCC000DFB89D089FB4DFD63" box="[1063,1212,710,735]" gridcol="3" gridrow="9" pageId="15" pageNumber="365">not observable</td>
<td id="9D2F5DC9FFCC000DFB5CD089FA8EFD63" box="[1266,1407,710,735]" gridcol="4" gridrow="9" pageId="15" pageNumber="365">369 / 370</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D0A3FA8EFCB9" box="[166,1407,748,773]" gridrow="10" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D0A3FDDEFCB9" box="[166,559,748,773]" gridcol="0" gridrow="10" pageId="15" pageNumber="365">length of left / right labial palpomere 3</th>
<td id="9D2F5DC9FFCC000DFD26D0A3FCF3FCB9" box="[648,770,748,773]" gridcol="1" gridrow="10" pageId="15" pageNumber="365">478 / 466</td>
<td id="9D2F5DC9FFCC000DFCF6D0A3FC14FCB9" box="[856,997,748,773]" gridcol="2" gridrow="10" pageId="15" pageNumber="365">461 / 442</td>
<td id="9D2F5DC9FFCC000DFB89D0A3FB4DFCB9" box="[1063,1212,748,773]" gridcol="3" gridrow="10" pageId="15" pageNumber="365">not observable</td>
<td id="9D2F5DC9FFCC000DFB5CD0A3FA8EFCB9" box="[1266,1407,748,773]" gridcol="4" gridrow="10" pageId="15" pageNumber="365">466 / 476</td>
</tr>
<tr id="DEFE34B5FFCC000DFF08D15DFA8EFC97" box="[166,1407,786,811]" gridrow="11" pageId="15" pageNumber="365">
<th id="9D2F5DC9FFCC000DFF08D15DFDDEFC97" box="[166,559,786,811]" gridcol="0" gridrow="11" pageId="15" pageNumber="365">length of left / right metafemur</th>
<td id="9D2F5DC9FFCC000DFD26D15DFCF3FC97" box="[648,770,786,811]" gridcol="1" gridrow="11" pageId="15" pageNumber="365">1937 / 1949</td>
<td id="9D2F5DC9FFCC000DFCF6D15DFC14FC97" box="[856,997,786,811]" gridcol="2" gridrow="11" pageId="15" pageNumber="365">1935 / 1955</td>
<td id="9D2F5DC9FFCC000DFB89D15DFB4DFC97" box="[1063,1212,786,811]" gridcol="3" gridrow="11" pageId="15" pageNumber="365">2194 / 2027</td>
<td id="9D2F5DC9FFCC000DFB5CD15DFA8EFC97" box="[1266,1407,786,811]" gridcol="4" gridrow="11" pageId="15" pageNumber="365">2120 / 2070</td>
</tr>
</table>
</paragraph>
<paragraph id="607136F7FFCCFFFDFF69D113FD86FC02" blockId="15.[151,1437,860,1030]" pageId="15" pageNumber="365">
Variation: The pronotum is less transverse, and its extension is larger, in the male
<typeStatus id="BF758855FFCCFFFDFBD8D113FB29FCCA" box="[1142,1240,860,886]" pageId="15" pageNumber="365" type="holotype">holotype</typeStatus>
than in the three female specimens. In the female
<typeStatus id="BF758855FFCCFFFDFDBDD1CFFD82FC26" box="[531,627,896,922]" pageId="15" pageNumber="365" type="paratype">paratype</typeStatus>
#3 the pronotum is less constricted towards the base than in the other specimens, and its legs are slightly shorter.
</paragraph>
</subSubSection>
<subSubSection id="28D4657CFFCCFFFDFF69D187FCFBFBBA" pageId="15" pageNumber="365" type="etymology">
<paragraph id="607136F7FFCCFFFDFF69D187FCFBFBBA" blockId="15.[151,1437,860,1030]" pageId="15" pageNumber="365">
<emphasis id="52BAEAE5FFCCFFFDFF69D187FEBFFC5E" bold="true" box="[199,334,968,994]" pageId="15" pageNumber="365">Etymology.</emphasis>
The species epithet,
<taxonomicName id="A7CE4D74FFCCFFFDFDEFD186FD3CFC5E" authorityName="Schmidt &amp; Scholz &amp; Kavanaugh" authorityYear="2019" box="[577,717,969,994]" class="Insecta" family="Carabidae" genus="Archaeonebria" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="365" phylum="Arthropoda" rank="species" species="inexspectata">
<emphasis id="52BAEAE5FFCCFFFDFDEFD186FD3CFC5E" box="[577,717,969,994]" italics="true" pageId="15" pageNumber="365">inexspectata</emphasis>
</taxonomicName>
, is a Latin adjective meaning “unexpected”, referring to unexpected occurrence of a nebriine carabid in Baltic amber.
</paragraph>
</subSubSection>
</treatment>
</document>