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<document id="EB844132CB539CD934BEF77A2C002CEA" ID-CLB-Dataset="6758" ID-DOI="10.5281/zenodo.4648379" ID-GBIF-Dataset="2883d728-9246-4b55-b0be-2b774fedcc7d" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648379" ID-ZooBank="881B2577-F367-469A-AB81-8AF742E7D00C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617135269639" checkinUser="carolina" docAuthor="Kovařík, František, Šťáhlavský, František &amp; Elmi, Hassan Sh Abdirahman" docDate="2020" docId="863187DADC41403B68DEF9B9FE9C55F2" docLanguage="en" docName="Euscorpius.307.1-32.pdf" docOrigin="Euscorpius 307" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Buthus zeylensis Pocock 1900" docType="treatment" docVersion="11" lastPageNumber="31" masterDocId="7A08FFA2DC5D401B6811FFFDFF8F5060" masterDocTitle="Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXIII Buthus (Buthidae), with description of two new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="27" updateTime="1698935455889" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="12B72B00E4C5FCC9D99FFFD44AACC08A">Scorpions of the Horn of Africa (Arachnida Scorpiones) Part XXIII Buthus (Buthidae), with description of two new species</mods:title>
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<mods:namePart id="C03C89043E224D582C48BD7349A76BD2">Kovařík, František</mods:namePart>
<mods:affiliation id="7551096E74E2AB4D4FAD66EA278BCDB9">Department of Zoology, Charles University, ViničnÁ 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="11894B113ACB6B03371212C90457145C">Šťáhlavský, František</mods:namePart>
<mods:affiliation id="E9A81824AA2713C6828DED1608A6B1E6">Department of Zoology, Charles University, ViničnÁ 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="2C038D0CDC777B882EDD0D8B58C600D9">Elmi, Hassan Sh Abdirahman</mods:namePart>
<mods:affiliation id="8DF5A67C9218B704EC00641ED4DC34CB">Amoud University, Borama, Republic of Somaliland</mods:affiliation>
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<mods:date id="2B700BEBAB1590E4D83E169EDEAA24AA">2020</mods:date>
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<treatment id="863187DADC41403B68DEF9B9FE9C55F2" ID-DOI="http://doi.org/10.5281/zenodo.4770197" ID-GBIF-Taxon="182215989" ID-Zenodo-Dep="4770197" LSID="urn:lsid:plazi:treatment:863187DADC41403B68DEF9B9FE9C55F2" httpUri="http://treatment.plazi.org/id/863187DADC41403B68DEF9B9FE9C55F2" lastPageId="32" lastPageNumber="31" pageId="28" pageNumber="27">
<subSubSection id="46826547DC41400768DEF9B9FD2C563E" box="[207,675,1604,1631]" pageId="28" pageNumber="27" type="nomenclature">
<paragraph id="0E2736CCDC41400768DEF9B9FD2C563E" blockId="28.[130,752,1604,1694]" box="[207,675,1604,1631]" pageId="28" pageNumber="27">
<heading id="556F81A0DC41400768DEF9B9FD2C563E" box="[207,675,1604,1631]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="27" reason="2">
<taxonomicName id="C9984D4FDC41400768DEF9B9FDCD563E" ID-CoL="NZ2C" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[207,578,1604,1631]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="zeylensis">
<emphasis id="3CECEADEDC41400768DEF9B9FE16563F" bold="true" box="[207,409,1604,1631]" italics="true" pageId="28" pageNumber="27">Buthus zeylensis</emphasis>
<bibRefCitation id="6A094B3DDC41400769B1F9B9FDCD563E" author="Pocock" box="[416,578,1604,1630]" firstAuthor="Pocock" pageId="28" pageNumber="27" pagination="55 - 63" refId="ref10359" refString="POCOCK, R. I. 1900. On a collection of insects and arachnids made in 1895 and 1897 by Mr. C. A. V. Peel, F. Z. S. in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somaliland and the Boran Country. Proceedings of the Zoological Society of London, 1900: 55 - 63." type="book chapter" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
,
<emphasis id="3CECEADEDC4140076A5FF9B9FD2C563E" bold="true" box="[590,675,1604,1630]" pageId="28" pageNumber="27">stat. n.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="46826547DC4140076893F99BFE7856FE" pageId="28" pageNumber="27" type="description">
<paragraph id="0E2736CCDC4140076893F99BFD7F561E" blockId="28.[130,752,1604,1694]" box="[130,752,1638,1662]" pageId="28" pageNumber="27">
(
<figureCitation id="96A32A49DC4140076898F99BFE84561E" box="[137,267,1638,1662]" captionStart="Figures 100–101" captionStartId="17.[113,190,2013,2035]" captionTargetBox="[113,1474,234,1995]" captionTargetId="figure-35@17.[113,1474,234,1998]" captionTargetPageId="17" captionText="Figures 100–101: Localities. Figure 100. Buthus pococki sp. n., type locality (19SC). Figure 101. B. zeylensis, locality 19SH." figureDoi="http://doi.org/10.5281/zenodo.4648410" httpUri="https://zenodo.org/record/4648410/files/figure.png" pageId="28" pageNumber="27">Figures 101</figureCitation>
,
<figureCitation id="96A32A49DC4140076906F99BFEF8561E" box="[279,375,1638,1662]" captionStart-0="Figures 140–143" captionStart-1="Figures 144–151" captionStart-2="Figures 152–167" captionStart-3="Figures 168–175" captionStartId-0="24.[116,193,1979,2001]" captionStartId-1="25.[115,192,1967,1989]" captionStartId-2="26.[115,192,1928,1950]" captionStartId-3="27.[113,190,1962,1984]" captionTargetBox-0="[125,1450,286,1924]" captionTargetBox-1="[126,1461,239,1956]" captionTargetBox-2="[113,1455,237,1909]" captionTargetBox-3="[113,1452,234,1947]" captionTargetId-0="figure-20@24.[113,1474,266,1952]" captionTargetId-1="figure-8@25.[113,1474,231,1960]" captionTargetId-2="figure-19@26.[113,1474,234,1959]" captionTargetId-3="figure-9@27.[113,1474,234,1950]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionText-0="Figures 140–143: Buthus zeylensis from locality 19SH. Figures 140–141. Male in dorsal (140) and ventral (141) views. Figures 142–143. Female in dorsal (142) and ventral (143) views. Scale bar: 10 mm." captionText-1="Figures 144–151: Buthus zeylensis from locality 19SH. Figures 144, 146–148. Male, telson lateral (144), and metasoma and telson lateral (146), dorsal (147), and ventral (148) views. Figures 145, 149–151. Female, telson lateral (145), and metasoma and telson lateral (149), dorsal (150), and ventral (151) views. Scale bar: 10 mm (146–151)." captionText-2="Figures 152–167: Buthus zeylensis from locality 19SH, pedipalp segments. Figures 152–161. Male. Chela dorsal (152), external (153) and ventral (154) views. Patella dorsal (155), external (156) and ventral (157) views. Trochanter and femur dorsal (158) and ventral (159) views. Movable (160) and fixed (161) finger dentition. Trichobothrial pattern is indicated by white circles (152–156, 158). Figures 162–167. Female. Chela dorsal (162), external (163) and ventral (164) views. Patella dorsal (165), external (166) and ventral (167) views." captionText-3="Figures 168–175: Buthus zeylensis from locality 19SH. Figures 168, 170, 172–175. Male, carapace and tergites I–IV (168), coxosternal area and sternites (170), left legs I–IV, retrolateral aspect (172–175 respectively). Figures 169, 171. Female, carapace and tergites I–III (169), coxosternal area and sternites III–VI (171)." figureDoi-0="http://doi.org/10.5281/zenodo.4648420" figureDoi-1="http://doi.org/10.5281/zenodo.4648422" figureDoi-2="http://doi.org/10.5281/zenodo.4648424" figureDoi-3="http://doi.org/10.5281/zenodo.4648426" httpUri-0="https://zenodo.org/record/4648420/files/figure.png" httpUri-1="https://zenodo.org/record/4648422/files/figure.png" httpUri-2="https://zenodo.org/record/4648424/files/figure.png" httpUri-3="https://zenodo.org/record/4648426/files/figure.png" pageId="28" pageNumber="27">140–175</figureCitation>
,
<figureCitation id="96A32A49DC4140076990F99BFD99561E" box="[385,534,1638,1662]" captionStart="Figures 176–190" captionStartId="28.[113,188,1497,1519]" captionTargetBox="[119,1474,233,1451]" captionTargetId="figure-196@28.[113,1474,232,1495]" captionText="Figures 176–190: Comparison of pdipalp chela (176, 178, 180, 182, 184), pedipalp patella (177, 179, 181, 183, 185), and telson (186–190) of males of Buthus. Figures 176–177, 186. B. awashensis, holotype. Figures 178–179, 187. B. berberensis from locality 19SO. Figures 180–181, 188. B. pococki sp. n., holotype. Figures 182–183, 189. B. somalilandus sp. n., holotype. Figures 184–185, 190. B. zeylensis from locality 19SH." figureDoi="http://doi.org/10.5281/zenodo.4770191" httpUri="https://zenodo.org/record/4770191/files/figure.png" pageId="28" pageNumber="27">184–185, 190</figureCitation>
, 194,
<figureCitation id="96A32A49DC4140076A46F99BFD38561E" box="[599,695,1638,1662]" captionStart="Figures 195–209" captionStartId="30.[113,190,1852,1874]" captionTargetBox="[181,1400,234,1839]" captionTargetId="figure-20@30.[159,1436,234,1839]" captionText="Figures 195–209: Male mitotic metaphases (195, 198, 201, 204, 207), postpachytenes (196, 199, 202, 205, 208) and ideograms (197, 200, 203, 206, 206) (y axis - % of the diploid chromosome length, dark grey marks chromosomes in multivalent association) of Buthus species from the Horn of Africa. Figures 195–197. B. awashensis, topotype from locality 12EX (2n=22, 6II+CX), Figures 198–200. B. berberensis from locality 19SO (2n=21, 9II+CIII), Figures 201–203. B. pococki sp. n., paratopotype (2n=22, 11II), Figures 204–206. B. pococki sp. n., paratype from locality 19SN (2n=22, 11II), Figures 207–209. B. zeylensis from locality 19SH (2n=22, 11II). Arrows show chromosomes in multivalent association during postpachytene. Asterisks show extra-large chromosome from multivalents. Arrowheads indicate large chromosome pair. Scale bar: 10 μm (195–196, 198–199, 201–202, 204–205, 207–208)." figureDoi="http://doi.org/10.5281/zenodo.4770195" httpUri="https://zenodo.org/record/4770195/files/figure.png" pageId="28" pageNumber="27">207–209</figureCitation>
, 210,
</paragraph>
<paragraph id="0E2736CCDC414007696AF97BFE7856FE" blockId="28.[130,752,1604,1694]" box="[379,503,1670,1694]" pageId="28" pageNumber="27">
<tableCitation id="431A0377DC414007696AF97BFE7F56FE" box="[379,496,1670,1694]" captionStart-0="Table 2" captionStart-1="Table 3" captionStartId-0="23.[115,171,942,963]" captionStartId-1="32.[115,171,447,468]" captionTargetId-0="graphics-574@23.[119,1476,234,918]" captionTargetId-1="graphics-722@32.[113,1474,244,422]" captionTargetPageId-0="23" captionTargetPageId-1="32" captionText-0="Table 2. Comparative measurements of adults of Buthus somalilandus sp. n. and B. zeylensis. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 3. The diploid numbers, the postpachytene configuration (Ppt configuration), the percentage of the total chromosome length of the diploid set (% TCL) and the localities of Buthus species from the Horn of Africa." httpUri-0="http://table.plazi.org/id/5AE76644DC4A400C6862FC53FDAD5380" httpUri-1="http://table.plazi.org/id/5AE76644DC7D403B6862FE42FCF25192" pageId="28" pageNumber="27" tableUuid-0="5AE76644DC4A400C6862FC53FDAD5380" tableUuid-1="5AE76644DC7D403B6862FE42FCF25192">Tables 2–3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="46826547DC4140076861F931FE9057E4" pageId="28" pageNumber="27" type="reference_group">
<paragraph id="0E2736CCDC4140076861F931FDF35684" blockId="28.[112,770,1740,1924]" box="[112,636,1740,1764]" pageId="28" pageNumber="27">
<treatmentCitationGroup id="2E8811E2DC4140076861F931FDF35684" box="[112,636,1740,1764]" pageId="28" pageNumber="27">
<treatmentCitation id="8F3910DDDC4140076861F931FDF75684" author="POCOCK, R. I. " box="[112,632,1740,1764]" page="57" pageId="28" pageNumber="27" year="1900">
<taxonomicName id="C9984D4FDC4140076861F931FDF75684" ID-CoL="NZ2C" authority="Pocock, 1900: 56 - 57" authorityName="Pocock" authorityPageNumber="56 - 57" authorityYear="1900" box="[112,632,1740,1764]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="zeylensis">
<emphasis id="3CECEADEDC4140076861F931FE025684" box="[112,397,1740,1764]" italics="true" pageId="28" pageNumber="27">Buthus occitanus zeylensis</emphasis>
<bibRefCitation id="6A094B3DDC4140076982F931FDF75684" author="POCOCK, R. I. " box="[403,632,1740,1764]" firstAuthor="Pocock" page="56 - 57" pageId="28" pageNumber="27" pagination="55 - 63" refId="ref10359" refString="POCOCK, R. I. 1900. On a collection of insects and arachnids made in 1895 and 1897 by Mr. C. A. V. Peel, F. Z. S. in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somaliland and the Boran Country. Proceedings of the Zoological Society of London, 1900: 55 - 63." type="book chapter" year="1900">Pocock, 1900: 56–57</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0E2736CCDC4140076861F911FE925724" blockId="28.[112,770,1740,1924]" pageId="28" pageNumber="27">
<treatmentCitationGroup id="2E8811E2DC4140076861F911FE925724" pageId="28" pageNumber="27">
<taxonomicName id="C9984D4FDC4140076861F911FE265764" ID-CoL="NYYS" authorityName="Pocock" authorityYear="1900" box="[112,425,1772,1796]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="berberensis">
<emphasis id="3CECEADEDC4140076861F911FE265764" box="[112,425,1772,1796]" italics="true" pageId="28" pageNumber="27">Buthus occitanus berberensis</emphasis>
</taxonomicName>
(in part):
<treatmentCitation id="8F3910DDDC4140076A07F910FF675744" author="LEVY, G. &amp; P. AMITAI" page="16, 21" pageId="28" pageNumber="27" year="1980">
<bibRefCitation id="6A094B3DDC4140076A07F910FF325744" author="LEVY, G. &amp; P. AMITAI" bookContentInfo="132 pp." journalOrPublisher="Jerusalem: The Israel Academy of Sciences and Humanities" pageId="28" pageNumber="27" refId="ref10220" refString="LEVY, G. &amp; P. AMITAI. 1980. Fauna Palaestina, Arachnida I. - Scorpiones. Jerusalem: The Israel Academy of Sciences and Humanities, 132 pp." title="Fauna Palaestina, Arachnida I. - Scorpiones." type="book" year="1980">Levy &amp; Amitai, 1980: 16</bibRefCitation>
, 21
</treatmentCitation>
;
<treatmentCitation id="8F3910DDDC41400768E7F8F0FE625744" author="FET, V. &amp; G. LOWE" box="[246,493,1804,1828]" page="95" pageId="28" pageNumber="27" year="2000">
<bibRefCitation id="6A094B3DDC41400768E7F8F0FE625744" author="FET, V. &amp; G. LOWE" bookContentInfo="689 pp." box="[246,493,1804,1828]" editor="Fet, V. &amp; W. D. Sissom &amp; G. Lowe &amp; M. E. Braunwalder" journalOrPublisher="New York: The New York Entomological Society" pageId="28" pageNumber="27" pagination="54 - 286" refId="ref9759" refString="FET, V. &amp; G. LOWE. 2000. Family Buthidae C. L. Koch, 1837. Pp. 54 - 286 in Fet, V., W. D. Sissom, G. Lowe &amp; M. E. Braunwalder. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." title="Family Buthidae C. L. Koch, 1837." type="book chapter" volumeTitle="Catalog of the Scorpions of the World (1758 - 1998)" year="2000">Fet &amp; Lowe, 2000: 95</bibRefCitation>
</treatmentCitation>
(complete references list until 1998).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0E2736CCDC4140076861F8B1FE9057E4" blockId="28.[112,770,1740,1924]" pageId="28" pageNumber="27">
<treatmentCitationGroup id="2E8811E2DC4140076861F8B1FE9057E4" pageId="28" pageNumber="27">
<treatmentCitation id="8F3910DDDC4140076861F8B1FDA65704" author="LOURENCO, W. R. " box="[112,553,1868,1892]" page="46" pageId="28" pageNumber="27" year="2008">
<taxonomicName id="C9984D4FDC4140076861F8B1FDA65704" ID-CoL="NYYS" authority=": Lourenco, 2008: 46" authorityName=": Lourenco" authorityPageNumber="46" authorityYear="2008" box="[112,553,1868,1892]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="berberensis">
<emphasis id="3CECEADEDC4140076861F8B1FEB15704" box="[112,318,1868,1892]" italics="true" pageId="28" pageNumber="27">Buthus berberensis</emphasis>
:
<bibRefCitation id="6A094B3DDC414007695EF8B0FDA65704" author="LOURENCO, W. R. " box="[335,553,1868,1892]" firstAuthor="Lourenco" page="46" pageId="28" pageNumber="27" pagination="45 - 52" refId="ref10253" refString="LOURENCO, W. R. 2008. About the presence of the genus Buthus Leach, 1815 in the Arabian Peninsula and description of a new species (Scorpiones, Buthidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 15 (179): 45 - 52." type="journal article" year="2008">Lourenço, 2008: 46</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="8F3910DDDC4140076A27F8B1FF3457E4" author="SOUSA P., M. A. ARNEDO &amp; D. J. HARRIS." page="38" pageId="28" pageNumber="27" year="2017">
<bibRefCitation id="6A094B3DDC4140076A27F8B1FF3457E4" author="SOUSA P., M. A. ARNEDO &amp; D. J. HARRIS" etAl="et al." firstAuthor="Sousa" page="38" pageId="28" pageNumber="27" pagination="15 - 84" refId="ref10552" refString="SOUSA P., M. A. ARNEDO &amp; D. J. HARRIS. 2017. Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae). ZooKeys, 686: 15 - 84." type="journal article" year="2017">Sousa et al., 2017: 38</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="46826547DC4140076861F84DFC5956BF" pageId="28" pageNumber="27" type="materials_examined">
<paragraph id="0E2736CCDC4140076861F84DFF475787" blockId="28.[112,770,1968,2023]" pageId="28" pageNumber="27">
<typeStatus id="D123886EDC4140076861F84DFF2957A7" box="[112,166,1968,1991]" pageId="28" pageNumber="27">TYPE</typeStatus>
LOCALITY AND
<typeStatus id="D123886EDC4140076972F84DFE1A57A7" box="[355,405,1968,1991]" pageId="28" pageNumber="27">TYPE</typeStatus>
REPOSITORY. Somaliland, Zeyla;
<collectionCode id="6889AE09DC4140076861F82DFF4C5787" box="[112,195,2000,2023]" pageId="28" pageNumber="27">BMNH</collectionCode>
.
</paragraph>
<paragraph id="0E2736CCDC4140076B3EF9B9FC5956BF" blockId="28.[815,1473,1604,1759]" pageId="28" pageNumber="27">
MATERIAL EXAMINED.
<materialsCitation id="BEF03C91DC4140076C29F9B9FA4E56FC" ID-GBIF-Occurrence="3754823343" collectingDate="2017-09-11" collectingDateMax="2017-09-12" collectingDateMin="2017-09-11" collectionCode="FKCP" collectorName="F. KovařÍk" elevation="245" latitude="10.600278" longLatPrecision="21" longitude="43.43528" pageId="28" pageNumber="27" specimenCount="29" specimenCount-female="7" specimenCount-male="22">
<emphasis id="3CECEADEDC4140076C29F9B9FB36563C" bold="true" box="[1080,1209,1604,1628]" pageId="28" pageNumber="27">
<collectorName id="A36D531ADC4140076C29F9B9FB36563C" box="[1080,1209,1604,1628]" pageId="28" pageNumber="27">Somaliland</collectorName>
</emphasis>
,
<collectorName id="A36D531ADC4140076CC7F9B9FAA6563C" box="[1238,1321,1604,1628]" pageId="28" pageNumber="27">Gerissa</collectorName>
,
<geoCoordinate id="6BAC500BDC4140076D52F9B9FA4E563C" box="[1347,1473,1604,1628]" degrees="10" direction="north" minutes="36" orientation="latitude" pageId="28" pageNumber="27" precision="15" seconds="01" value="10.600278">10°36'01&quot;N</geoCoordinate>
<geoCoordinate id="6BAC500BDC4140076B3EF999FC23561C" box="[815,940,1636,1660]" degrees="43" direction="east" minutes="26" orientation="longitude" pageId="28" pageNumber="27" precision="15" seconds="07" value="43.43528">43°26'07&quot;E</geoCoordinate>
,
<elevation id="85B5D1FFDC4140076BA8F999FBC8561C" box="[953,1095,1636,1660]" metricMagnitude="2" metricUnit="m" metricValue="2.45" pageId="28" pageNumber="27" unit="m" value="245.0">
<quantity id="C9609B29DC4140076BA8F999FC71561C" box="[953,1022,1636,1660]" metricMagnitude="2" metricUnit="m" metricValue="2.45" pageId="28" pageNumber="27" unit="m" value="245.0">245 m</quantity>
a. s. l.
</elevation>
(
<collectorName id="A36D531ADC4140076C46F999FB6E561C" box="[1111,1249,1636,1660]" pageId="28" pageNumber="27">Locality No.</collectorName>
<emphasis id="3CECEADEDC4140076CF8F999FAA9561C" bold="true" box="[1257,1318,1636,1660]" pageId="28" pageNumber="27">19SH</emphasis>
= 17ST),
<date id="7A26100CDC4140076D89F999FC2456FC" pageId="28" pageNumber="27" value="2017-09-11" valueMax="2017-09-12" valueMin="2017-09-11">
<collectingDate id="6A62E9E4DC4140076D89F999FC2456FC" pageId="28" pageNumber="27" value="2017-09-11" valueMax="2017-09-12" valueMin="2017-09-11">11.- 12.IX.2017</collectingDate>
</date>
,
<specimenCount id="189EFD45DC4140076BA9F979FC6556FC" box="[952,1002,1668,1692]" pageId="28" pageNumber="27" type="male">22♂</specimenCount>
<specimenCount id="189EFD45DC4140076BF8F979FB8A56FC" box="[1001,1029,1668,1692]" pageId="28" pageNumber="27" type="female">7♀</specimenCount>
21juvs. (
<collectorName id="A36D531ADC4140076C76F979FB1556FC" box="[1127,1178,1668,1692]" pageId="28" pageNumber="27">Nos.</collectorName>
1294, 1295, 1297),
<collectionCode id="6889AE09DC4140076D6BF979FA3256FC" box="[1402,1469,1668,1692]" pageId="28" pageNumber="27">FKCP</collectionCode>
,
</materialsCitation>
<materialsCitation id="BEF03C91DC4140076B3EF959FC5D56BF" ID-GBIF-Occurrence="3754823325" collectingDate="2017-09-11" collectingDateMax="2017-09-12" collectingDateMin="2017-09-11" collectionCode="RTOC" collectorName="F. KovařÍk" elevation="245" latitude="10.600278" longLatPrecision="21" longitude="43.43528" pageId="28" pageNumber="27" specimenCount="29" specimenCount-female="7" specimenCount-male="22">
<specimenCount id="189EFD45DC4140076B3EF959FCDA56DC" box="[815,853,1700,1724]" pageId="28" pageNumber="27" type="male">1♂</specimenCount>
<specimenCount id="189EFD45DC4140076B44F959FCFC56DC" box="[853,883,1700,1724]" pageId="28" pageNumber="27" type="female">1♀</specimenCount>
,
<collectionCode id="6889AE09DC4140076B90F959FC4656DC" box="[897,969,1700,1724]" pageId="28" pageNumber="27">RTOC</collectionCode>
,
<date id="7A26100CDC4140076BC7F959FBC156DC" box="[982,1102,1700,1724]" pageId="28" pageNumber="27" value="2019-07-03">
<collectingDate id="6A62E9E4DC4140076BC7F959FBC156DC" box="[982,1102,1700,1724]" pageId="28" pageNumber="27" value="2019-07-03">3.VII.2019</collectingDate>
</date>
,
<specimenCount id="189EFD45DC4140076C4BF959FBF456DC" box="[1114,1147,1700,1724]" pageId="28" pageNumber="27" type="male">8♂</specimenCount>
<specimenCount id="189EFD45DC4140076C6AF959FB1956DC" box="[1147,1174,1700,1724]" pageId="28" pageNumber="27" type="female">3♀</specimenCount>
12juvs. (No. 1686
<emphasis id="3CECEADEDC4140076D73F959FAE456DC" bold="true" box="[1378,1387,1700,1724]" pageId="28" pageNumber="27">)</emphasis>
,
<collectionCode id="6889AE09DC4140076D6BF959FA3256DC" box="[1402,1469,1700,1724]" pageId="28" pageNumber="27">FKCP</collectionCode>
, leg.
<collectorName id="A36D531ADC4140076B4CF938FC5D56BF" box="[861,978,1733,1759]" pageId="28" pageNumber="27">F. KovařÍk</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46826547DC4140076B3EF8FCFA2A5799" pageId="28" pageNumber="27" type="diagnosis">
<paragraph id="0E2736CCDC4140076B3EF8FCFA2A5799" blockId="28.[815,1474,1793,2041]" pageId="28" pageNumber="27">
DIAGNOSIS. Total length
<quantity id="C9609B29DC4140076C2AF8FCFB3C5778" box="[1083,1203,1793,1817]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.4" metricValueMin="3.7" pageId="28" pageNumber="27" unit="mm" value="45.5" valueMax="54.0" valueMin="37.0">37–54 mm</quantity>
. Base color yellow dark pattern reduced; telson brown; chelicerae pale yellow without reticulation. Pedipalp movable fingers bear 10–11 rows of granules, with 12 outer and inner denticles and fixed finger with 10–11 outer and inner denticles. Chela of pedipalp narrower in male, its length to width ratio 3.4–3.6 in females and 4.1–4.3 in males. Telson bulbous, with aculeus shorter than vesicle. Pectinal teeth number
<quantity id="C9609B29DC4140076C09F81CFBFB5799" box="[1048,1140,2017,2041]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" metricValueMax="7.366" metricValueMin="6.35" pageId="28" pageNumber="27" unit="in" value="27.0" valueMax="29.0" valueMin="25.0">25–29 in</quantity>
females and
<quantity id="C9609B29DC4140076CEFF81CFAD55799" box="[1278,1370,2017,2041]" metricMagnitude="-1" metricUnit="m" metricValue="8.001" metricValueMax="8.89" metricValueMin="7.112" pageId="28" pageNumber="27" unit="in" value="31.5" valueMax="35.0" valueMin="28.0">28–35 in</quantity>
males.
</paragraph>
</subSubSection>
<subSubSection id="46826547DC7D403B6860FDE7FE9C55F2" pageId="32" pageNumber="31" type="discussion">
<paragraph id="0E2736CCDC7D403B6860FDE7FEA253F2" blockId="32.[113,771,538,914]" pageId="32" pageNumber="31">
COMMENTS.
<bibRefCitation id="6A094B3DDC7D403B68E7FDE6FD975252" author="LEVY, G. &amp; P. AMITAI" bookContentInfo="132 pp." box="[246,536,538,563]" journalOrPublisher="Jerusalem: The Israel Academy of Sciences and Humanities" pageId="32" pageNumber="31" refId="ref10220" refString="LEVY, G. &amp; P. AMITAI. 1980. Fauna Palaestina, Arachnida I. - Scorpiones. Jerusalem: The Israel Academy of Sciences and Humanities, 132 pp." title="Fauna Palaestina, Arachnida I. - Scorpiones." type="book" year="1980">Levy &amp; Amitai (1980: 21)</bibRefCitation>
synonymized
<taxonomicName id="C9984D4FDC7D403B6AA8FDE7FE7D5232" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="zeylensis">
<emphasis id="3CECEADEDC7D403B6AA8FDE7FEC85232" italics="true" pageId="32" pageNumber="31">Buthus occitanus zeylensis</emphasis>
<bibRefCitation id="6A094B3DDC7D403B6946FDC7FE7D5232" author="Pocock" box="[343,498,570,594]" firstAuthor="Pocock" pageId="32" pageNumber="31" pagination="55 - 63" refId="ref10359" refString="POCOCK, R. I. 1900. On a collection of insects and arachnids made in 1895 and 1897 by Mr. C. A. V. Peel, F. Z. S. in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somaliland and the Boran Country. Proceedings of the Zoological Society of London, 1900: 55 - 63." type="book chapter" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="C9984D4FDC7D403B6A53FDC7FE1A5212" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="berberensis">
<emphasis id="3CECEADEDC7D403B6A53FDC7FF635212" italics="true" pageId="32" pageNumber="31">Buthus occitanus berberensis</emphasis>
<bibRefCitation id="6A094B3DDC7D403B68EBFDA7FE1A5212" author="Pocock" box="[250,405,602,626]" firstAuthor="Pocock" pageId="32" pageNumber="31" pagination="55 - 63" refId="ref10359" refString="POCOCK, R. I. 1900. On a collection of insects and arachnids made in 1895 and 1897 by Mr. C. A. V. Peel, F. Z. S. in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somaliland and the Boran Country. Proceedings of the Zoological Society of London, 1900: 55 - 63." type="book chapter" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
. They argued that “examination of Pocock´s
<typeStatus id="D123886EDC7D403B68EEFD86FEB852F3" box="[255,311,635,659]" pageId="32" pageNumber="31">types</typeStatus>
and of additional material from
<collectingCountry id="768F765CDC7D403B6ABDFD87FC8C52F2" box="[684,771,634,658]" name="Somalia" pageId="32" pageNumber="31">Somalia</collectingCountry>
showed that these two forms are in fact the two sexes of the same subspecies, the first the male, the second the female.” We studied in detail a large collection of Somaliland specimens collected during expeditions of 2011–2019; this study, including cytogenetic and DNA analysis, confirmed that these two taxa are valid species. Here, we restore
<taxonomicName id="C9984D4FDC7D403B6AA8FCC7FE5B5312" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="occitanus" subSpecies="zeylensis">
<emphasis id="3CECEADEDC7D403B6AA8FCC7FEB25312" italics="true" pageId="32" pageNumber="31">Buthus occitanus zeylensis</emphasis>
<bibRefCitation id="6A094B3DDC7D403B6952FCA7FE5B5312" author="Pocock" box="[323,468,858,882]" firstAuthor="Pocock" pageId="32" pageNumber="31" pagination="55 - 63" refId="ref10359" refString="POCOCK, R. I. 1900. On a collection of insects and arachnids made in 1895 and 1897 by Mr. C. A. V. Peel, F. Z. S. in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somaliland and the Boran Country. Proceedings of the Zoological Society of London, 1900: 55 - 63." type="book chapter" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
from synonymy and elevate it to species rank.
</paragraph>
<paragraph id="0E2736CCDC7D403B6860FC47FE9C55F2" blockId="32.[113,771,954,1426]" pageId="32" pageNumber="31">
COMMENTS ON LOCALITIES AND LIFE STRATEGY. In 2019, two of us (FK, HE) visited the city of Zeyla, the
<typeStatus id="D123886EDC7D403B6A4DFC26FD045393" box="[604,651,987,1011]" pageId="32" pageNumber="31">type</typeStatus>
locality of
<taxonomicName id="C9984D4FDC7D403B6860FC07FE495472" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[113,454,1018,1042]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="zeylensis">
<emphasis id="3CECEADEDC7D403B6860FC07FEA95472" box="[113,294,1018,1042]" italics="true" pageId="32" pageNumber="31">Buthus zeylensis</emphasis>
<bibRefCitation id="6A094B3DDC7D403B693CFC07FE495472" author="Pocock" box="[301,454,1018,1042]" firstAuthor="Pocock" pageId="32" pageNumber="31" pagination="55 - 63" refId="ref10359" refString="POCOCK, R. I. 1900. On a collection of insects and arachnids made in 1895 and 1897 by Mr. C. A. V. Peel, F. Z. S. in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somaliland and the Boran Country. Proceedings of the Zoological Society of London, 1900: 55 - 63." type="book chapter" year="1900">Pocock, 1900</bibRefCitation>
</taxonomicName>
. It is an extremely hot place, with heavily salted soils. We collected scorpions several km from the city (
<geoCoordinate id="6BAC500BDC7D403B6902FBC7FE285432" box="[275,423,1082,1106]" degrees="11" direction="north" minutes="19" orientation="latitude" pageId="32" pageNumber="31" precision="1" seconds="31.2" value="11.325334">11°19'31.2&quot;N</geoCoordinate>
<geoCoordinate id="6BAC500BDC7D403B69A0FBC7FDCC5432" box="[433,579,1082,1106]" degrees="43" direction="east" minutes="22" orientation="longitude" pageId="32" pageNumber="31" precision="1" seconds="16.9" value="43.37136">43°22'16.9&quot;E</geoCoordinate>
) and found there
<taxonomicName id="C9984D4FDC7D403B6860FBA6FD9B5412" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[113,532,1114,1138]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="polystictus">
<emphasis id="3CECEADEDC7D403B6860FBA6FEE75412" box="[113,360,1114,1138]" italics="true" pageId="32" pageNumber="31">Hottentotta polystictus</emphasis>
(Pocock, 1896)
</taxonomicName>
,
<taxonomicName id="C9984D4FDC7D403B6A0CFBA7FE8254F2" authority="(Pavesi, 1885)" baseAuthorityName="Pavesi" baseAuthorityYear="1885" class="Arachnida" family="Buthidae" genus="Microbuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="3CECEADEDC7D403B6A0CFBA7FC8C5412" box="[541,771,1114,1138]" italics="true" pageId="32" pageNumber="31">Microbuthus litoralis</emphasis>
(Pavesi, 1885)
</taxonomicName>
(first record for Somaliland),
<taxonomicName id="C9984D4FDC7D403B6A5AFB87FEE454D2" authority="KovarIk et Lowe, 2016" authorityName="KovarIk et Lowe" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Orthochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="afar">
<emphasis id="3CECEADEDC7D403B6A5AFB87FC8C54F2" box="[587,771,1146,1170]" italics="true" pageId="32" pageNumber="31">Orthochirus afar</emphasis>
KovařÍk et Lowe, 2016
</taxonomicName>
and
<taxonomicName id="C9984D4FDC7D403B698EFB67FF2854B2" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" class="Arachnida" family="Buthidae" genus="Parabuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="granimanus">
<emphasis id="3CECEADEDC7D403B698EFB67FD2954D2" box="[415,678,1178,1202]" italics="true" pageId="32" pageNumber="31">Parabuthus granimanus</emphasis>
Pocock, 1895
</taxonomicName>
(topotypes) but no
<taxonomicName id="C9984D4FDC7D403B696FFB47FE4654B2" authorityName="Leach" authorityYear="1815" box="[382,457,1210,1234]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="3CECEADEDC7D403B696FFB47FE4654B2" box="[382,457,1210,1234]" italics="true" pageId="32" pageNumber="31">Buthus</emphasis>
</taxonomicName>
specimens.
<taxonomicName id="C9984D4FDC7D403B6A41FB47FC8C54B2" authorityName="Pocock" authorityYear="1900" box="[592,771,1210,1234]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="zeylensis">
<emphasis id="3CECEADEDC7D403B6A41FB47FC8C54B2" box="[592,771,1210,1234]" italics="true" pageId="32" pageNumber="31">Buthus zeylensis</emphasis>
</taxonomicName>
is common in Gerissa, a very hot, sandy semidesert area (
<figureCitation id="96A32A49DC7D403B6868FB07FF565572" box="[121,217,1274,1298]" captionStart="Figures 100–101" captionStartId="17.[113,190,2013,2035]" captionTargetBox="[113,1474,234,1995]" captionTargetId="figure-35@17.[113,1474,234,1998]" captionTargetPageId="17" captionText="Figures 100–101: Localities. Figure 100. Buthus pococki sp. n., type locality (19SC). Figure 101. B. zeylensis, locality 19SH." figureDoi="http://doi.org/10.5281/zenodo.4648410" httpUri="https://zenodo.org/record/4648410/files/figure.png" pageId="32" pageNumber="31">Fig. 101</figureCitation>
) where we recorded also
<taxonomicName id="C9984D4FDC7D403B6A11FB06FD155572" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[512,666,1274,1298]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="polystictus">
<emphasis id="3CECEADEDC7D403B6A11FB06FD155572" box="[512,666,1274,1298]" italics="true" pageId="32" pageNumber="31">H. polystictus</emphasis>
</taxonomicName>
,
<taxonomicName id="C9984D4FDC7D403B6ABAFB06FD735572" authorityName="KovarIk et Lowe" authorityYear="2016" box="[683,764,1274,1298]" class="Arachnida" family="Buthidae" genus="Orthochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="afar">
<emphasis id="3CECEADEDC7D403B6ABAFB06FD735572" box="[683,764,1274,1298]" italics="true" pageId="32" pageNumber="31">O. afar</emphasis>
</taxonomicName>
, and
<taxonomicName id="C9984D4FDC7D403B68B4FAE6FEC75552" authorityName="Pocock" authorityYear="1895" box="[165,328,1307,1330]" class="Arachnida" family="Buthidae" genus="Parabuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="granimanus">
<emphasis id="3CECEADEDC7D403B68B4FAE6FEC75552" box="[165,328,1307,1330]" italics="true" pageId="32" pageNumber="31">P. granimanus</emphasis>
</taxonomicName>
, but also
<taxonomicName id="C9984D4FDC7D403B69D5FAE7FE9B5532" authority="KovarIk, 2012" authorityName="KovarIk" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="3CECEADEDC7D403B69D5FAE7FC8C5552" box="[452,771,1306,1330]" italics="true" pageId="32" pageNumber="31">Compsobuthus somalilandus</emphasis>
KovařÍk, 2012
</taxonomicName>
,
<taxonomicName id="C9984D4FDC7D403B6933FAC6FD4E5532" authority="KovarIk et al., 2018" authorityName="KovarIk" authorityYear="2018" box="[290,705,1338,1365]" class="Arachnida" family="Buthidae" genus="Gint" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="gubanensis">
<emphasis id="3CECEADEDC7D403B6933FAC6FE595532" box="[290,470,1338,1362]" italics="true" pageId="32" pageNumber="31">Gint gubanensis</emphasis>
KovařÍk et al., 2018
</taxonomicName>
(
<typeStatus id="D123886EDC7D403B6AC2FAC7FC8C5532" box="[723,771,1338,1362]" pageId="32" pageNumber="31">type</typeStatus>
locality), and
<taxonomicName id="C9984D4FDC7D403B6902FAA7FC8C5512" authority="KovarIk et al., 2018" authorityName="KovarIk" authorityYear="2018" box="[275,771,1370,1397]" class="Arachnida" family="Buthidae" genus="Neobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="gubanensis">
<emphasis id="3CECEADEDC7D403B6902FAA7FD825512" box="[275,525,1370,1394]" italics="true" pageId="32" pageNumber="31">Neobuthus gubanensis</emphasis>
KovařÍk et al., 2018
</taxonomicName>
(
<typeStatus id="D123886EDC7D403B6868FA87FF2655F2" box="[121,169,1402,1426]" pageId="32" pageNumber="31">type</typeStatus>
locality).
</paragraph>
</subSubSection>
</treatment>
</document>
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