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<mods:title id="2BF34FC690ACE65C1171164B889BD9E7">Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXIII Buthus (Buthidae), with description of two new species</mods:title>
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<taxonomicName id="C9984D4FDC4F40096BC5FD55FB5A52A3" ID-CoL="NZ27" authority="Kovařík &amp; Šťáhlavský &amp; Elmi, 2020" authorityName="Kovařík &amp; Šťáhlavský &amp; Elmi" authorityYear="2020" box="[980,1237,680,707]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="somalilandus" status="sp. nov.">Buthus somalilandus</taxonomicName>
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<paragraph id="0E2736CCDC4F40096B58FD37FA265282" blockId="18.[827,1464,680,802]" box="[841,1449,714,738]" pageId="18" pageNumber="17">
(
<figureCitation id="96A32A49DC4F40096B41FD37FB875282" box="[848,1032,714,738]" captionStart-0="Figures 102105" captionStart-1="Figures 106113" captionStart-2="Figures 114131" captionStart-3="Figures 132139" captionStartId-0="19.[113,190,1973,1995]" captionStartId-1="20.[116,193,1958,1980]" captionStartId-2="21.[113,190,1904,1926]" captionStartId-3="22.[115,192,1958,1980]" captionTargetBox-0="[133,1468,266,1933]" captionTargetBox-1="[134,1471,244,1910]" captionTargetBox-2="[113,1465,243,1889]" captionTargetBox-3="[113,1474,250,1884]" captionTargetId-0="figure-49@19.[113,1474,248,1939]" captionTargetId-1="figure-81@20.[114,1475,234,1936]" captionTargetId-2="figure-8@21.[113,1474,234,1903]" captionTargetId-3="figure-18@22.[113,1474,250,1920]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="Figures 102105: Buthus somalilandus sp. n. Figures 102103. Male in dorsal (102) and ventral (103) views. Figures 104105. Female in dorsal (104) and ventral (105) views. Scale bar: 10 mm." captionText-1="Figures 106113: Buthus somalilandus sp. n. Figures 106, 108110. Male, telson lateral (106), and metasoma and telson lateral (108), dorsal (109), and ventral (111) views. Figures 107, 111113. Female, telson lateral (107), and metasoma and telson lateral (111), dorsal (112), and ventral (113) views. Scale bar: 10 mm (108113)." captionText-2="Figures 114131: Buthus somalilandus sp. n. Figures 114123. Male. Chela dorsal (114), external (115) and ventral (116) views. Patella dorsal (117), external (118) and ventral (119) views. Trochanter and femur dorsal (120) and femur ventral (121) views. Movable (122) and fixed (123) finger dentition. Trichobothrial pattern is indicated by white circles (114118, 120). Figures 124129. Female. Chela dorsal (124), external (125) and ventral (126) views. Patella dorsal (127), external (128) and ventral (129) views. Trochanter and femur dorsal (130) and ventral (131) views." captionText-3="Figures 132139: Buthus somalilandus sp. n. Figures 132, 136139. Male holotype, carapace and tergites IIV (132), coxosternal area and sternites (134), left legs IIV, retrolateral aspect (136139 respectively). Figures 133134. Female paratopotype, carapace and tergites IIII (133), coxosternal area and sternites IIIVI (134)." figureDoi-0="http://doi.org/10.5281/zenodo.4648412" figureDoi-1="http://doi.org/10.5281/zenodo.4648414" figureDoi-2="http://doi.org/10.5281/zenodo.4648416" figureDoi-3="http://doi.org/10.5281/zenodo.4648418" httpUri-0="https://zenodo.org/record/4648412/files/figure.png" httpUri-1="https://zenodo.org/record/4648414/files/figure.png" httpUri-2="https://zenodo.org/record/4648416/files/figure.png" httpUri-3="https://zenodo.org/record/4648418/files/figure.png" pageId="18" pageNumber="17">Figures 102139</figureCitation>
,
<figureCitation id="96A32A49DC4F40096C02FD37FB275282" box="[1043,1192,714,738]" captionStart="Figures 176190" captionStartId="28.[113,188,1497,1519]" captionTargetBox="[119,1474,233,1451]" captionTargetId="figure-196@28.[113,1474,232,1495]" captionText="Figures 176190: Comparison of pdipalp chela (176, 178, 180, 182, 184), pedipalp patella (177, 179, 181, 183, 185), and telson (186190) of males of Buthus. Figures 176177, 186. B. awashensis, holotype. Figures 178179, 187. B. berberensis from locality 19SO. Figures 180181, 188. B. pococki sp. n., holotype. Figures 182183, 189. B. somalilandus sp. n., holotype. Figures 184185, 190. B. zeylensis from locality 19SH." figureDoi="http://doi.org/10.5281/zenodo.4770191" httpUri="https://zenodo.org/record/4770191/files/figure.png" pageId="18" pageNumber="17">182183, 189</figureCitation>
, 193, 210211,
<tableCitation id="431A0377DC4F40096D43FD37FA2E5282" box="[1362,1441,714,738]" captionStart="Table 2" captionStartId="23.[115,171,942,963]" captionTargetId="graphics-574@23.[119,1476,234,918]" captionTargetPageId="23" captionText="Table 2. Comparative measurements of adults of Buthus somalilandus sp. n. and B. zeylensis. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/5AE76644DC4A400C6862FC53FDAD5380" pageId="18" pageNumber="17" tableUuid="5AE76644DC4A400C6862FC53FDAD5380">Table 2</tableCitation>
)
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<emphasis id="3CECEADEDC4F40096B2AFD17FAA35342" bold="true" pageId="18" pageNumber="17">http://zoobank.org/urn:lsid:zoobank.org:act:2D740F55- 9522-4911-87A8-5119B727F228</emphasis>
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<taxonomicName id="C9984D4FDC4F40096B21FCB7FB4E5302" ID-CoL="NYYS" authority=": KovarIk, 2011: 2" authorityName=": KovarIk" authorityPageNumber="2" authorityYear="2011" box="[816,1217,842,869]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="berberensis">
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:
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, 46, figs. 14, 1823.
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LOCALITY AND
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REPOSITORY. Somaliland, near Berbera,
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11SO);
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TYPE MATERIAL
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<geoCoordinate id="6BAC500BDC4F40096B21FB77FC2254C2" box="[816,941,1162,1186]" degrees="44" direction="east" minutes="47" orientation="longitude" pageId="18" pageNumber="17" precision="15" seconds="21" value="44.789165">44°47'21&quot;E</geoCoordinate>
,
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a. s. l.
</elevation>
(
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<emphasis id="3CECEADEDC4F40096CCBFB77FA9754C2" bold="true" box="[1242,1304,1162,1186]" pageId="18" pageNumber="17">17SG</emphasis>
),
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<collectingDate id="6A62E9E4DC4F40096D21FB77FA3154C2" box="[1328,1470,1162,1186]" pageId="18" pageNumber="17" value="2018-08-21">21.VIII.2018</collectingDate>
</date>
,
</materialsCitation>
<materialsCitation id="BEF03C91DC4F40096B21FB57FB2954A5" ID-GBIF-Occurrence="3754823338" box="[816,1190,1194,1221]" collectingDate="2018-08-21" collectionCode="FKCP" collectorName="F. KovarIk &amp; Locality No." county="Berbera" elevation="60" latitude="10.037784" location="Hamas" longLatPrecision="1" longitude="44.788315" municipality="Hargeisa" pageId="18" pageNumber="17" specimenCount="11" specimenCount-female="5" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="189EFD45DC4F40096B21FB57FCD954A2" box="[816,854,1194,1218]" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
<specimenCount id="189EFD45DC4F40096B47FB57FCFC54A2" box="[854,883,1194,1218]" pageId="18" pageNumber="17" type="female">1♀</specimenCount>
(
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), leg.
<collectorName id="A36D531ADC4F40096C3CFB56FB2D54A5" box="[1069,1186,1195,1221]" pageId="18" pageNumber="17">F. KovařÍk</collectorName>
;
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<location id="0B476017DC4F40096CBCFB56FB7554A2" LSID="urn:lsid:plazi:treatment:863187DADC4F400C6BC5FD55FC6E5799:0B476017DC4F40096CBCFB56FB7554A2" box="[1197,1274,1195,1218]" county="Berbera" latitude="10.037784" longLatPrecision="1" longitude="44.788315" municipality="Hargeisa" name="Hamas" pageId="18" pageNumber="17">Hamas</location>
, between
<collectingMunicipality id="EE43ACB6DC4F40096D75FB57FA4D54A2" box="[1380,1474,1194,1218]" pageId="18" pageNumber="17">Hargeisa</collectingMunicipality>
and
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,
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<geoCoordinate id="6BAC500BDC4F40096C5CFB37FB5B5482" box="[1101,1236,1226,1250]" degrees="44" direction="east" minutes="47.299" orientation="longitude" pageId="18" pageNumber="17" precision="1" value="44.788315">44°47.299'E</geoCoordinate>
,
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<quantity id="C9609B29DC4F40096CCEFB37FAAC5482" box="[1247,1315,1226,1250]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="17" unit="m" value="650.0">650 m</quantity>
a. s. l.
</elevation>
),
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<specimenCount id="189EFD45DC4F40096DB9FB37FA4D5482" box="[1448,1474,1226,1250]" pageId="18" pageNumber="17" type="female">1♀</specimenCount>
(
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),
<date id="7A26100CDC4F40096BA9FB17FB9D5562" box="[952,1042,1258,1282]" pageId="18" pageNumber="17" value="2010-11">XI.2010</date>
, leg. T.
<location id="0B476017DC4F40096C7CFB17FB495562" LSID="urn:lsid:plazi:treatment:863187DADC4F400C6BC5FD55FC6E5799:0B476017DC4F40096C7CFB17FB495562" box="[1133,1222,1258,1282]" county="Berbera" latitude="9.859722" longLatPrecision="21" longitude="44.535557" municipality="Berbera" name="Mazuch" pageId="18" pageNumber="17">Mazuch</location>
;
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beetwen
<location id="0B476017DC4F40096D25FB17FA1D5562" LSID="urn:lsid:plazi:treatment:863187DADC4F400C6BC5FD55FC6E5799:0B476017DC4F40096D25FB17FA1D5562" box="[1332,1426,1258,1282]" county="Berbera" latitude="9.859722" longLatPrecision="21" longitude="44.535557" municipality="Berbera" name="Hargeisa" pageId="18" pageNumber="17">Hargeisa</location>
and
<collectingMunicipality id="EE43ACB6DC4F40096B21FAF7FC085542" box="[816,903,1290,1314]" pageId="18" pageNumber="17">Berbera</collectingMunicipality>
,
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<geoCoordinate id="6BAC500BDC4F40096C06FAF7FB1B5542" box="[1047,1172,1290,1314]" degrees="44" direction="east" minutes="41" orientation="longitude" pageId="18" pageNumber="17" precision="15" seconds="03" value="44.684166">44°41'03&quot;E</geoCoordinate>
,
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<quantity id="C9609B29DC4F40096CB1FAF7FB6B5542" box="[1184,1252,1290,1314]" metricMagnitude="2" metricUnit="m" metricValue="8.17" pageId="18" pageNumber="17" unit="m" value="817.0">817 m</quantity>
a. s. l.
</elevation>
(
<collectorName id="A36D531ADC4F40096D28FAF7FA4C5542" box="[1337,1475,1290,1314]" pageId="18" pageNumber="17">Locality No.</collectorName>
<emphasis id="3CECEADEDC4F40096B21FAD7FCE45522" bold="true" box="[816,875,1322,1346]" pageId="18" pageNumber="17">11SD</emphasis>
),
<date id="7A26100CDC4F40096B94FAD7FC745522" box="[901,1019,1322,1346]" pageId="18" pageNumber="17" value="2011-07-08">
<collectingDate id="6A62E9E4DC4F40096B94FAD7FC745522" box="[901,1019,1322,1346]" pageId="18" pageNumber="17" value="2011-07-08">8.VII.2011</collectingDate>
</date>
,
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<specimenCount id="189EFD45DC4F40096C18FAD7FBA45522" box="[1033,1067,1322,1346]" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
(
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), leg.
<collectorName id="A36D531ADC4F40096CF5FAD6FAD35525" box="[1252,1372,1323,1349]" pageId="18" pageNumber="17">F. KovařÍk</collectorName>
;
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<materialsCitation id="BEF03C91DC4F40096D78FAD7FC5255C5" ID-GBIF-Occurrence="3754823305" collectingDate="2018-08-21" collectingDateMax="2011-07-10" collectingDateMin="2011-07-08" collectionCode="FKCP" collectorName="F. KovarIk &amp; Locality No." county="Berbera" elevation="60" latitude="10.037784" location="Hamas" longLatPrecision="1" longitude="44.788315" municipality="Hargeisa" pageId="18" pageNumber="17" specimenCount="11" specimenCount-female="5" specimenCount-male="6" typeStatus="paratype">
beetwen
<collectingCounty id="E7464E40DC4F40096B21FAB7FC0B5502" box="[816,900,1354,1378]" pageId="18" pageNumber="17">Berbera</collectingCounty>
and Sheikh,
<geoCoordinate id="6BAC500BDC4F40096C0EFAB7FB3F5502" box="[1055,1200,1354,1378]" degrees="10" direction="north" minutes="05" orientation="latitude" pageId="18" pageNumber="17" precision="1" seconds="49.9" value="10.097194">10°05'49.9&quot;N</geoCoordinate>
<geoCoordinate id="6BAC500BDC4F40096CADFAB7FAC45502" box="[1212,1355,1354,1378]" degrees="45" direction="east" minutes="11" orientation="longitude" pageId="18" pageNumber="17" precision="1" seconds="40.1" value="45.194473">45°11'40.1&quot;E</geoCoordinate>
,
<elevation id="85B5D1FFDC4F40096D4DFAB7FCD355E2" metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" pageId="18" pageNumber="17" unit="m" value="628.0">
<quantity id="C9609B29DC4F40096D4DFAB7FA2B5502" box="[1372,1444,1354,1378]" metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" pageId="18" pageNumber="17" unit="m" value="628.0">628 m</quantity>
a. s. l.
</elevation>
(
<collectorName id="A36D531ADC4F40096B61FA97FC7055E2" box="[880,1023,1386,1410]" pageId="18" pageNumber="17">Locality No.</collectorName>
<emphasis id="3CECEADEDC4F40096C1AFA97FBC755E2" bold="true" box="[1035,1096,1386,1410]" pageId="18" pageNumber="17">11SH</emphasis>
),
<date id="7A26100CDC4F40096C75FA97FB6755E2" box="[1124,1256,1386,1410]" pageId="18" pageNumber="17" value="2011-07-10">
<collectingDate id="6A62E9E4DC4F40096C75FA97FB6755E2" box="[1124,1256,1386,1410]" pageId="18" pageNumber="17" value="2011-07-10">10.VII.2011</collectingDate>
</date>
,
<specimenCount id="189EFD45DC4F40096CE8FA97FA9E55E2" box="[1273,1297,1386,1410]" pageId="18" pageNumber="17" type="female">1♀</specimenCount>
juv. (
<typeStatus id="D123886EDC4F40096D44FA97FA3955E2" box="[1365,1462,1386,1410]" pageId="18" pageNumber="17" type="paratype">paratype</typeStatus>
), leg.
<collectorName id="A36D531ADC4F40096B70FA76FC5655C5" box="[865,985,1419,1445]" pageId="18" pageNumber="17">F. KovařÍk</collectorName>
;
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,
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<geoCoordinate id="6BAC500BDC4F40096B21FA57FC2255A2" box="[816,941,1450,1474]" degrees="44" direction="east" minutes="32" orientation="longitude" pageId="18" pageNumber="17" precision="15" seconds="08" value="44.535557">44°32'08&quot;E</geoCoordinate>
,
<elevation id="85B5D1FFDC4F40096BAAFA57FBC155A2" box="[955,1102,1450,1474]" metricMagnitude="2" metricUnit="m" metricValue="9.33" pageId="18" pageNumber="17" unit="m" value="933.0">
<quantity id="C9609B29DC4F40096BAAFA57FB8E55A2" box="[955,1025,1450,1474]" metricMagnitude="2" metricUnit="m" metricValue="9.33" pageId="18" pageNumber="17" unit="m" value="933.0">933 m</quantity>
a. s. l.
</elevation>
(
<collectorName id="A36D531ADC4F40096C4EFA57FB6455A2" box="[1119,1259,1450,1474]" pageId="18" pageNumber="17">Locality No.</collectorName>
<emphasis id="3CECEADEDC4F40096CE5FA57FAB955A2" bold="true" box="[1268,1334,1450,1474]" pageId="18" pageNumber="17">18SM</emphasis>
),
<date id="7A26100CDC4F40096D5EFA57FA3155A2" box="[1359,1470,1450,1474]" pageId="18" pageNumber="17" value="2018-09-02">2.IX.2018</date>
,
<specimenCount id="189EFD45DC4F40096B21FA37FCDD5582" box="[816,850,1482,1506]" pageId="18" pageNumber="17" type="female">1♀</specimenCount>
(
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), leg. F. KovařÍk.
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</paragraph>
</subSubSection>
<subSubSection id="46826547DC4F40096B21F9F7FAD65642" box="[816,1369,1546,1571]" pageId="18" pageNumber="17" type="etymology">
<paragraph id="0E2736CCDC4F40096B21F9F7FAD65642" blockId="18.[816,1369,1546,1571]" box="[816,1369,1546,1571]" pageId="18" pageNumber="17">ETYMOLOGY. Named after the country of occurence.</paragraph>
</subSubSection>
<subSubSection id="46826547DC4F40096B21F9B7FA915702" pageId="18" pageNumber="17" type="diagnosis">
<paragraph id="0E2736CCDC4F40096B21F9B7FA915702" blockId="18.[816,1474,1610,1890]" pageId="18" pageNumber="17">
DIAGNOSIS. Total length
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. Base color yellow dark pattern reduced; telson orange; chelicerae pale yellow without reticulation. Pedipalp movable fingers bear 1011 rows of granules, with 11 outer and inner denticles and fixed finger with 9 outer and 10 inner denticles. Chela of pedipalp narrower in male, its length to width ratio 3.63.9 in females and 4.24.4 in males. Telson bulbous, with aculeus shorter than vesicle. Pectinal teeth number
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females and
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males.
</paragraph>
</subSubSection>
<subSubSection id="46826547DC4F400C6B21F877FBE85779" lastPageId="23" lastPageNumber="22" pageId="18" pageNumber="17" type="description">
<paragraph id="0E2736CCDC4F400C6B21F877FEB75459" blockId="18.[816,1474,1930,2050]" lastBlockId="23.[113,772,1025,2041]" lastPageId="23" lastPageNumber="22" pageId="18" pageNumber="17">
DESCRIPTION. The adults are
<quantity id="C9609B29DC4F40096C74F877FB2057C2" box="[1125,1199,1930,1954]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="18" pageNumber="17" unit="mm" value="42.0">42 mm</quantity>
(male)
<quantity id="C9609B29DC4F40096D07F877FAEE57C2" box="[1302,1377,1930,1954]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="17" unit="mm" value="55.0">55 mm</quantity>
(female) long. The habitus is shown in
<figureCitation id="96A32A49DC4F40096C7BF857FA8B57A2" box="[1130,1284,1962,1986]" captionStart="Figures 102105" captionStartId="19.[113,190,1973,1995]" captionTargetBox="[133,1468,266,1933]" captionTargetId="figure-49@19.[113,1474,248,1939]" captionTargetPageId="19" captionText="Figures 102105: Buthus somalilandus sp. n. Figures 102103. Male in dorsal (102) and ventral (103) views. Figures 104105. Female in dorsal (104) and ventral (105) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648412" httpUri="https://zenodo.org/record/4648412/files/figure.png" pageId="18" pageNumber="17">Figs. 102105</figureCitation>
,
<figureCitation id="96A32A49DC4F40096D1CF857FAB757A2" box="[1293,1336,1962,1986]" captionStart="Figures 191194" captionStartId="29.[113,190,2007,2029]" captionTargetBox="[113,1474,221,1997]" captionTargetId="figure-9@29.[113,1474,218,2000]" captionTargetPageId="29" captionText="Figures 191194: Buthus, in vivo habitus. Figure 191. B. awashensis, female paratype. Figure 192. B. pococki sp. n., male paratype. Figure 193. B. somalilandus sp. n., female, paratype. Figure 194. B. zeylensis from locality 19SH." figureDoi="http://doi.org/10.5281/zenodo.4648428" httpUri="https://zenodo.org/record/4648428/files/figure.png" pageId="18" pageNumber="17">193</figureCitation>
. For position and distribution of trichobothria of pedipalps see
<figureCitation id="96A32A49DC4F40096D41F837FCD65862" captionStart="Figures 114131" captionStartId="21.[113,190,1904,1926]" captionTargetBox="[113,1465,243,1889]" captionTargetId="figure-8@21.[113,1474,234,1903]" captionTargetPageId="21" captionText="Figures 114131: Buthus somalilandus sp. n. Figures 114123. Male. Chela dorsal (114), external (115) and ventral (116) views. Patella dorsal (117), external (118) and ventral (119) views. Trochanter and femur dorsal (120) and femur ventral (121) views. Movable (122) and fixed (123) finger dentition. Trichobothrial pattern is indicated by white circles (114118, 120). Figures 124129. Female. Chela dorsal (124), external (125) and ventral (126) views. Patella dorsal (127), external (128) and ventral (129) views. Trochanter and femur dorsal (130) and ventral (131) views." figureDoi="http://doi.org/10.5281/zenodo.4648416" httpUri="https://zenodo.org/record/4648416/files/figure.png" pageId="18" pageNumber="17">Figs. 114 118</figureCitation>
, and 120. Sexual dimorphism minor: adult males with pedipalp chela broader. Female with a wide basal middle lamella (
<figureCitation id="96A32A49DC4A400C68DEFBDCFEA35459" box="[207,300,1057,1081]" captionStart="Figures 132139" captionStartId="22.[115,192,1958,1980]" captionTargetBox="[113,1474,250,1884]" captionTargetId="figure-18@22.[113,1474,250,1920]" captionTargetPageId="22" captionText="Figures 132139: Buthus somalilandus sp. n. Figures 132, 136139. Male holotype, carapace and tergites IIV (132), coxosternal area and sternites (134), left legs IIV, retrolateral aspect (136139 respectively). Figures 133134. Female paratopotype, carapace and tergites IIII (133), coxosternal area and sternites IIIVI (134)." figureDoi="http://doi.org/10.5281/zenodo.4648418" httpUri="https://zenodo.org/record/4648418/files/figure.png" pageId="23" pageNumber="22">Fig. 134</figureCitation>
).
</paragraph>
<paragraph id="0E2736CCDC4A400C6860FBBDFDEA54D9" blockId="23.[113,772,1025,2041]" pageId="23" pageNumber="22">
<emphasis id="3CECEADEDC4A400C6860FBBDFF635438" bold="true" box="[113,236,1088,1112]" pageId="23" pageNumber="22">Coloration</emphasis>
(
<figureCitation id="96A32A49DC4A400C68EBFBBCFE195438" box="[250,406,1088,1113]" captionStart="Figures 102105" captionStartId="19.[113,190,1973,1995]" captionTargetBox="[133,1468,266,1933]" captionTargetId="figure-49@19.[113,1474,248,1939]" captionTargetPageId="19" captionText="Figures 102105: Buthus somalilandus sp. n. Figures 102103. Male in dorsal (102) and ventral (103) views. Figures 104105. Female in dorsal (104) and ventral (105) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648412" httpUri="https://zenodo.org/record/4648412/files/figure.png" pageId="23" pageNumber="22">Figs. 102105</figureCitation>
,
<figureCitation id="96A32A49DC4A400C69B0FBBCFE435439" box="[417,460,1089,1113]" captionStart="Figures 191194" captionStartId="29.[113,190,2007,2029]" captionTargetBox="[113,1474,221,1997]" captionTargetId="figure-9@29.[113,1474,218,2000]" captionTargetPageId="29" captionText="Figures 191194: Buthus, in vivo habitus. Figure 191. B. awashensis, female paratype. Figure 192. B. pococki sp. n., male paratype. Figure 193. B. somalilandus sp. n., female, paratype. Figure 194. B. zeylensis from locality 19SH." figureDoi="http://doi.org/10.5281/zenodo.4648428" httpUri="https://zenodo.org/record/4648428/files/figure.png" pageId="23" pageNumber="22">193</figureCitation>
). The base color is uniformly yellow with dark pattern reduced in juveniles and often absent in adults, tergites yellow. The pedipalps and legs are yellow without dark carina in adults. Telson is orange.
</paragraph>
<paragraph id="0E2736CCDC4A400C6860FB3DFF515639" blockId="23.[113,772,1025,2041]" pageId="23" pageNumber="22">
<emphasis id="3CECEADEDC4A400C6860FB3DFE1654B9" bold="true" box="[113,409,1216,1241]" pageId="23" pageNumber="22">Carapace and mesosoma</emphasis>
(
<figureCitation id="96A32A49DC4A400C69BCFB3CFDDF54B8" box="[429,592,1216,1241]" captionStart="Figures 102105" captionStartId="19.[113,190,1973,1995]" captionTargetBox="[133,1468,266,1933]" captionTargetId="figure-49@19.[113,1474,248,1939]" captionTargetPageId="19" captionText="Figures 102105: Buthus somalilandus sp. n. Figures 102103. Male in dorsal (102) and ventral (103) views. Figures 104105. Female in dorsal (104) and ventral (105) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648412" httpUri="https://zenodo.org/record/4648412/files/figure.png" pageId="23" pageNumber="22">Figs. 102105</figureCitation>
,
<figureCitation id="96A32A49DC4A400C6A71FB3DFD4D54B8" box="[608,706,1216,1240]" captionStart="Figures 132139" captionStartId="22.[115,192,1958,1980]" captionTargetBox="[113,1474,250,1884]" captionTargetId="figure-18@22.[113,1474,250,1920]" captionTargetPageId="22" captionText="Figures 132139: Buthus somalilandus sp. n. Figures 132, 136139. Male holotype, carapace and tergites IIV (132), coxosternal area and sternites (134), left legs IIV, retrolateral aspect (136139 respectively). Figures 133134. Female paratopotype, carapace and tergites IIII (133), coxosternal area and sternites IIIVI (134)." figureDoi="http://doi.org/10.5281/zenodo.4648418" httpUri="https://zenodo.org/record/4648418/files/figure.png" pageId="23" pageNumber="22">132135</figureCitation>
). The entire carapace is covered with granules small along much of the middline and large anteriorly, posteriorly and laterally. The carinae are typical for genus
<taxonomicName id="C9984D4FDC4A400C69CFFADCFDA75559" authorityName="Leach" authorityYear="1815" box="[478,552,1313,1337]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="3CECEADEDC4A400C69CFFADCFDA75559" box="[478,552,1313,1337]" italics="true" pageId="23" pageNumber="22">Buthus</emphasis>
</taxonomicName>
, strongly developed and composed of large, round granules. The anterior margin of the carapace is straight, and bears 1012 long, symmetrically placed macrosetae. The tergites are granulated in posterior parts and bear very strong median carinae. A pair of denticulate lateral carinae is strong on tergites IIVI. Tergite VII is pentacarinate, with all carinae strong and granulated. The pectinal tooth count is 2326 (1x23, 6x24, 6x25, 1x26) in the females and 3035 (2x30, 4x31, 5x33, 3x34, 2x35) in the males.
</paragraph>
<paragraph id="0E2736CCDC4A400C6860F99CFEA15739" blockId="23.[113,772,1025,2041]" pageId="23" pageNumber="22">The pectinal marginal tips extend to the end of the fourth sternite or to the quoter of the fifth sternite in males and to the end of the third sternite in females. The pectines have three marginal lamellae and seven or eight middle lamellae. The lamellae bear numerous dark setae, each fulcrum with three to six dark setae. All sternites are smooth or finely granulated. The seventh sternite bears four strongly developed ventral crenulate carinae.</paragraph>
<paragraph id="0E2736CCDC4A400C6860F89CFBC85579" blockId="23.[113,772,1025,2041]" lastBlockId="23.[816,1475,1025,1817]" pageId="23" pageNumber="22">
<emphasis id="3CECEADEDC4A400C6860F89CFEE75719" bold="true" box="[113,360,1889,1913]" pageId="23" pageNumber="22">Metasoma and telson</emphasis>
(
<figureCitation id="96A32A49DC4A400C6966F89CFD9A5718" box="[375,533,1888,1913]" captionStart="Figures 106113" captionStartId="20.[116,193,1958,1980]" captionTargetBox="[134,1471,244,1910]" captionTargetId="figure-81@20.[114,1475,234,1936]" captionTargetPageId="20" captionText="Figures 106113: Buthus somalilandus sp. n. Figures 106, 108110. Male, telson lateral (106), and metasoma and telson lateral (108), dorsal (109), and ventral (111) views. Figures 107, 111113. Female, telson lateral (107), and metasoma and telson lateral (111), dorsal (112), and ventral (113) views. Scale bar: 10 mm (108113)." figureDoi="http://doi.org/10.5281/zenodo.4648414" httpUri="https://zenodo.org/record/4648414/files/figure.png" pageId="23" pageNumber="22">Figs. 106113</figureCitation>
). The first metasomal segment bears 10 carinae, the second throught fourth segments bear eight to 10 carinae (the lateral surface of the second and third segments bears a row of granules that exceeds three-quarters of the second and one-half of the third segment length). The fourth segment bear eight carinae and the fifth segment bears five carinae. The ventrolateral carinae of the fifth segment terminate in two lobes. The ventral keels on the second and third segments posteriorly bear two to four large granules. The surface between the carinae is smooth. All segments are sparsely setose. The telson is bulbous, with aculeus shorter than the vesicle. The surface of the telson is unevenly granulated and bears an incomplete lateral carina.
</paragraph>
<paragraph id="0E2736CCDC4A400C6B21FADDFB255679" blockId="23.[816,1475,1025,1817]" pageId="23" pageNumber="22">
<emphasis id="3CECEADEDC4A400C6B21FADDFC105558" bold="true" box="[816,927,1312,1336]" pageId="23" pageNumber="22">Pedipalps</emphasis>
(
<figureCitation id="96A32A49DC4A400C6BA7FADCFBD55559" box="[950,1114,1313,1337]" captionStart="Figures 114131" captionStartId="21.[113,190,1904,1926]" captionTargetBox="[113,1465,243,1889]" captionTargetId="figure-8@21.[113,1474,234,1903]" captionTargetPageId="21" captionText="Figures 114131: Buthus somalilandus sp. n. Figures 114123. Male. Chela dorsal (114), external (115) and ventral (116) views. Patella dorsal (117), external (118) and ventral (119) views. Trochanter and femur dorsal (120) and femur ventral (121) views. Movable (122) and fixed (123) finger dentition. Trichobothrial pattern is indicated by white circles (114118, 120). Figures 124129. Female. Chela dorsal (124), external (125) and ventral (126) views. Patella dorsal (127), external (128) and ventral (129) views. Trochanter and femur dorsal (130) and ventral (131) views." figureDoi="http://doi.org/10.5281/zenodo.4648416" httpUri="https://zenodo.org/record/4648416/files/figure.png" pageId="23" pageNumber="22">Figs. 114131</figureCitation>
). The pedipalps are smooth or finely granulated. The femur bears four to five carinae; the ventroexternal carina is incomplete, the other carinae are coarsely granular. The patella bears seven coarsely granular carinae. The chela bears five carinae, which may be weak and incomplete. The movable and fixed fingers bear 1011 rows of granules, with 11 outer and inner denticles and fixed finger with 9 outer and 10 inner denticles.
</paragraph>
<paragraph id="0E2736CCDC4A400C6B21F9DCFABC5698" blockId="23.[816,1475,1025,1817]" pageId="23" pageNumber="22">
<emphasis id="3CECEADEDC4A400C6B21F9DCFCEA5659" bold="true" box="[816,869,1569,1593]" pageId="23" pageNumber="22">Legs</emphasis>
(
<figureCitation id="96A32A49DC4A400C6B64F9DCFB9E5658" box="[885,1041,1568,1593]" captionStart="Figures 132139" captionStartId="22.[115,192,1958,1980]" captionTargetBox="[113,1474,250,1884]" captionTargetId="figure-18@22.[113,1474,250,1920]" captionTargetPageId="22" captionText="Figures 132139: Buthus somalilandus sp. n. Figures 132, 136139. Male holotype, carapace and tergites IIV (132), coxosternal area and sternites (134), left legs IIV, retrolateral aspect (136139 respectively). Figures 133134. Female paratopotype, carapace and tergites IIII (133), coxosternal area and sternites IIIVI (134)." figureDoi="http://doi.org/10.5281/zenodo.4648418" httpUri="https://zenodo.org/record/4648418/files/figure.png" pageId="23" pageNumber="22">Figs. 136139</figureCitation>
). Pairs III and IV bear long tibial spurs. Retrolateral and prolateral pedal spurs are present on all legs. The tarsomeres bear two rows of macrosetae on the ventral surface and numerous macrosetae on the other surfaces. Bristlecombs are present on the first to third legs. The femur bears four carinae and the patella bears four to six carinae. The femur and patella bear only solitary macrosetae.
</paragraph>
<paragraph id="0E2736CCDC4A400C6B21F8FCFBE85779" blockId="23.[816,1475,1025,1817]" box="[816,1127,1793,1817]" pageId="23" pageNumber="22">
<emphasis id="3CECEADEDC4A400C6B21F8FCFC585779" bold="true" box="[816,983,1793,1817]" pageId="23" pageNumber="22">Measurements</emphasis>
. See
<tableCitation id="431A0377DC4A400C6C00F8FCFBEC5779" box="[1041,1123,1793,1817]" captionStart="Table 2" captionStartId="23.[115,171,942,963]" captionTargetId="graphics-574@23.[119,1476,234,918]" captionTargetPageId="23" captionText="Table 2. Comparative measurements of adults of Buthus somalilandus sp. n. and B. zeylensis. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/5AE76644DC4A400C6862FC53FDAD5380" pageId="23" pageNumber="22" tableUuid="5AE76644DC4A400C6862FC53FDAD5380">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46826547DC4A400C6B21F8BCFC6E5799" pageId="23" pageNumber="22" type="discussion">
<paragraph id="0E2736CCDC4A400C6B21F8BCFC6E5799" blockId="23.[816,1474,1857,2041]" pageId="23" pageNumber="22">
AFFINITIES. The described features distinguish
<taxonomicName id="C9984D4FDC4A400C6D09F8BCFA4D5739" authority="Kovařík &amp; Šťáhlavský &amp; Elmi, 2020" authorityName="Kovařík &amp; Šťáhlavský &amp; Elmi" authorityYear="2020" box="[1304,1474,1857,1881]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="somalilandus" status="sp. nov.">
<emphasis id="3CECEADEDC4A400C6D09F8BCFA4D5739" box="[1304,1474,1857,1881]" italics="true" pageId="23" pageNumber="22">B. somalilandus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27DF57A5DC4A400C6B21F89CFCE25718" box="[816,877,1889,1913]" pageId="23" pageNumber="22" rank="species">
<emphasis id="3CECEADEDC4A400C6B21F89CFCC65719" bold="true" box="[816,841,1889,1913]" pageId="23" pageNumber="22">sp</emphasis>
.
<emphasis id="3CECEADEDC4A400C6B46F89CFCE95719" bold="true" box="[855,870,1889,1913]" pageId="23" pageNumber="22">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. They are recounted in the key below.
<taxonomicName id="C9984D4FDC4A400C6C17F87CFB3757F9" authority="Kovařík &amp; Šťáhlavský &amp; Elmi, 2020" authorityName="Kovařík &amp; Šťáhlavský &amp; Elmi" authorityYear="2020" box="[1030,1208,1921,1945]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="somalilandus" status="sp. nov.">
<emphasis id="3CECEADEDC4A400C6C17F87CFB3757F9" box="[1030,1208,1921,1945]" italics="true" pageId="23" pageNumber="22">B. somalilandus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27DF57A5DC4A400C6CD4F87CFA8857F8" box="[1221,1287,1921,1945]" pageId="23" pageNumber="22" rank="species">
<emphasis id="3CECEADEDC4A400C6CD4F87CFB5157F9" bold="true" box="[1221,1246,1921,1945]" pageId="23" pageNumber="22">sp</emphasis>
.
<emphasis id="3CECEADEDC4A400C6CE0F87CFA8F57F9" bold="true" box="[1265,1280,1921,1945]" pageId="23" pageNumber="22">n</emphasis>
.
</taxonomicNameLabel>
is similar to
<taxonomicName id="C9984D4FDC4A400C6DBAF87CFC0057D9" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="zeylensis">
<emphasis id="3CECEADEDC4A400C6DBAF87CFC0057D9" italics="true" pageId="23" pageNumber="22">B. zeylensis</emphasis>
</taxonomicName>
. It is possible to mainly differentiate these two species mainly according to pedipalp movable finger dentition (see key below).
</paragraph>
</subSubSection>
</treatment>
</document>