781 lines
101 KiB
XML
781 lines
101 KiB
XML
<document id="2E6311EE54D3E88BA1501F6F7CA34B19" ID-CLB-Dataset="3486" ID-DOI="10.11646/zootaxa.5068.1.3" ID-GBIF-Dataset="87a6a8b8-2167-4ff1-89c5-d3a94e8ba51b" ID-ISSN="1175-5326" ID-Zenodo-Dep="5702104" ID-ZooBank="D6B213E5-8A79-4438-8481-01C074EB74CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636961428043" checkinUser="plazi" docAuthor="Maeda, Ken, Shinzato, Chuya, Koyanagi, Ryo, Kunishima, Taiga, Kobayashi, Hirozumi, Satoh, Noriyuki & Palla, Herminie P." docDate="2021" docId="635C87825F6CFFC251AD961CB2FBFBBB" docLanguage="en" docName="zootaxa.5068.1.3.pdf" docOrigin="Zootaxa 5068 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinogobius tandikan Maeda, Kobayashi & Palla 2021, new species" docType="treatment" docUuid="47FBF5CB-033A-4857-AFE6-91E5706ACFEF" docUuidSource="ZooBank" docVersion="11" lastPageNumber="94" masterDocId="9F65FFFA5F64FFCF513A9137B033FFE2" masterDocTitle="Two new species of Rhinogobius (Gobiiformes: Oxudercidae) from Palawan, Philippines, with their phylogenetic placement" masterLastPageNumber="98" masterPageNumber="81" pageNumber="89" updateTime="1702002386712" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6A6660D0F9B1630D205BA60827C3E520">Two new species of Rhinogobius (Gobiiformes: Oxudercidae) from Palawan, Philippines, with their phylogenetic placement</mods:title>
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<mods:namePart id="E81925362ED49B4DFD843856172C189F">Maeda, Ken</mods:namePart>
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<mods:nameIdentifier id="771762404944C86BB1FED4526987570A" type="ORCID">0000-0003-3631-811X</mods:nameIdentifier>
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<mods:affiliation id="9527260AB5B1BB11CD284731B60D4D79">Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 - 0495, Japan. * Corresponding author. goby @ live. jp; https: // orcid. org / 0000 - 0003 - 3631 - 811 X</mods:affiliation>
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<mods:nameIdentifier id="01637C2E54BC7D96FC4A580E95A75C0C" type="email">goby@live.jp</mods:nameIdentifier>
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<mods:roleTerm id="7F3BA46F2E553414A407107E9F35A8B8">Author</mods:roleTerm>
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<mods:namePart id="F5835D9C73E9253D0C31B2E83DF5D0E0">Shinzato, Chuya</mods:namePart>
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<mods:affiliation id="830F197C7034265CDF1CA551CEF88AFC">Atmosphere and Ocean Research Institute, The University of Tokyo, Chiba 277 - 8564, Japan.</mods:affiliation>
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<mods:namePart id="454F3EBD8B8F4D107B8AB8EA439024AD">Koyanagi, Ryo</mods:namePart>
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<mods:nameIdentifier id="B213012F19E872887BC3429EBE011AFE" type="ORCID">0000-0003-3631-811X</mods:nameIdentifier>
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<mods:affiliation id="AA5C7067897FFA162335764D7ECDA667">Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 - 0495, Japan. * Corresponding author. goby @ live. jp; https: // orcid. org / 0000 - 0003 - 3631 - 811 X & https: // orcid. org / 0000 - 0002 - 9694 - 0288</mods:affiliation>
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<mods:nameIdentifier id="F8A0E2A816624C7F83D196299EE69C9A" type="email">goby@live.jp</mods:nameIdentifier>
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<mods:namePart id="8091533F8C5C5022C0FF24241BE5164E">Kunishima, Taiga</mods:namePart>
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<mods:affiliation id="CD995BC42DA524175A77E0B61B893ED0">Wakayama Prefectural Museum of Natural History, 370 - 1 Funoo, Kainan, Wakayama 642 - 0001, Japan.</mods:affiliation>
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<mods:namePart id="FF6901B8300686FBFB653DF4FE5AF830">Kobayashi, Hirozumi</mods:namePart>
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<mods:affiliation id="394EA353CBEB1CCB43A13D05A32E1B91">University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903 - 0213, Japan.</mods:affiliation>
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<mods:namePart id="A199A9E70B2C7469E7DFA69B1AF700E3">Satoh, Noriyuki</mods:namePart>
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<mods:nameIdentifier id="D38DE0567E181BE665CBB6B0D9FD8919" type="ORCID">0000-0003-3631-811X</mods:nameIdentifier>
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<mods:affiliation id="6FC217758B50388780CA7EF153F57A3D">Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 - 0495, Japan. * Corresponding author. goby @ live. jp; https: // orcid. org / 0000 - 0003 - 3631 - 811 X & https: // orcid. org / 0000 - 0002 - 4480 - 3572</mods:affiliation>
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<mods:nameIdentifier id="83B1F974EB11672701C611CA506A7A05" type="email">goby@live.jp</mods:nameIdentifier>
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<mods:namePart id="6FCE8DBD9B1AA6CFD06C715260C8F8E5">Palla, Herminie P.</mods:namePart>
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<mods:nameIdentifier id="AE1C0D4841F9177FD9295D28FB69CC1F" type="ORCID">0000-0003-2080-6602</mods:nameIdentifier>
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<mods:affiliation id="67E664DB0ED7CF6FAEA5003EF3557AAE">College of Fisheries and Aquatic Sciences, Western Philippines University - Puerto Princesa Campus, Puerto Princesa City, 5300 Palawan, Philippines. https: // orcid. org / 0000 - 0003 - 2080 - 6602</mods:affiliation>
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<mods:title id="1533C4CA169DDF212B72FE59FCDD79CB">Zootaxa</mods:title>
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<mods:part id="64C3521495B86062DE52230280B308C3">
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<mods:date id="CF8E6D0BD4E79FA0FD50E57349FA0FC3">2021</mods:date>
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<mods:number id="E39A3921C824B6D817F3F4E9A46269AC">2021-11-15</mods:number>
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<mods:number id="BFC8A068DEF9821E41DBA8A3120F010D">5068</mods:number>
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<mods:classification id="81AC99572A9A21D860FEEBDEE9F9420D">journal article</mods:classification>
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<mods:identifier id="08FC6CB221BC05AB6E38083602A0983A" type="DOI">10.11646/zootaxa.5068.1.3</mods:identifier>
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<treatment id="635C87825F6CFFC251AD961CB2FBFBBB" ID-DOI="http://doi.org/10.5281/zenodo.5706125" ID-GBIF-Taxon="190303668" ID-Zenodo-Dep="5706125" LSID="urn:lsid:zoobank.org:act:47FBF5CB-033A-4857-AFE6-91E5706ACFEF" httpUri="http://treatment.plazi.org/id/635C87825F6CFFC251AD961CB2FBFBBB" lastPageId="13" lastPageNumber="94" pageId="8" pageNumber="89">
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<subSubSection id="A3EF651F5F6CFFC751AD961CB200F88B" pageId="8" pageNumber="89" type="nomenclature">
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<paragraph id="EB4A36945F6CFFC751AD961CB3AFF8A7" blockId="8.[151,924,1835,1934]" box="[151,924,1835,1862]" pageId="8" pageNumber="89">
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<heading id="B00281F85F6CFFC751AD961CB3AFF8A7" bold="true" box="[151,924,1835,1862]" fontSize="11" level="1" pageId="8" pageNumber="89" reason="1">
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<emphasis id="D981EA865F6CFFC751AD961CB3AFF8A7" bold="true" box="[151,924,1835,1862]" pageId="8" pageNumber="89">
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<taxonomicName id="2CF54D175F6CFFC751AD961CB2CCF8A7" authority="Maeda, Kobayashi & Palla" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[151,767,1835,1862]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="89" phylum="Chordata" rank="species" species="tandikan" status="sp. nov.">
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<emphasis id="D981EA865F6CFFC751AD961CB196F8A4" bold="true" box="[151,421,1835,1862]" italics="true" pageId="8" pageNumber="89">Rhinogobius tandikan</emphasis>
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Maeda, Kobayashi & Palla
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</taxonomicName>
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,
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<taxonomicNameLabel id="C2B257FD5F6CFFC75236961CB3AFF8A7" box="[780,924,1835,1861]" pageId="8" pageNumber="89" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EB4A36945F6CFFC751AD9667B200F88B" blockId="8.[151,924,1835,1934]" box="[151,563,1870,1898]" pageId="8" pageNumber="89">[New English name: Tandikan goby]</paragraph>
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||
</subSubSection>
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<subSubSection id="A3EF651F5F6CFFC751AD9644B1E1F86C" box="[151,466,1907,1934]" pageId="8" pageNumber="89" type="description">
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<paragraph id="EB4A36945F6CFFC751AD9644B1E1F86C" blockId="8.[151,924,1835,1934]" box="[151,466,1907,1934]" pageId="8" pageNumber="89">
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(
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<figureCitation id="73CE2A115F6CFFC751A59644B0DEF86F" box="[159,237,1907,1933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1837,1861]" captionTargetBox="[157,1419,1035,1802]" captionTargetId="figure-341@3.[151,1436,1027,1812]" captionTargetPageId="3" captionText="FIGURE 1. Preserved specimens of Rhinogobius estrellae (a–d, paratypes) and Rhinogobius tandikan (e–h, paratypes). Specimens in the left side are males, the right side are females. a, URM-P 49305, 39.6 mm SL; b, WPU-PPC-P 55, 38.4 mm SL; c, URM-P 49296, 36.1 mm SL; d, URM-P 49306, 36.4 mm SL; e, URM-P 49314, 46.9 mm SL; f, URM-P 49315, 46.5 mm SL; g, URM-P 49318, 45.2 mm SL; h, URM-P 49320, 37.9 mm SL (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.5702106" httpUri="https://zenodo.org/record/5702106/files/figure.png" pageId="8" pageNumber="89">Figs. 1</figureCitation>
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,
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<figureCitation id="73CE2A115F6CFFC751C19644B139F86F" box="[251,266,1907,1933]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[217,1372,191,1911]" captionTargetId="figure-21@6.[200,1387,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Head length, upper-jaw length, second dorsal-fin length, anal-fin length, and caudal-fin length of Rhinogobius estrellae (red circles) and Rhinogobius tandikan (blue triangles). Solid and open symbols represent males and females, respectively." figureDoi="http://doi.org/10.5281/zenodo.5702110" httpUri="https://zenodo.org/record/5702110/files/figure.png" pageId="8" pageNumber="89">3</figureCitation>
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,
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<figureCitation id="73CE2A115F6CFFC7502D9644B177F86F" box="[279,324,1907,1933]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="10.[151,250,1153,1177]" captionStartId-1="11.[151,250,1943,1967]" captionStartId-2="12.[151,250,687,711]" captionTargetBox-0="[151,1436,345,1130]" captionTargetBox-1="[328,1254,188,1905]" captionTargetBox-2="[151,1437,181,663]" captionTargetId-0="figure-99@10.[151,1436,345,1130]" captionTargetId-1="figure-20@11.[306,1282,181,1919]" captionTargetId-2="figure-21@12.[151,1437,181,663]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 6. Rhinogobius tandikan immediately after fixation. a, NSMT-P 140093, holotype, male, 43.6 mm SL; b, URM-P 49316, paratype, female, 44.8 mm SL (photo by K. Maeda)." captionText-1="FIGURE 7. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius tandikan (URM-P 49307, paratype). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." captionText-2="FIGURE 8. In situ underwater photographs of Rhinogobius tandikan in Cayulo River, Palawan. a, male, 31 May 2018; b, female, 1 June 2018 (photo by K. Maeda)." figureDoi-0="http://doi.org/10.5281/zenodo.5702116" figureDoi-1="http://doi.org/10.5281/zenodo.5702118" figureDoi-2="http://doi.org/10.5281/zenodo.5702120" httpUri-0="https://zenodo.org/record/5702116/files/figure.png" httpUri-1="https://zenodo.org/record/5702118/files/figure.png" httpUri-2="https://zenodo.org/record/5702120/files/figure.png" pageId="8" pageNumber="89">6–8</figureCitation>
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;
|
||
<tableCitation id="A677032F5F6CFFC750759644B1F9F86C" box="[335,458,1907,1934]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStart-3="TABLE 5" captionStart-4="TABLE 6" captionStartId-0="5.[151,239,151,175]" captionStartId-1="5.[151,239,675,699]" captionStartId-2="5.[151,239,869,893]" captionStartId-3="5.[151,239,1063,1087]" captionStartId-4="12.[151,239,795,819]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="12" captionText-0="TABLE 2. Pectoral-fin ray counts of Rhinogobius estrellae and Rhinogobius tandikan." captionText-1="TABLE 3. Longitudinal scale counts of Rhinogobius estrellae and Rhinogobius tandikan." captionText-2="TABLE 4. Transverse scale counts of Rhinogobius estrellae and Rhinogobius tandikan." captionText-3="TABLE 5. Predorsal scale counts of Rhinogobius estrellae and Rhinogobius tandikan." captionText-4="TABLE 6. Morphometrics of Rhinogobius estrellae and Rhinogobius tandikan expressed as a percentage of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 1, pectoral fin; P 2, pelvic fin." httpUri-0="http://table.plazi.org/id/BF8A661C5F61FFCA51AD91A0B406FF4D" httpUri-4="http://table.plazi.org/id/BF8A661C5F68FFC351AD922CB4C0FCB5" pageId="8" pageNumber="89" tableUuid-0="BF8A661C5F61FFCA51AD91A0B406FF4D" tableUuid-4="BF8A661C5F68FFC351AD922CB4C0FCB5">Tables 2–6</tableCitation>
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||
)
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="A3EF651F5F6CFFC651AD968CB1A6FE6A" lastPageId="9" lastPageNumber="90" pageId="8" pageNumber="89" type="materials_examined">
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<paragraph id="EB4A36945F6CFFC751AD968CB19DF81B" blockId="8.[151,1436,1979,2041]" pageId="8" pageNumber="89">
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<emphasis id="D981EA865F6CFFC751AD968CB1B7F837" bold="true" box="[151,388,1979,2005]" pageId="8" pageNumber="89">Material examined.</emphasis>
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<materialsCitation id="5B9D3CC95F6CFFC750B4968CB198F81B" ID-GBIF-Occurrence="3407138309" country="Philippines" county="Island" location="Cayulo River" municipality="Bahile" pageId="8" pageNumber="89" specimenCount="16" specimenCount-female="16" stateProvince="Puerto Princesa">
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Thirteen males and
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<specimenCount id="FDF3FD1D5F6CFFC7534F968CB2C1F837" box="[629,754,1979,2005]" pageId="8" pageNumber="89" type="female">16 females</specimenCount>
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from
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,
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<collectingMunicipality id="0B2EACEE5F6CFFC752DD968CB400F837" box="[999,1075,1979,2005]" pageId="8" pageNumber="89">Bahile</collectingMunicipality>
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,
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<collectingRegion id="2931F8765F6CFFC7557B968BB51CF837" box="[1089,1327,1979,2006]" country="Philippines" name="Puerto Princesa" pageId="8" pageNumber="89">Puerto Princesa City</collectingRegion>
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,
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<collectingRegion id="2931F8765F6CFFC75407968CB5AFF837" box="[1341,1436,1979,2005]" country="Philippines" name="Palawan" pageId="8" pageNumber="89">Palawan</collectingRegion>
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<collectingCounty id="022B4E185F6CFFC751AD96E8B0E8F81B" box="[151,219,2015,2041]" pageId="8" pageNumber="89">Island</collectingCounty>
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in the
|
||
<collectingCountry id="93E276045F6CFFC7501396E8B198F81B" box="[297,427,2015,2041]" name="Philippines" pageId="8" pageNumber="89">Philippines</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F6DFFC651FD91A1B5A6FF52" blockId="9.[151,1437,150,2038]" box="[199,1429,150,177]" pageId="9" pageNumber="90">
|
||
<materialsCitation id="5B9D3CC95F6DFFC651FD91A1B5A6FF52" ID-GBIF-Occurrence="3407138304" box="[199,1429,150,177]" collectingDate="2018-06-01" collectionCode="NSMT-P" collectorName="K. Maeda & Maeda, H & H. P. Palla." pageId="9" pageNumber="90" specimenCode="NSMT-P 140093" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="D981EA865F6DFFC651FD91A1B108FF52" bold="true" box="[199,315,150,176]" pageId="9" pageNumber="90">
|
||
<typeStatus id="344E88365F6DFFC651FD91A1B105FF52" box="[199,310,150,176]" pageId="9" pageNumber="90" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="BB539EEF5F6DFFC6507891A1B230FF52" box="[322,515,150,176]" collectionCode="NSMT-P" pageId="9" pageNumber="90">NSMT-P 140093</specimenCode>
|
||
, male (
|
||
<quantity id="2C0D9B715F6DFFC6536991A1B28AFF53" box="[595,697,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.36" pageId="9" pageNumber="90" unit="mm" value="43.6">43.6 mm</quantity>
|
||
SL),
|
||
<date id="9F4B10545F6DFFC653CD91A1B3B1FF52" box="[759,898,150,177]" pageId="9" pageNumber="90" value="2018-06-01">
|
||
<collectingDate id="8F0FE9BC5F6DFFC653CD91A1B3B1FF52" box="[759,898,150,177]" pageId="9" pageNumber="90" value="2018-06-01">1 June 2018</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="460053425F6DFFC652FF91A0B405FF52" box="[965,1078,150,177]" pageId="9" pageNumber="90">K. Maeda</collectorName>
|
||
, H. Kobayashi, and
|
||
<collectorName id="460053425F6DFFC6542391A0B5A6FF52" box="[1305,1429,150,177]" pageId="9" pageNumber="90">H. P. Palla.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F6DFFC651FD918DB1A6FE6A" blockId="9.[151,1437,150,2038]" pageId="9" pageNumber="90">
|
||
<materialsCitation id="5B9D3CC95F6DFFC651FD918DB1CEFEA2" ID-GBIF-Occurrence="3407138310" collectingDate="2018-05-31" collectionCode="NSMT-P" collectorName="K. Maeda & Maeda, H & H. P. Palla" pageId="9" pageNumber="90" specimenCode="NSMT-P 140094" specimenCount="17" specimenCount-female="8" specimenCount-male="9" typeStatus="paratype">
|
||
<emphasis id="D981EA865F6DFFC651FD918DB175FF36" bold="true" box="[199,326,186,212]" pageId="9" pageNumber="90">
|
||
<typeStatus id="344E88365F6DFFC651FD918DB172FF36" box="[199,321,186,212]" pageId="9" pageNumber="90" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="BB539EEF5F6DFFC65075918DB220FF36" box="[335,531,186,212]" collectionCode="NSMT-P" pageId="9" pageNumber="90">NSMT-P 140094</specimenCode>
|
||
, female (
|
||
<quantity id="2C0D9B715F6DFFC65347918DB2D6FF37" box="[637,741,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.57" pageId="9" pageNumber="90" unit="mm" value="45.7">45.7 mm</quantity>
|
||
SL),
|
||
<date id="9F4B10545F6DFFC65212918DB3F1FF36" box="[808,962,186,213]" pageId="9" pageNumber="90" value="2018-05-31">
|
||
<collectingDate id="8F0FE9BC5F6DFFC65212918DB3F1FF36" box="[808,962,186,213]" pageId="9" pageNumber="90" value="2018-05-31">31 May 2018</collectingDate>
|
||
</date>
|
||
coll.
|
||
<collectorName id="460053425F6DFFC6553F918CB44AFF36" box="[1029,1145,186,213]" pageId="9" pageNumber="90">K. Maeda</collectorName>
|
||
, H. Kobayashi, and
|
||
<collectorName id="460053425F6DFFC6545E918CB0E0FF1A" pageId="9" pageNumber="90">H. P. Palla</collectorName>
|
||
; URM-P 49307–49315,
|
||
<specimenCount id="FDF3FD1D5F6DFFC650D991E8B20AFF1A" box="[483,569,222,249]" pageId="9" pageNumber="90" type="male">7 males</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC6537291E9B2DFFF1B" box="[584,748,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.6899999999999995" metricValueMin="3.61" pageId="9" pageNumber="90" unit="mm" value="41.5" valueMax="46.9" valueMin="36.1">36.1–46.9 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="FDF3FD1D5F6DFFC6526891E9B38DFF1A" box="[850,958,222,248]" pageId="9" pageNumber="90" type="female">2 females</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC652F691E9B443FF1B" box="[972,1136,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.8099999999999996" metricValueMax="3.8899999999999997" metricValueMin="3.7299999999999995" pageId="9" pageNumber="90" unit="mm" value="38.099999999999994" valueMax="38.9" valueMin="37.3">37.3–38.9 mm</quantity>
|
||
SL),
|
||
<date id="9F4B10545F6DFFC6559491E9B570FF1A" box="[1198,1347,222,249]" pageId="9" pageNumber="90" value="2018-05-31">
|
||
<collectingDate id="8F0FE9BC5F6DFFC6559491E9B570FF1A" box="[1198,1347,222,249]" pageId="9" pageNumber="90" value="2018-05-31">31 May 2018</collectingDate>
|
||
</date>
|
||
coll.
|
||
<collectorName id="460053425F6DFFC654BB91E8B0D5FEFE" pageId="9" pageNumber="90">K. Maeda</collectorName>
|
||
, H. Kobayashi, and
|
||
<collectorName id="460053425F6DFFC650FE9034B208FEFE" box="[452,571,258,285]" pageId="9" pageNumber="90">H. P. Palla</collectorName>
|
||
; URM-P 49316–49323,
|
||
<specimenCount id="FDF3FD1D5F6DFFC652729035B3AEFEFE" box="[840,925,258,284]" pageId="9" pageNumber="90" type="male">2 males</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC652919035B47CFEFF" box="[939,1103,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.279999999999999" metricValueMax="4.4799999999999995" metricValueMin="4.08" pageId="9" pageNumber="90" unit="mm" value="42.8" valueMax="44.8" valueMin="40.8">40.8–44.8 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="FDF3FD1D5F6DFFC655889035B52EFEFE" box="[1202,1309,258,284]" pageId="9" pageNumber="90" type="female">6 females</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC654109035B0F0FEA3" metricMagnitude="-2" metricUnit="m" metricValue="4.04" metricValueMax="4.52" metricValueMin="3.56" pageId="9" pageNumber="90" unit="mm" value="40.4" valueMax="45.2" valueMin="35.6">35.6–45.2 mm</quantity>
|
||
SL), same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5B9D3CC95F6DFFC6533D9011B1A1FE6A" ID-GBIF-Occurrence="3407138303" collectingDate="2018-05-31" collectionCode="NSMT-P" collectorName="K. Maeda & Maeda, H & H. P. Palla" pageId="9" pageNumber="90" specimenCount="10" specimenCount-female="7" specimenCount-male="3" typeStatus="paratype">
|
||
WPU-PPC-P 60–64,
|
||
<specimenCount id="FDF3FD1D5F6DFFC653CE9011B353FEA2" box="[756,864,294,321]" pageId="9" pageNumber="90" type="female">5 females</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC652549011B420FEA3" box="[878,1043,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="3.3649999999999998" metricValueMax="3.65" metricValueMin="3.08" pageId="9" pageNumber="90" unit="mm" value="33.65" valueMax="36.5" valueMin="30.8">30.8–36.5 mm</quantity>
|
||
SL),
|
||
<date id="9F4B10545F6DFFC655689011B4DBFEA2" box="[1106,1256,294,321]" pageId="9" pageNumber="90" value="2018-05-31">
|
||
<collectingDate id="8F0FE9BC5F6DFFC655689011B4DBFEA2" box="[1106,1256,294,321]" pageId="9" pageNumber="90" value="2018-05-31">31 May 2018</collectingDate>
|
||
</date>
|
||
coll.
|
||
<collectorName id="460053425F6DFFC6541D9010B5ABFEA2" box="[1319,1432,294,321]" pageId="9" pageNumber="90">K. Maeda</collectorName>
|
||
, H. Kobayashi, and
|
||
<collectorName id="460053425F6DFFC65057907CB1D6FE86" box="[365,485,330,357]" pageId="9" pageNumber="90">H. P. Palla</collectorName>
|
||
; WPU-PPC-P 65–69,
|
||
<specimenCount id="FDF3FD1D5F6DFFC653E0907DB31CFE86" box="[730,815,330,356]" pageId="9" pageNumber="90" type="male">3 males</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC65204907DB3D1FE87" box="[830,994,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="4.485" metricValueMax="4.61" metricValueMin="4.36" pageId="9" pageNumber="90" unit="mm" value="44.85" valueMax="46.1" valueMin="43.6">43.6–46.1 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="FDF3FD1D5F6DFFC6557D907DB480FE86" box="[1095,1203,330,356]" pageId="9" pageNumber="90" type="female">2 females</specimenCount>
|
||
(
|
||
<quantity id="2C0D9B715F6DFFC655FB907DB556FE87" box="[1217,1381,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="4.045" metricValueMax="4.13" metricValueMin="3.96" pageId="9" pageNumber="90" unit="mm" value="40.45" valueMax="41.3" valueMin="39.6">39.6–41.3 mm</quantity>
|
||
SL), same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A3EF651F5F6DFFC651FD90A5B4CCFD66" pageId="9" pageNumber="90" type="diagnosis">
|
||
<paragraph id="EB4A36945F6DFFC651FD90A5B4CCFD66" blockId="9.[151,1437,150,2038]" pageId="9" pageNumber="90">
|
||
<emphasis id="D981EA865F6DFFC651FD90A5B171FE4E" bold="true" box="[199,322,402,428]" pageId="9" pageNumber="90">Diagnosis.</emphasis>
|
||
Pectoral fin with 16 or 17 rays. Nape and posterior part of occipital region covered by cycloid scales; scaled area extending anteriorly to around a vertical line through posterior margin of preopercle or a little posterior to this line. Longitudinal scales 29–34, predorsal scales 5–13. Number of vertebrae 26. Cephalic sensory pore system usually with B´, C, D(S), E, F, H´, K´, L´, M´, N, and O´, but often lacking one or both side(s) of pore C. Transverse rows of sensory papillae on cheek. In preservative, lateral and dorsal sides of body without clear meshlike markings, obscure markings on cheek, and upper half of pectoral fin with dark brown band vertically across proximal part. In life, pectoral-fin base with a bright white vertical band reaching near bottom of fin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A3EF651F5F6DFFC551FD93B9B251F98C" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90" type="description">
|
||
<paragraph id="EB4A36945F6DFFC651FD93B9B4BAFBBA" blockId="9.[151,1437,150,2038]" pageId="9" pageNumber="90">
|
||
<emphasis id="D981EA865F6DFFC651FD93B9B16AFD4A" bold="true" box="[199,345,654,680]" pageId="9" pageNumber="90">Description.</emphasis>
|
||
Body nearly cylindrical anteriorly and somewhat compressed posteriorly. Abdomen of male often thin. Head depressed and larger in male than female (head length 32.1–37.4 vs 30.8–33.4% of SL;
|
||
<figureCitation id="73CE2A115F6DFFC654339385B57DFD2E" box="[1289,1358,690,716]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[217,1372,191,1911]" captionTargetId="figure-21@6.[200,1387,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Head length, upper-jaw length, second dorsal-fin length, anal-fin length, and caudal-fin length of Rhinogobius estrellae (red circles) and Rhinogobius tandikan (blue triangles). Solid and open symbols represent males and females, respectively." figureDoi="http://doi.org/10.5281/zenodo.5702110" httpUri="https://zenodo.org/record/5702110/files/figure.png" pageId="9" pageNumber="90">Fig. 3</figureCitation>
|
||
). Eyes located dorsolaterally. Mouth terminal and oblique with thick upper and lower lips. Anterior tips of upper and lower jaws almost even or upper jaw slightly protruding beyond lower jaw. Posterior end of upper jaw always exceeding a vertical line through anterior margin of eye. Mouth larger in male than female (upper-jaw length 13.3–20.9 vs 10.1–12.7% of SL;
|
||
<figureCitation id="73CE2A115F6DFFC650AF9275B1EAFCBE" box="[405,473,834,860]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[217,1372,191,1911]" captionTargetId="figure-21@6.[200,1387,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Head length, upper-jaw length, second dorsal-fin length, anal-fin length, and caudal-fin length of Rhinogobius estrellae (red circles) and Rhinogobius tandikan (blue triangles). Solid and open symbols represent males and females, respectively." figureDoi="http://doi.org/10.5281/zenodo.5702110" httpUri="https://zenodo.org/record/5702110/files/figure.png" pageId="9" pageNumber="90">Fig. 3</figureCitation>
|
||
). Anterior nostril short tubular, posterior nostril a pore. Cephalic sensory pore system usually with B´, C, D(S), E, F, H´, K´, and L´ in oculoscapular canal and M´, N, and O´ in preopercular canal, but 9/
|
||
<specimenCount id="FDF3FD1D5F6DFFC6519792BDB176FC46" box="[173,325,906,932]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
lacking one pore C, 7/
|
||
<specimenCount id="FDF3FD1D5F6DFFC6537992BDB2E9FC46" box="[579,730,906,932]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
lacking both pores C, 4/
|
||
<specimenCount id="FDF3FD1D5F6DFFC652D792BDB4B7FC46" box="[1005,1156,906,932]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
lacking one pore E, 1/
|
||
<specimenCount id="FDF3FD1D5F6DFFC6544492BDB132FC2A" pageId="9" pageNumber="90" type="generic">29 specimen</specimenCount>
|
||
having two pores E on one side, 4/
|
||
<specimenCount id="FDF3FD1D5F6DFFC653B99299B32AFC2A" box="[643,793,942,968]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
having pore G on one side, 1/
|
||
<specimenCount id="FDF3FD1D5F6DFFC655659299B4DDFC2A" box="[1119,1262,942,968]" pageId="9" pageNumber="90" type="generic">29 specimen</specimenCount>
|
||
having pores G on both sides, and 5/
|
||
<specimenCount id="FDF3FD1D5F6DFFC6504192E5B220FC0E" box="[379,531,978,1005]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
lacking one pore N. One specimen having combined pores K´ and L´ on one side (to be a groove without roof). Arrangement of cutaneous sensory papillae of head shown in
|
||
<figureCitation id="73CE2A115F6DFFC655BD92C1B4F9FBF3" box="[1159,1226,1014,1041]" captionStart="FIGURE 7" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[328,1254,188,1905]" captionTargetId="figure-20@11.[306,1282,181,1919]" captionTargetPageId="11" captionText="FIGURE 7. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius tandikan (URM-P 49307, paratype). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.5702118" httpUri="https://zenodo.org/record/5702118/files/figure.png" pageId="9" pageNumber="90">Fig. 7</figureCitation>
|
||
. Cheek having two longitudinal rows of papillae and 3–6 transverse rows both between eye and upper longitudinal row and between upper and lower longitudinal rows. Vertebrae 10+16=26 (n=10), P-V 3/II II I I 0/9 (n=10).
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F6DFFC651FD9555B375F9AB" blockId="9.[151,1437,150,2038]" pageId="9" pageNumber="90">
|
||
All males had damage to the caudal fin, except
|
||
<typeStatus id="344E88365F6DFFC653CB9555B364FB9E" box="[753,855,1122,1148]" pageId="9" pageNumber="90" type="holotype">holotype</typeStatus>
|
||
; 10/
|
||
<specimenCount id="FDF3FD1D5F6DFFC652B39555B3CAFB9E" box="[905,1017,1122,1148]" pageId="9" pageNumber="90" type="female">16 female</specimenCount>
|
||
specimens with damaged caudal fin (
|
||
<figureCitation id="73CE2A115F6DFFC651A595B1B0DDFB42" box="[159,238,1158,1184]" captionStart="FIGURE 1" captionStartId="3.[151,250,1837,1861]" captionTargetBox="[157,1419,1035,1802]" captionTargetId="figure-341@3.[151,1436,1027,1812]" captionTargetPageId="3" captionText="FIGURE 1. Preserved specimens of Rhinogobius estrellae (a–d, paratypes) and Rhinogobius tandikan (e–h, paratypes). Specimens in the left side are males, the right side are females. a, URM-P 49305, 39.6 mm SL; b, WPU-PPC-P 55, 38.4 mm SL; c, URM-P 49296, 36.1 mm SL; d, URM-P 49306, 36.4 mm SL; e, URM-P 49314, 46.9 mm SL; f, URM-P 49315, 46.5 mm SL; g, URM-P 49318, 45.2 mm SL; h, URM-P 49320, 37.9 mm SL (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.5702106" httpUri="https://zenodo.org/record/5702106/files/figure.png" pageId="9" pageNumber="90">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="73CE2A115F6DFFC651C795B1B13EFB42" box="[253,269,1158,1184]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[217,1372,191,1911]" captionTargetId="figure-21@6.[200,1387,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Head length, upper-jaw length, second dorsal-fin length, anal-fin length, and caudal-fin length of Rhinogobius estrellae (red circles) and Rhinogobius tandikan (blue triangles). Solid and open symbols represent males and females, respectively." figureDoi="http://doi.org/10.5281/zenodo.5702110" httpUri="https://zenodo.org/record/5702110/files/figure.png" pageId="9" pageNumber="90">3</figureCitation>
|
||
). Other fins of male and female sometimes damaged also. First dorsal fin with six spines supported by six pterygiophores. Second dorsal fin usually with one spine and eight soft rays. First and second dorsal-fin bases separated each other by a small interval (0.2–4.4% of SL). First dorsal fin rounded, usually almost semi-circular; posterior tips of fin (tips of fourth to sixth spines) not reaching second dorsal fin origin (N=9), just touching base of second dorsal-fin spine (n=12), or exceeding base of second dorsal-fin spine but not reaching base of first soft ray of second dorsal fin (n=8). Anal fin usually with one spine and eight soft rays, but 4/
|
||
<specimenCount id="FDF3FD1D5F6DFFC655119400B4F0FAB3" box="[1067,1219,1335,1361]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
with one spine and seven soft rays. Caudal fin usually with 17 segmented rays but 3/
|
||
<specimenCount id="FDF3FD1D5F6DFFC65259946CB3C9FA96" box="[867,1018,1370,1397]" pageId="9" pageNumber="90" type="generic">29 specimens</specimenCount>
|
||
with 16 segmented rays, including 11 (n=1), 12 (n=6), 13 (n=15), or 14 (n=22) branched rays (unknown in
|
||
<specimenCount id="FDF3FD1D5F6DFFC652999449B47DFA7A" box="[931,1102,1406,1432]" pageId="9" pageNumber="90" type="generic">four specimens</specimenCount>
|
||
due to fin damage); posterior margin rounded in non-damaged individuals. Male having larger second dorsal and anal fins than female (second dorsal-fin length 33.8–43.6 vs 29.5–35.3% of SL and anal-fin length 30.2–38.3 vs 26.7–32.7% of SL), and nondamaged male with larger caudal fin than female (
|
||
<figureCitation id="73CE2A115F6DFFC653F694DFB323F9E0" box="[716,784,1512,1538]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[217,1372,191,1911]" captionTargetId="figure-21@6.[200,1387,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Head length, upper-jaw length, second dorsal-fin length, anal-fin length, and caudal-fin length of Rhinogobius estrellae (red circles) and Rhinogobius tandikan (blue triangles). Solid and open symbols represent males and females, respectively." figureDoi="http://doi.org/10.5281/zenodo.5702110" httpUri="https://zenodo.org/record/5702110/files/figure.png" pageId="9" pageNumber="90">Fig. 3</figureCitation>
|
||
). Pectoral fin with 16 or 17 rays (
|
||
<tableCitation id="A677032F5F6DFFC655B194DFB4D1F9E0" box="[1163,1250,1512,1538]" captionStart="TABLE 2" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 2. Pectoral-fin ray counts of Rhinogobius estrellae and Rhinogobius tandikan." httpUri="http://table.plazi.org/id/BF8A661C5F61FFCA51AD91A0B406FF4D" pageId="9" pageNumber="90" tableUuid="BF8A661C5F61FFCA51AD91A0B406FF4D">Table 2</tableCitation>
|
||
). Pelvic fin with one spine and five soft rays; pelvic fins joined together to form a cuplike disk with fleshy bilobed frenum. Posterior tip of pelvic fin located below middle of first-dorsal-fin base.
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F6DFFC651FD9764B4F6F884" blockId="9.[151,1437,150,2038]" pageId="9" pageNumber="90">
|
||
Ctenoid scales covering lateral side of body and dorsal and ventral sides of caudal peduncle. Nape and posterior part of occipital region covered by cycloid scales; scaled area extending anteriorly around vertical line through posterior margin of preopercle (above pore H´) or to a little posterior to this line (above area between pores H´ and K´). Other regions in head naked. Belly and first dorsal-fin base covered with cycloid scales. A few cycloid scales also present on trunk behind pectoral fin, along bases of second dorsal, anal, and caudal fins, and on proximal part of caudal fin. Pectoral-fin base and breast (prepelvic area) probably naked (at least invisible without staining by arizarin red; see
|
||
<bibRefCitation id="8F644B655F6DFFC650689610B216F8A0" author="Suzuki, T. & Shibukawa, K. & Senou, H. & Chen, I. - S." box="[338,549,1831,1858]" pageId="9" pageNumber="90" pagination="227 - 238" refId="ref11378" refString="Suzuki, T., Shibukawa, K., Senou, H. & Chen, I. - S. (2016) Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, 63 (2), 227 - 238. https: // doi. org / 10.1007 / s 10228 - 015 - 0494 - 3" type="journal article" year="2016">
|
||
Suzuki
|
||
<emphasis id="D981EA865F6DFFC65092961FB1D3F8A3" box="[424,480,1831,1857]" italics="true" pageId="9" pageNumber="90">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). Longitudinal scales 29–34 (usually 30 or 31), transverse scales 8–10 (usually 9 or 10), transverse scales in caudal peduncle always 7, and predorsal scales 5–13 (
|
||
<tableCitation id="A677032F5F6DFFC65503967CB48BF884" box="[1081,1208,1867,1894]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStart-2="TABLE 5" captionStartId-0="5.[151,239,675,699]" captionStartId-1="5.[151,239,869,893]" captionStartId-2="5.[151,239,1063,1087]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="TABLE 3. Longitudinal scale counts of Rhinogobius estrellae and Rhinogobius tandikan." captionText-1="TABLE 4. Transverse scale counts of Rhinogobius estrellae and Rhinogobius tandikan." captionText-2="TABLE 5. Predorsal scale counts of Rhinogobius estrellae and Rhinogobius tandikan." pageId="9" pageNumber="90">Tables 3–5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F6DFFC551FD9658B173FEA3" blockId="9.[151,1437,150,2038]" lastBlockId="10.[151,1437,151,321]" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90">
|
||
Colour in preservative (
|
||
<figureCitation id="73CE2A115F6DFFC650F49658B223F86B" box="[462,528,1903,1929]" captionStart="FIGURE 1" captionStartId="3.[151,250,1837,1861]" captionTargetBox="[157,1419,1035,1802]" captionTargetId="figure-341@3.[151,1436,1027,1812]" captionTargetPageId="3" captionText="FIGURE 1. Preserved specimens of Rhinogobius estrellae (a–d, paratypes) and Rhinogobius tandikan (e–h, paratypes). Specimens in the left side are males, the right side are females. a, URM-P 49305, 39.6 mm SL; b, WPU-PPC-P 55, 38.4 mm SL; c, URM-P 49296, 36.1 mm SL; d, URM-P 49306, 36.4 mm SL; e, URM-P 49314, 46.9 mm SL; f, URM-P 49315, 46.5 mm SL; g, URM-P 49318, 45.2 mm SL; h, URM-P 49320, 37.9 mm SL (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.5702106" httpUri="https://zenodo.org/record/5702106/files/figure.png" pageId="9" pageNumber="90">Fig. 1</figureCitation>
|
||
): Background of head and body brown dorsally and laterally and cream ventrally. Snout with obscure dark brown stripe connecting upper lip and eye. Cheek often with two obscure, dark brown intermittent lines. Posterior margin of opercle fringed by dark brown vertical band. Pectoral-fin base blackish except cream or pale brown bottom. Four to eight dark brown transverse bars or round blotches aligned along lateral midline of body. First and second dorsal fins pale grey with 3–7 brown horizontal stripes on proximal and middle parts, but anterior half of first dorsal fin lacking this marking. Anal fin greyish brown but light brown proximally. Caudal fin pale grey or greyish brown with 6–11 dark brown vertical bands, but dorsal and ventral parts lacking these bands. Pectoral fin pale greyish brown with a dark brown vertical band proximally on upper half. Pelvic fin greyish brown.
|
||
</paragraph>
|
||
<caption id="BF8A661C5F6EFFC551AD95B6B2CBFB5C" ID-DOI="http://doi.org/10.5281/zenodo.5702116" ID-Zenodo-Dep="5702116" httpUri="https://zenodo.org/record/5702116/files/figure.png" pageId="10" pageNumber="91" startId="10.[151,250,1153,1177]" targetBox="[151,1436,345,1130]" targetPageId="10">
|
||
<paragraph id="EB4A36945F6EFFC551AD95B6B2CBFB5C" blockId="10.[151,1436,1153,1214]" pageId="10" pageNumber="91">
|
||
<emphasis id="D981EA865F6EFFC551AD95B6B126FB78" bold="true" box="[151,277,1153,1178]" pageId="10" pageNumber="91">FIGURE 6.</emphasis>
|
||
<taxonomicName id="2CF54D175F6EFFC5502795B5B1CCFB78" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[285,511,1154,1178]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F6EFFC5502795B5B1CCFB78" box="[285,511,1154,1178]" italics="true" pageId="10" pageNumber="91">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
immediately after fixation. a, NSMT-P 140093, holotype, male, 43.6 mm SL; b, URM-P 49316, paratype, female, 44.8 mm SL (photo by K. Maeda).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EB4A36945F6EFFC551FD95DBB251F98C" blockId="10.[151,1437,1260,2042]" pageId="10" pageNumber="91">
|
||
Colour in life (
|
||
<figureCitation id="73CE2A115F6EFFC5505495DBB180FAE4" box="[366,435,1260,1286]" captionStart="FIGURE 8" captionStartId="12.[151,250,687,711]" captionTargetBox="[151,1437,181,663]" captionTargetId="figure-21@12.[151,1437,181,663]" captionTargetPageId="12" captionText="FIGURE 8. In situ underwater photographs of Rhinogobius tandikan in Cayulo River, Palawan. a, male, 31 May 2018; b, female, 1 June 2018 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.5702120" httpUri="https://zenodo.org/record/5702120/files/figure.png" pageId="10" pageNumber="91">Fig. 8</figureCitation>
|
||
): Background of head and body olive green or yellowish grey. Snout with reddish brown stripe connecting upper lip and eye. Cheek with two reddish brown longitudinal intermittent lines and several reddish brown spots. Posterior margin of opercle fringed by reddish-brown vertical band. Pectoral-fin base black or dusky with a large greyish blue blotch anteriorly, and with a bright white vertical band posteriorly. Four to eight, dark brown transverse bars or round blotches aligned along lateral midline of body. Irregularly arranged dark brown markings dorsally on body. Centre of lateral scales tinged with sky blue. First and second dorsal fins light yellowish brown with 3–7 dark brown horizontal stripes on proximal and middle parts and reddish brown on distal part, but anterior half of first dorsal fin reddish brown without clear marking. Anal fin reddish brown or orange but yellowish proximally. Caudal fin reddish brown dorsally and ventrally and whitish with 6–11 dark reddish-brown transverse bands on middle part. Pectoral fin translucent but proximal parts of some ventral rays white. Pelvic fin reddish brown proximally and light grey distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A3EF651F5F6EFFC551FD974FB44CF888" pageId="10" pageNumber="91" type="distribution">
|
||
<paragraph id="EB4A36945F6EFFC551FD974FB44CF888" blockId="10.[151,1437,1260,2042]" pageId="10" pageNumber="91">
|
||
<emphasis id="D981EA865F6EFFC551FD974FB152F970" bold="true" box="[199,353,1656,1682]" pageId="10" pageNumber="91">Distribution.</emphasis>
|
||
<taxonomicName id="2CF54D175F6EFFC55051974FB256F970" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[363,613,1656,1682]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F6EFFC55051974FB256F970" box="[363,613,1656,1682]" italics="true" pageId="10" pageNumber="91">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
is endemic to Palawan Island. It was found only in the Cayulo River, Bahile, northern part of Puerto Princesa City. The Cayulo River is a small stream (length approximately
|
||
<quantity id="2C0D9B715F6EFFC5540197ABB546F954" box="[1339,1397,1692,1718]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="91" unit="km" value="4.0">4 km</quantity>
|
||
including
|
||
<quantity id="2C0D9B715F6EFFC551C997F7B11FF938" box="[243,300,1728,1754]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="91" unit="km" value="1.0">1 km</quantity>
|
||
of an estuary) flowing into Ulugan Bay on South
|
||
<collectingCountry id="93E276045F6EFFC5525E97F7B394F938" box="[868,935,1728,1754]" name="China" pageId="10" pageNumber="91">China</collectingCountry>
|
||
Sea-side of the island. The stream system is surrounded by mountains of about
|
||
<quantity id="2C0D9B715F6EFFC5531B97D3B293F91C" box="[545,672,1764,1790]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="10" pageNumber="91" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</quantity>
|
||
above sea level. Cayulo Falls, composed of a series of small waterfalls, is located at the middle eaches of this stream (
|
||
<geoCoordinate id="8EC150535F6EFFC553E7963FB351F8C0" box="[733,866,1800,1826]" degrees="10" direction="north" minutes="02" orientation="latitude" pageId="10" pageNumber="91" precision="15" seconds="12" value="10.036667">10°02'12"N</geoCoordinate>
|
||
<geoCoordinate id="8EC150535F6EFFC55253963FB3C9F8C1" box="[873,1018,1800,1827]" degrees="118" direction="east" minutes="45" orientation="longitude" pageId="10" pageNumber="91" precision="15" seconds="03" value="118.75083">118°45'03"E</geoCoordinate>
|
||
).
|
||
<taxonomicName id="2CF54D175F6EFFC55534963FB536F8C0" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[1038,1285,1800,1826]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F6EFFC55534963FB536F8C0" box="[1038,1285,1800,1826]" italics="true" pageId="10" pageNumber="91">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
was found at both the lower and upper reaches of this waterfall, as well as pools in the middle of the falls. The new species was abundant in this site with many other goby species, cyprinids, and a jungle perch species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A3EF651F5F6EFFC251FD9643B2FBFBBB" lastPageId="13" lastPageNumber="94" pageId="10" pageNumber="91" type="etymology">
|
||
<paragraph id="EB4A36945F6EFFC551FD9643B47CF818" blockId="10.[151,1437,1260,2042]" pageId="10" pageNumber="91">
|
||
<emphasis id="D981EA865F6EFFC551FD9643B17DF86C" bold="true" box="[199,334,1908,1934]" pageId="10" pageNumber="91">Etymology.</emphasis>
|
||
Tandikan is a local name of
|
||
<collectingRegion id="2931F8765F6EFFC553A39643B2CBF86C" box="[665,760,1908,1934]" country="Philippines" name="Palawan" pageId="10" pageNumber="91">Palawan</collectingRegion>
|
||
peacock-pheasant,
|
||
<taxonomicName id="2CF54D175F6EFFC552EC9643B570F86C" authority="Lesson" authorityName="Lesson" box="[982,1347,1908,1934]" class="Aves" family="Phasianidae" genus="Polyplectron" kingdom="Animalia" order="Galliformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="napoleonis">
|
||
<emphasis id="D981EA865F6EFFC552EC9643B4DAF86C" box="[982,1257,1908,1934]" italics="true" pageId="10" pageNumber="91">Polyplectron napoleonis</emphasis>
|
||
Lesson
|
||
</taxonomicName>
|
||
, known as a symbol of
|
||
<collectingRegion id="2931F8765F6EFFC5500596AFB214F850" box="[319,551,1944,1970]" country="Philippines" name="Puerto Princesa" pageId="10" pageNumber="91">Puerto Princesa City</collectingRegion>
|
||
. The new goby species is named
|
||
<taxonomicName id="2CF54D175F6EFFC552A396AFB4BCF850" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[921,1167,1944,1970]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="91" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F6EFFC552A396AFB4BCF850" box="[921,1167,1944,1970]" italics="true" pageId="10" pageNumber="91">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
, as it has blue markings on the body, suggestive of the plumage of the
|
||
<collectingRegion id="2931F8765F6EFFC5539B968BB333F834" box="[673,768,1980,2006]" country="Philippines" name="Palawan" pageId="10" pageNumber="91">Palawan</collectingRegion>
|
||
peacock-pheasant, and as the
|
||
<typeStatus id="344E88365F6EFFC5556C968BB4B5F834" box="[1110,1158,1980,2006]" pageId="10" pageNumber="91">type</typeStatus>
|
||
locality is located in the northern part of
|
||
<collectingRegion id="2931F8765F6EFFC5507496D7B205F818" box="[334,566,2016,2042]" country="Philippines" name="Puerto Princesa" pageId="10" pageNumber="91">Puerto Princesa City</collectingRegion>
|
||
. The new specific name is a noun in apposition.
|
||
</paragraph>
|
||
<caption id="BF8A661C5F6FFFC451AD96A0B455F836" ID-DOI="http://doi.org/10.5281/zenodo.5702118" ID-Zenodo-Dep="5702118" httpUri="https://zenodo.org/record/5702118/files/figure.png" pageId="11" pageNumber="92" startId="11.[151,250,1943,1967]" targetBox="[328,1254,188,1905]" targetPageId="11">
|
||
<paragraph id="EB4A36945F6FFFC451AD96A0B455F836" blockId="11.[151,1436,1943,2004]" pageId="11" pageNumber="92">
|
||
<emphasis id="D981EA865F6FFFC451AD96A0B127F852" bold="true" box="[151,276,1943,1968]" pageId="11" pageNumber="92">FIGURE 7.</emphasis>
|
||
Arrangement of cephalic sensory pores and cutaneous sensory papillae in
|
||
<taxonomicName id="2CF54D175F6FFFC4552A96AFB4C2F852" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[1040,1265,1944,1968]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="92" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F6FFFC4552A96AFB4C2F852" box="[1040,1265,1944,1968]" italics="true" pageId="11" pageNumber="92">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
(URM-P 49307, paratype). a, lateral view; b, dorsal view; c, ventral view. AN, anterior naris; PN, posterior naris.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BF8A661C5F68FFC351AD9398B201FD09" ID-DOI="http://doi.org/10.5281/zenodo.5702120" ID-Zenodo-Dep="5702120" httpUri="https://zenodo.org/record/5702120/files/figure.png" pageId="12" pageNumber="93" startId="12.[151,250,687,711]" targetBox="[151,1437,181,663]" targetPageId="12">
|
||
<paragraph id="EB4A36945F68FFC351AD9398B201FD09" blockId="12.[151,1436,687,748]" pageId="12" pageNumber="93">
|
||
<emphasis id="D981EA865F68FFC351AD9398B127FD25" bold="true" box="[151,276,687,711]" pageId="12" pageNumber="93">FIGURE 8.</emphasis>
|
||
In situ underwater photographs of
|
||
<taxonomicName id="2CF54D175F68FFC353479398B36EFD25" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[637,861,687,711]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F68FFC353479398B36EFD25" box="[637,861,687,711]" italics="true" pageId="12" pageNumber="93">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
in Cayulo River, Palawan. a, male, 31 May 2018; b, female, 1 June 2018 (photo by K. Maeda).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BF8A661C5F68FFC351AD922CB4C0FCB5" ID-Table-UUID="BF8A661C5F68FFC351AD922CB4C0FCB5" httpUri="http://table.plazi.org/id/BF8A661C5F68FFC351AD922CB4C0FCB5" pageId="12" pageNumber="93" startId="12.[151,239,795,819]" targetBox="[164,1413,872,1857]" targetIsTable="true" targetPageId="12">
|
||
<paragraph id="EB4A36945F68FFC351AD922CB4C0FCB5" blockId="12.[151,1436,795,862]" pageId="12" pageNumber="93">
|
||
<emphasis id="D981EA865F68FFC351AD922CB13FFCD1" bold="true" box="[151,268,795,819]" pageId="12" pageNumber="93">TABLE 6.</emphasis>
|
||
Morphometrics of
|
||
<taxonomicName id="2CF54D175F68FFC350DE922CB2FCFCD1" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[484,719,795,819]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F68FFC350DE922CB2FCFCD1" box="[484,719,795,819]" italics="true" pageId="12" pageNumber="93">Rhinogobius estrellae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2CF54D175F68FFC35232922CB3C6FCD1" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[776,1013,795,819]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F68FFC35232922CB3C6FCD1" box="[776,1013,795,819]" italics="true" pageId="12" pageNumber="93">Rhinogobius tandikan</emphasis>
|
||
</taxonomicName>
|
||
expressed as a percentage of standard length. D
|
||
<subScript id="777134D15F68FFC351C09267B131FCBC" attach="left" box="[250,258,848,862]" fontSize="6" pageId="12" pageNumber="93">1</subScript>
|
||
, first dorsal fin; D
|
||
<subScript id="777134D15F68FFC350F29267B1E3FCBC" attach="left" box="[456,464,848,862]" fontSize="6" pageId="12" pageNumber="93">2</subScript>
|
||
, second dorsal fin; A, anal fin; C, caudal fin; P
|
||
<subScript id="777134D15F68FFC352FF9267B3FEFCBC" attach="left" box="[965,973,848,862]" fontSize="6" pageId="12" pageNumber="93">1</subScript>
|
||
, pectoral fin; P
|
||
<subScript id="777134D15F68FFC3554B9267B44AFCBC" attach="left" box="[1137,1145,848,862]" fontSize="6" pageId="12" pageNumber="93">2</subScript>
|
||
, pelvic fin.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EB4A36945F68FFC353A4925FB54DF8A3" pageId="12" pageNumber="93">
|
||
<table id="99F5C4345F680030519E925FB5B6F8A3" box="[164,1413,872,1857]" gridcols="7" gridrows="24" pageId="12" pageNumber="93">
|
||
<tr id="55C534D65F680030519E925FB5B6FC62" box="[164,1413,872,896]" gridrow="0" pageId="12" pageNumber="93" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
|
||
<th id="16145DAA5F68003053A1925FB323FC62" box="[667,784,872,896]" gridcol="2" gridrow="0" pageId="12" pageNumber="93">
|
||
<taxonomicName id="2CF54D175F68FFC353A4925FB323FC62" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[670,784,872,896]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F68FFC353A4925FB323FC62" box="[670,784,872,896]" italics="true" pageId="12" pageNumber="93">R. estrellae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="16145DAA5F68003055BA925FB4C6FC62" box="[1152,1269,872,896]" gridcol="5" gridrow="0" pageId="12" pageNumber="93">
|
||
<taxonomicName id="2CF54D175F68FFC355BA925FB4C6FC62" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[1152,1269,872,896]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F68FFC355BA925FB4C6FC62" box="[1152,1269,872,896]" italics="true" pageId="12" pageNumber="93">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E92A7B5B6FC4A" box="[164,1413,912,936]" gridrow="1" pageId="12" pageNumber="93" rowspan-0="1">
|
||
<td id="16145DAA5F680030533692A7B259FC4A" box="[524,618,912,936]" gridcol="1" gridrow="1" pageId="12" pageNumber="93">Holotype</td>
|
||
<td id="16145DAA5F68003053A192A7B323FC4A" box="[667,784,912,936]" gridcol="2" gridrow="1" pageId="12" pageNumber="93">Paratypes</td>
|
||
<td id="16145DAA5F680030520D92A7B3AEFC4A" box="[823,925,912,936]" gridcol="3" gridrow="1" pageId="12" pageNumber="93">Paratypes</td>
|
||
<td id="16145DAA5F68003052D592A7B47EFC4A" box="[1007,1101,912,936]" gridcol="4" gridrow="1" pageId="12" pageNumber="93">Holotype</td>
|
||
<td id="16145DAA5F68003055BA92A7B4C6FC4A" box="[1152,1269,912,936]" gridcol="5" gridrow="1" pageId="12" pageNumber="93">Paratypes</td>
|
||
<td id="16145DAA5F680030542592A7B5B6FC4A" box="[1311,1413,912,936]" gridcol="6" gridrow="1" pageId="12" pageNumber="93">Paratypes</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E928FB5B6FC32" box="[164,1413,952,976]" gridrow="2" pageId="12" pageNumber="93" rowspan-0="1">
|
||
<td id="16145DAA5F6800305336928FB259FC32" box="[524,618,952,976]" gridcol="1" gridrow="2" pageId="12" pageNumber="93">Male</td>
|
||
<td id="16145DAA5F68003053A1928FB323FC32" box="[667,784,952,976]" gridcol="2" gridrow="2" pageId="12" pageNumber="93">Male</td>
|
||
<td id="16145DAA5F680030520D928FB3AEFC32" box="[823,925,952,976]" gridcol="3" gridrow="2" pageId="12" pageNumber="93">Female</td>
|
||
<td id="16145DAA5F68003052D5928FB47EFC32" box="[1007,1101,952,976]" gridcol="4" gridrow="2" pageId="12" pageNumber="93">Male</td>
|
||
<td id="16145DAA5F68003055BA928FB4C6FC32" box="[1152,1269,952,976]" gridcol="5" gridrow="2" pageId="12" pageNumber="93">Male</td>
|
||
<td id="16145DAA5F6800305425928FB5B6FC32" box="[1311,1413,952,976]" gridcol="6" gridrow="2" pageId="12" pageNumber="93">Female</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E92D7B5B6FC1B" box="[164,1413,992,1017]" gridrow="3" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E92D7B18BFC1B" box="[164,440,992,1017]" gridcol="0" gridrow="3" pageId="12" pageNumber="93">Number of specimens</th>
|
||
<td id="16145DAA5F680030533692D7B259FC1B" box="[524,618,992,1017]" gridcol="1" gridrow="3" pageId="12" pageNumber="93">1</td>
|
||
<td id="16145DAA5F68003053A192D7B323FC1B" box="[667,784,992,1017]" gridcol="2" gridrow="3" pageId="12" pageNumber="93">7</td>
|
||
<td id="16145DAA5F680030520D92D7B3AEFC1B" box="[823,925,992,1017]" gridcol="3" gridrow="3" pageId="12" pageNumber="93">16</td>
|
||
<td id="16145DAA5F68003052D592D7B47EFC1B" box="[1007,1101,992,1017]" gridcol="4" gridrow="3" pageId="12" pageNumber="93">1</td>
|
||
<td id="16145DAA5F68003055BA92D7B4C6FC1B" box="[1152,1269,992,1017]" gridcol="5" gridrow="3" pageId="12" pageNumber="93">12</td>
|
||
<td id="16145DAA5F680030542592D7B5B6FC1B" box="[1311,1413,992,1017]" gridcol="6" gridrow="3" pageId="12" pageNumber="93">16</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E953FB5B6FBC3" box="[164,1413,1032,1057]" gridrow="4" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E953FB18BFBC3" box="[164,440,1032,1057]" gridcol="0" gridrow="4" pageId="12" pageNumber="93">Standard length (mm)</th>
|
||
<td id="16145DAA5F6800305336953FB259FBC3" box="[524,618,1032,1057]" gridcol="1" gridrow="4" pageId="12" pageNumber="93">40.6</td>
|
||
<td id="16145DAA5F68003053A1953FB323FBC3" box="[667,784,1032,1057]" gridcol="2" gridrow="4" pageId="12" pageNumber="93">37.3–40.7</td>
|
||
<td id="16145DAA5F680030520D953FB3AEFBC3" box="[823,925,1032,1057]" gridcol="3" gridrow="4" pageId="12" pageNumber="93">33.2–45.3</td>
|
||
<td id="16145DAA5F68003052D5953FB47EFBC3" box="[1007,1101,1032,1057]" gridcol="4" gridrow="4" pageId="12" pageNumber="93">43.6</td>
|
||
<td id="16145DAA5F68003055BA953FB4C6FBC3" box="[1152,1269,1032,1057]" gridcol="5" gridrow="4" pageId="12" pageNumber="93">36.1–46.9</td>
|
||
<td id="16145DAA5F6800305425953FB5B6FBC3" box="[1311,1413,1032,1057]" gridcol="6" gridrow="4" pageId="12" pageNumber="93">30.8–45.7</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E9506B5B6FBAB" box="[164,1413,1073,1097]" gridrow="5" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E9506B18BFBAB" box="[164,440,1073,1097]" gridcol="0" gridrow="5" pageId="12" pageNumber="93">Head length</th>
|
||
<td id="16145DAA5F68003053369506B259FBAB" box="[524,618,1073,1097]" gridcol="1" gridrow="5" pageId="12" pageNumber="93">33.5</td>
|
||
<td id="16145DAA5F68003053A19506B323FBAB" box="[667,784,1073,1097]" gridcol="2" gridrow="5" pageId="12" pageNumber="93">33.9–36.0</td>
|
||
<td id="16145DAA5F680030520D9506B3AEFBAB" box="[823,925,1073,1097]" gridcol="3" gridrow="5" pageId="12" pageNumber="93">30.3–32.9</td>
|
||
<td id="16145DAA5F68003052D59506B47EFBAB" box="[1007,1101,1073,1097]" gridcol="4" gridrow="5" pageId="12" pageNumber="93">37.4</td>
|
||
<td id="16145DAA5F68003055BA9506B4C6FBAB" box="[1152,1269,1073,1097]" gridcol="5" gridrow="5" pageId="12" pageNumber="93">32.1–35.1</td>
|
||
<td id="16145DAA5F68003054259506B5B6FBAB" box="[1311,1413,1073,1097]" gridcol="6" gridrow="5" pageId="12" pageNumber="93">30.8–33.4</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E956EB5B6FB93" box="[164,1413,1113,1137]" gridrow="6" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E956EB18BFB93" box="[164,440,1113,1137]" gridcol="0" gridrow="6" pageId="12" pageNumber="93">Snout length</th>
|
||
<td id="16145DAA5F6800305336956EB259FB93" box="[524,618,1113,1137]" gridcol="1" gridrow="6" pageId="12" pageNumber="93">10.6</td>
|
||
<td id="16145DAA5F68003053A1956EB323FB93" box="[667,784,1113,1137]" gridcol="2" gridrow="6" pageId="12" pageNumber="93">11.5–12.7</td>
|
||
<td id="16145DAA5F680030520D956EB3AEFB93" box="[823,925,1113,1137]" gridcol="3" gridrow="6" pageId="12" pageNumber="93">9.0–10.7</td>
|
||
<td id="16145DAA5F68003052D5956EB47EFB93" box="[1007,1101,1113,1137]" gridcol="4" gridrow="6" pageId="12" pageNumber="93">13.5</td>
|
||
<td id="16145DAA5F68003055BA956EB4C6FB93" box="[1152,1269,1113,1137]" gridcol="5" gridrow="6" pageId="12" pageNumber="93">10.5–12.8</td>
|
||
<td id="16145DAA5F6800305425956EB5B6FB93" box="[1311,1413,1113,1137]" gridcol="6" gridrow="6" pageId="12" pageNumber="93">9.4–11.1</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E95B6B5B6FB78" box="[164,1413,1153,1178]" gridrow="7" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E95B6B18BFB78" box="[164,440,1153,1178]" gridcol="0" gridrow="7" pageId="12" pageNumber="93">Eye diameter</th>
|
||
<td id="16145DAA5F680030533695B6B259FB78" box="[524,618,1153,1178]" gridcol="1" gridrow="7" pageId="12" pageNumber="93">6.7</td>
|
||
<td id="16145DAA5F68003053A195B6B323FB78" box="[667,784,1153,1178]" gridcol="2" gridrow="7" pageId="12" pageNumber="93">6.3–7.4</td>
|
||
<td id="16145DAA5F680030520D95B6B3AEFB78" box="[823,925,1153,1178]" gridcol="3" gridrow="7" pageId="12" pageNumber="93">6.9–7.8</td>
|
||
<td id="16145DAA5F68003052D595B6B47EFB78" box="[1007,1101,1153,1178]" gridcol="4" gridrow="7" pageId="12" pageNumber="93">6.4</td>
|
||
<td id="16145DAA5F68003055BA95B6B4C6FB78" box="[1152,1269,1153,1178]" gridcol="5" gridrow="7" pageId="12" pageNumber="93">5.7–7.2</td>
|
||
<td id="16145DAA5F680030542595B6B5B6FB78" box="[1311,1413,1153,1178]" gridcol="6" gridrow="7" pageId="12" pageNumber="93">5.6–7.7</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E959EB5B6FB20" box="[164,1413,1193,1218]" gridrow="8" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E959EB18BFB20" box="[164,440,1193,1218]" gridcol="0" gridrow="8" pageId="12" pageNumber="93">Postorbital length of head</th>
|
||
<td id="16145DAA5F6800305336959EB259FB20" box="[524,618,1193,1218]" gridcol="1" gridrow="8" pageId="12" pageNumber="93">17.0</td>
|
||
<td id="16145DAA5F68003053A1959EB323FB20" box="[667,784,1193,1218]" gridcol="2" gridrow="8" pageId="12" pageNumber="93">16.9–17.7</td>
|
||
<td id="16145DAA5F680030520D959EB3AEFB20" box="[823,925,1193,1218]" gridcol="3" gridrow="8" pageId="12" pageNumber="93">14.8–17.0</td>
|
||
<td id="16145DAA5F68003052D5959EB47EFB20" box="[1007,1101,1193,1218]" gridcol="4" gridrow="8" pageId="12" pageNumber="93">18.3</td>
|
||
<td id="16145DAA5F68003055BA959EB4C6FB20" box="[1152,1269,1193,1218]" gridcol="5" gridrow="8" pageId="12" pageNumber="93">15.8–17.6</td>
|
||
<td id="16145DAA5F6800305425959EB5B6FB20" box="[1311,1413,1193,1218]" gridcol="6" gridrow="8" pageId="12" pageNumber="93">15.7–17.2</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E95E5B5B6FB08" box="[164,1413,1234,1258]" gridrow="9" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E95E5B18BFB08" box="[164,440,1234,1258]" gridcol="0" gridrow="9" pageId="12" pageNumber="93">Upper-jaw length</th>
|
||
<td id="16145DAA5F680030533695E5B259FB08" box="[524,618,1234,1258]" gridcol="1" gridrow="9" pageId="12" pageNumber="93">15.5</td>
|
||
<td id="16145DAA5F68003053A195E5B323FB08" box="[667,784,1234,1258]" gridcol="2" gridrow="9" pageId="12" pageNumber="93">14.1–16.1</td>
|
||
<td id="16145DAA5F680030520D95E5B3AEFB08" box="[823,925,1234,1258]" gridcol="3" gridrow="9" pageId="12" pageNumber="93">10.5–11.9</td>
|
||
<td id="16145DAA5F68003052D595E5B47EFB08" box="[1007,1101,1234,1258]" gridcol="4" gridrow="9" pageId="12" pageNumber="93">20.9</td>
|
||
<td id="16145DAA5F68003055BA95E5B4C6FB08" box="[1152,1269,1234,1258]" gridcol="5" gridrow="9" pageId="12" pageNumber="93">13.3–17.2</td>
|
||
<td id="16145DAA5F680030542595E5B5B6FB08" box="[1311,1413,1234,1258]" gridcol="6" gridrow="9" pageId="12" pageNumber="93">10.1–12.7</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E95CDB5B6FAFA" box="[164,1413,1274,1304]" gridrow="10" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E95CDB18BFAFA" box="[164,440,1274,1304]" gridcol="0" gridrow="10" pageId="12" pageNumber="93">Body depth at P2 origin</th>
|
||
<td id="16145DAA5F680030533695CDB259FAFA" box="[524,618,1274,1304]" gridcol="1" gridrow="10" pageId="12" pageNumber="93">18.5</td>
|
||
<td id="16145DAA5F68003053A195CDB323FAFA" box="[667,784,1274,1304]" gridcol="2" gridrow="10" pageId="12" pageNumber="93">16.5–18.0</td>
|
||
<td id="16145DAA5F680030520D95CDB3AEFAFA" box="[823,925,1274,1304]" gridcol="3" gridrow="10" pageId="12" pageNumber="93">15.7–19.7</td>
|
||
<td id="16145DAA5F68003052D595CDB47EFAFA" box="[1007,1101,1274,1304]" gridcol="4" gridrow="10" pageId="12" pageNumber="93">17.4</td>
|
||
<td id="16145DAA5F68003055BA95CDB4C6FAFA" box="[1152,1269,1274,1304]" gridcol="5" gridrow="10" pageId="12" pageNumber="93">14.5–17.8</td>
|
||
<td id="16145DAA5F680030542595CDB5B6FAFA" box="[1311,1413,1274,1304]" gridcol="6" gridrow="10" pageId="12" pageNumber="93">15.5–18.8</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E9415B5B6FAD9" box="[164,1413,1314,1339]" gridrow="11" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E9415B18BFAD9" box="[164,440,1314,1339]" gridcol="0" gridrow="11" pageId="12" pageNumber="93">Body depth at A origin</th>
|
||
<td id="16145DAA5F68003053369415B259FAD9" box="[524,618,1314,1339]" gridcol="1" gridrow="11" pageId="12" pageNumber="93">16.3</td>
|
||
<td id="16145DAA5F68003053A19415B323FAD9" box="[667,784,1314,1339]" gridcol="2" gridrow="11" pageId="12" pageNumber="93">14.9–16.9</td>
|
||
<td id="16145DAA5F680030520D9415B3AEFAD9" box="[823,925,1314,1339]" gridcol="3" gridrow="11" pageId="12" pageNumber="93">14.8–17.9</td>
|
||
<td id="16145DAA5F68003052D59415B47EFAD9" box="[1007,1101,1314,1339]" gridcol="4" gridrow="11" pageId="12" pageNumber="93">16.1</td>
|
||
<td id="16145DAA5F68003055BA9415B4C6FAD9" box="[1152,1269,1314,1339]" gridcol="5" gridrow="11" pageId="12" pageNumber="93">13.6–16.6</td>
|
||
<td id="16145DAA5F68003054259415B5B6FAD9" box="[1311,1413,1314,1339]" gridcol="6" gridrow="11" pageId="12" pageNumber="93">15.7–18.8</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E947DB5B6FA81" box="[164,1413,1354,1379]" gridrow="12" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E947DB18BFA81" box="[164,440,1354,1379]" gridcol="0" gridrow="12" pageId="12" pageNumber="93">Depth at caudal peduncle</th>
|
||
<td id="16145DAA5F6800305336947DB259FA81" box="[524,618,1354,1379]" gridcol="1" gridrow="12" pageId="12" pageNumber="93">12.8</td>
|
||
<td id="16145DAA5F68003053A1947DB323FA81" box="[667,784,1354,1379]" gridcol="2" gridrow="12" pageId="12" pageNumber="93">11.5–12.7</td>
|
||
<td id="16145DAA5F680030520D947DB3AEFA81" box="[823,925,1354,1379]" gridcol="3" gridrow="12" pageId="12" pageNumber="93">10.8–12.8</td>
|
||
<td id="16145DAA5F68003052D5947DB47EFA81" box="[1007,1101,1354,1379]" gridcol="4" gridrow="12" pageId="12" pageNumber="93">12.2</td>
|
||
<td id="16145DAA5F68003055BA947DB4C6FA81" box="[1152,1269,1354,1379]" gridcol="5" gridrow="12" pageId="12" pageNumber="93">10.1–11.9</td>
|
||
<td id="16145DAA5F6800305425947DB5B6FA81" box="[1311,1413,1354,1379]" gridcol="6" gridrow="12" pageId="12" pageNumber="93">10.5–12.6</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E9444B5B6FA69" box="[164,1413,1395,1419]" gridrow="13" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E9444B18BFA69" box="[164,440,1395,1419]" gridcol="0" gridrow="13" pageId="12" pageNumber="93">Length of caudal peduncle</th>
|
||
<td id="16145DAA5F68003053369444B259FA69" box="[524,618,1395,1419]" gridcol="1" gridrow="13" pageId="12" pageNumber="93">24.9</td>
|
||
<td id="16145DAA5F68003053A19444B323FA69" box="[667,784,1395,1419]" gridcol="2" gridrow="13" pageId="12" pageNumber="93">24.0–25.1</td>
|
||
<td id="16145DAA5F680030520D9444B3AEFA69" box="[823,925,1395,1419]" gridcol="3" gridrow="13" pageId="12" pageNumber="93">23.2–27.5</td>
|
||
<td id="16145DAA5F68003052D59444B47EFA69" box="[1007,1101,1395,1419]" gridcol="4" gridrow="13" pageId="12" pageNumber="93">23.2</td>
|
||
<td id="16145DAA5F68003055BA9444B4C6FA69" box="[1152,1269,1395,1419]" gridcol="5" gridrow="13" pageId="12" pageNumber="93">23.1–26.4</td>
|
||
<td id="16145DAA5F68003054259444B5B6FA69" box="[1311,1413,1395,1419]" gridcol="6" gridrow="13" pageId="12" pageNumber="93">24.3–27.4</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E94ACB5B6FA51" box="[164,1413,1435,1459]" gridrow="14" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E94ACB18BFA51" box="[164,440,1435,1459]" gridcol="0" gridrow="14" pageId="12" pageNumber="93">Predorsal length</th>
|
||
<td id="16145DAA5F680030533694ACB259FA51" box="[524,618,1435,1459]" gridcol="1" gridrow="14" pageId="12" pageNumber="93">39.4</td>
|
||
<td id="16145DAA5F68003053A194ACB323FA51" box="[667,784,1435,1459]" gridcol="2" gridrow="14" pageId="12" pageNumber="93">39.3–41.4</td>
|
||
<td id="16145DAA5F680030520D94ACB3AEFA51" box="[823,925,1435,1459]" gridcol="3" gridrow="14" pageId="12" pageNumber="93">37.5–40.4</td>
|
||
<td id="16145DAA5F68003052D594ACB47EFA51" box="[1007,1101,1435,1459]" gridcol="4" gridrow="14" pageId="12" pageNumber="93">41.5</td>
|
||
<td id="16145DAA5F68003055BA94ACB4C6FA51" box="[1152,1269,1435,1459]" gridcol="5" gridrow="14" pageId="12" pageNumber="93">38.7–41.1</td>
|
||
<td id="16145DAA5F680030542594ACB5B6FA51" box="[1311,1413,1435,1459]" gridcol="6" gridrow="14" pageId="12" pageNumber="93">37.3–40.4</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E94F4B5B6FA39" box="[164,1413,1475,1499]" gridrow="15" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E94F4B18BFA39" box="[164,440,1475,1499]" gridcol="0" gridrow="15" pageId="12" pageNumber="93">Preanal length</th>
|
||
<td id="16145DAA5F680030533694F4B259FA39" box="[524,618,1475,1499]" gridcol="1" gridrow="15" pageId="12" pageNumber="93">56.2</td>
|
||
<td id="16145DAA5F68003053A194F4B323FA39" box="[667,784,1475,1499]" gridcol="2" gridrow="15" pageId="12" pageNumber="93">55.2–57.6</td>
|
||
<td id="16145DAA5F680030520D94F4B3AEFA39" box="[823,925,1475,1499]" gridcol="3" gridrow="15" pageId="12" pageNumber="93">54.7–59.2</td>
|
||
<td id="16145DAA5F68003052D594F4B47EFA39" box="[1007,1101,1475,1499]" gridcol="4" gridrow="15" pageId="12" pageNumber="93">59.6</td>
|
||
<td id="16145DAA5F68003055BA94F4B4C6FA39" box="[1152,1269,1475,1499]" gridcol="5" gridrow="15" pageId="12" pageNumber="93">55.4–58.3</td>
|
||
<td id="16145DAA5F680030542594F4B5B6FA39" box="[1311,1413,1475,1499]" gridcol="6" gridrow="15" pageId="12" pageNumber="93">54.5–57.3</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E94DCB5B6F9EB" box="[164,1413,1515,1545]" gridrow="16" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E94DCB18BF9EB" box="[164,440,1515,1545]" gridcol="0" gridrow="16" pageId="12" pageNumber="93">Length of D1 base</th>
|
||
<td id="16145DAA5F680030533694DCB259F9EB" box="[524,618,1515,1545]" gridcol="1" gridrow="16" pageId="12" pageNumber="93">18.5</td>
|
||
<td id="16145DAA5F68003053A194DCB323F9EB" box="[667,784,1515,1545]" gridcol="2" gridrow="16" pageId="12" pageNumber="93">16.4–19.2</td>
|
||
<td id="16145DAA5F680030520D94DCB3AEF9EB" box="[823,925,1515,1545]" gridcol="3" gridrow="16" pageId="12" pageNumber="93">16.6–20.9</td>
|
||
<td id="16145DAA5F68003052D594DCB47EF9EB" box="[1007,1101,1515,1545]" gridcol="4" gridrow="16" pageId="12" pageNumber="93">19.3</td>
|
||
<td id="16145DAA5F68003055BA94DCB4C6F9EB" box="[1152,1269,1515,1545]" gridcol="5" gridrow="16" pageId="12" pageNumber="93">17.3–19.7</td>
|
||
<td id="16145DAA5F680030542594DCB5B6F9EB" box="[1311,1413,1515,1545]" gridcol="6" gridrow="16" pageId="12" pageNumber="93">17.9–21.2</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E9723B5B6F9D0" box="[164,1413,1556,1586]" gridrow="17" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E9723B18BF9D0" box="[164,440,1556,1586]" gridcol="0" gridrow="17" pageId="12" pageNumber="93">D 1 length</th>
|
||
<td id="16145DAA5F68003053369723B259F9D0" box="[524,618,1556,1586]" gridcol="1" gridrow="17" pageId="12" pageNumber="93">20.0</td>
|
||
<td id="16145DAA5F68003053A19723B323F9D0" box="[667,784,1556,1586]" gridcol="2" gridrow="17" pageId="12" pageNumber="93">19.8–22.9</td>
|
||
<td id="16145DAA5F680030520D9723B3AEF9D0" box="[823,925,1556,1586]" gridcol="3" gridrow="17" pageId="12" pageNumber="93">19.1–21.1</td>
|
||
<td id="16145DAA5F68003052D59723B47EF9D0" box="[1007,1101,1556,1586]" gridcol="4" gridrow="17" pageId="12" pageNumber="93">22.7</td>
|
||
<td id="16145DAA5F68003055BA9723B4C6F9D0" box="[1152,1269,1556,1586]" gridcol="5" gridrow="17" pageId="12" pageNumber="93">20.4–22.8</td>
|
||
<td id="16145DAA5F68003054259723B5B6F9D0" box="[1311,1413,1556,1586]" gridcol="6" gridrow="17" pageId="12" pageNumber="93">19.2–21.6</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E970BB5B6F99A" box="[164,1413,1596,1656]" gridrow="18" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E970BB18BF99A" box="[164,440,1596,1656]" gridcol="0" gridrow="18" pageId="12" pageNumber="93">Interval between D1 and D2 bases</th>
|
||
<td id="16145DAA5F6800305336970BB259F99A" box="[524,618,1596,1656]" gridcol="1" gridrow="18" pageId="12" pageNumber="93">1.2</td>
|
||
<td id="16145DAA5F68003053A1970BB323F99A" box="[667,784,1596,1656]" gridcol="2" gridrow="18" pageId="12" pageNumber="93">0.3–3.7</td>
|
||
<td id="16145DAA5F680030520D970BB3AEF99A" box="[823,925,1596,1656]" gridcol="3" gridrow="18" pageId="12" pageNumber="93">1.4–3.9</td>
|
||
<td id="16145DAA5F68003052D5970BB47EF99A" box="[1007,1101,1596,1656]" gridcol="4" gridrow="18" pageId="12" pageNumber="93">0.2</td>
|
||
<td id="16145DAA5F68003055BA970BB4C6F99A" box="[1152,1269,1596,1656]" gridcol="5" gridrow="18" pageId="12" pageNumber="93">0.4–2.3</td>
|
||
<td id="16145DAA5F6800305425970BB5B6F99A" box="[1311,1413,1596,1656]" gridcol="6" gridrow="18" pageId="12" pageNumber="93">0.3–4.4</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E97BFB5B6F944" box="[164,1413,1672,1702]" gridrow="19" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E97BFB18BF944" box="[164,440,1672,1702]" gridcol="0" gridrow="19" pageId="12" pageNumber="93">Length of D2 base</th>
|
||
<td id="16145DAA5F680030533697BFB259F944" box="[524,618,1672,1702]" gridcol="1" gridrow="19" pageId="12" pageNumber="93">19.5</td>
|
||
<td id="16145DAA5F68003053A197BFB323F944" box="[667,784,1672,1702]" gridcol="2" gridrow="19" pageId="12" pageNumber="93">16.5–18.5</td>
|
||
<td id="16145DAA5F680030520D97BFB3AEF944" box="[823,925,1672,1702]" gridcol="3" gridrow="19" pageId="12" pageNumber="93">16.5–18.9</td>
|
||
<td id="16145DAA5F68003052D597BFB47EF944" box="[1007,1101,1672,1702]" gridcol="4" gridrow="19" pageId="12" pageNumber="93">18.1</td>
|
||
<td id="16145DAA5F68003055BA97BFB4C6F944" box="[1152,1269,1672,1702]" gridcol="5" gridrow="19" pageId="12" pageNumber="93">17.1–19.4</td>
|
||
<td id="16145DAA5F680030542597BFB5B6F944" box="[1311,1413,1672,1702]" gridcol="6" gridrow="19" pageId="12" pageNumber="93">16.6–19.1</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E9787B5B6F92C" box="[164,1413,1712,1742]" gridrow="20" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E9787B18BF92C" box="[164,440,1712,1742]" gridcol="0" gridrow="20" pageId="12" pageNumber="93">D2 length</th>
|
||
<td id="16145DAA5F68003053369787B259F92C" box="[524,618,1712,1742]" gridcol="1" gridrow="20" pageId="12" pageNumber="93">38.2</td>
|
||
<td id="16145DAA5F68003053A19787B323F92C" box="[667,784,1712,1742]" gridcol="2" gridrow="20" pageId="12" pageNumber="93">34.6–38.7</td>
|
||
<td id="16145DAA5F680030520D9787B3AEF92C" box="[823,925,1712,1742]" gridcol="3" gridrow="20" pageId="12" pageNumber="93">27.7–30.2</td>
|
||
<td id="16145DAA5F68003052D59787B47EF92C" box="[1007,1101,1712,1742]" gridcol="4" gridrow="20" pageId="12" pageNumber="93">42.7</td>
|
||
<td id="16145DAA5F68003055BA9787B4C6F92C" box="[1152,1269,1712,1742]" gridcol="5" gridrow="20" pageId="12" pageNumber="93">33.8–43.6</td>
|
||
<td id="16145DAA5F68003054259787B5B6F92C" box="[1311,1413,1712,1742]" gridcol="6" gridrow="20" pageId="12" pageNumber="93">29.5–35.3</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E97EEB5B6F913" box="[164,1413,1753,1777]" gridrow="21" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E97EEB18BF913" box="[164,440,1753,1777]" gridcol="0" gridrow="21" pageId="12" pageNumber="93">Length of A base</th>
|
||
<td id="16145DAA5F680030533697EEB259F913" box="[524,618,1753,1777]" gridcol="1" gridrow="21" pageId="12" pageNumber="93">16.5</td>
|
||
<td id="16145DAA5F68003053A197EEB323F913" box="[667,784,1753,1777]" gridcol="2" gridrow="21" pageId="12" pageNumber="93">15.4–17.1</td>
|
||
<td id="16145DAA5F680030520D97EEB3AEF913" box="[823,925,1753,1777]" gridcol="3" gridrow="21" pageId="12" pageNumber="93">13.7–17.7</td>
|
||
<td id="16145DAA5F68003052D597EEB47EF913" box="[1007,1101,1753,1777]" gridcol="4" gridrow="21" pageId="12" pageNumber="93">14.9</td>
|
||
<td id="16145DAA5F68003055BA97EEB4C6F913" box="[1152,1269,1753,1777]" gridcol="5" gridrow="21" pageId="12" pageNumber="93">14.0–16.2</td>
|
||
<td id="16145DAA5F680030542597EEB5B6F913" box="[1311,1413,1753,1777]" gridcol="6" gridrow="21" pageId="12" pageNumber="93">14.3–15.6</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E9636B5B6F8FB" box="[164,1413,1793,1817]" gridrow="22" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E9636B18BF8FB" box="[164,440,1793,1817]" gridcol="0" gridrow="22" pageId="12" pageNumber="93">A length</th>
|
||
<td id="16145DAA5F68003053369636B259F8FB" box="[524,618,1793,1817]" gridcol="1" gridrow="22" pageId="12" pageNumber="93">33.7</td>
|
||
<td id="16145DAA5F68003053A19636B323F8FB" box="[667,784,1793,1817]" gridcol="2" gridrow="22" pageId="12" pageNumber="93">31.3–33.1</td>
|
||
<td id="16145DAA5F680030520D9636B3AEF8FB" box="[823,925,1793,1817]" gridcol="3" gridrow="22" pageId="12" pageNumber="93">24.2–29.3</td>
|
||
<td id="16145DAA5F68003052D59636B47EF8FB" box="[1007,1101,1793,1817]" gridcol="4" gridrow="22" pageId="12" pageNumber="93">38.3</td>
|
||
<td id="16145DAA5F68003055BA9636B4C6F8FB" box="[1152,1269,1793,1817]" gridcol="5" gridrow="22" pageId="12" pageNumber="93">30.2–37.0</td>
|
||
<td id="16145DAA5F68003054259636B5B6F8FB" box="[1311,1413,1793,1817]" gridcol="6" gridrow="22" pageId="12" pageNumber="93">26.7–32.7</td>
|
||
</tr>
|
||
<tr id="55C534D65F680030519E961EB5B6F8A3" box="[164,1413,1833,1857]" gridrow="23" pageId="12" pageNumber="93">
|
||
<th id="16145DAA5F680030519E961EB18BF8A3" box="[164,440,1833,1857]" gridcol="0" gridrow="23" pageId="12" pageNumber="93">C length</th>
|
||
<td id="16145DAA5F6800305336961EB259F8A3" box="[524,618,1833,1857]" gridcol="1" gridrow="23" pageId="12" pageNumber="93">29.6</td>
|
||
<td id="16145DAA5F68003053A1961EB323F8A3" box="[667,784,1833,1857]" gridcol="2" gridrow="23" pageId="12" pageNumber="93">28.6–31.7</td>
|
||
<td id="16145DAA5F680030520D961EB3AEF8A3" box="[823,925,1833,1857]" gridcol="3" gridrow="23" pageId="12" pageNumber="93">24.4–28.0</td>
|
||
<td id="16145DAA5F68003052D5961EB47EF8A3" box="[1007,1101,1833,1857]" gridcol="4" gridrow="23" pageId="12" pageNumber="93">34.9</td>
|
||
<td id="16145DAA5F68003055BA961EB4C6F8A3" box="[1152,1269,1833,1857]" gridcol="5" gridrow="23" pageId="12" pageNumber="93">9.3–33.8</td>
|
||
<td id="16145DAA5F6800305425961EB5B6F8A3" box="[1311,1413,1833,1857]" gridcol="6" gridrow="23" pageId="12" pageNumber="93">8.4–28.2</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F68FFC251FD9646B207FF18" blockId="12.[151,1437,1905,2040]" lastBlockId="13.[151,1437,151,1113]" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93">
|
||
<emphasis id="D981EA865F68FFC351FD9646B151F869" bold="true" box="[199,354,1905,1931]" pageId="12" pageNumber="93">Comparison.</emphasis>
|
||
<taxonomicName id="2CF54D175F68FFC350509646B2F7F86E" authority="Gill, 1859" authorityName="Gill" authorityYear="1859" box="[362,708,1905,1932]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="D981EA865F68FFC350509646B27AF869" box="[362,585,1905,1931]" italics="true" pageId="12" pageNumber="93">Rhinogobius similis</emphasis>
|
||
<bibRefCitation id="8F644B655F68FFC3536B9646B2F7F86E" author="Gill, T. N." box="[593,708,1905,1932]" pageId="12" pageNumber="93" pagination="144 - 150" refId="ref9769" refString="Gill, T. N. (1859) Notes on a collection of Japanese fishes, made by Dr. J. Morrow. Proceedings of the Academy of Natural Sciences of Philadelphia, 11, 144 - 150." type="journal article" year="1859">Gill, 1859</bibRefCitation>
|
||
</taxonomicName>
|
||
is the only species previously known in the genus having transverse rows of sensory papillae on cheek (
|
||
<bibRefCitation id="8F644B655F68FFC3534A96A2B37EF852" author="Suzuki, T. & Shibukawa, K. & Senou, H. & Chen, I. - S." box="[624,845,1941,1968]" pageId="12" pageNumber="93" pagination="227 - 238" refId="ref11378" refString="Suzuki, T., Shibukawa, K., Senou, H. & Chen, I. - S. (2016) Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, 63 (2), 227 - 238. https: // doi. org / 10.1007 / s 10228 - 015 - 0494 - 3" type="journal article" year="2016">
|
||
Suzuki
|
||
<emphasis id="D981EA865F68FFC353F296A1B331F84D" box="[712,770,1941,1967]" italics="true" pageId="12" pageNumber="93">et al.</emphasis>
|
||
, 2016
|
||
</bibRefCitation>
|
||
), and our two new species share this unique feature with
|
||
<taxonomicName id="2CF54D175F68FFC351EB968DB109F831" authorityName="Gill" authorityYear="1859" box="[209,314,1977,2003]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="D981EA865F68FFC351EB968DB109F831" box="[209,314,1977,2003]" italics="true" pageId="12" pageNumber="93">R. similis</emphasis>
|
||
</taxonomicName>
|
||
. However, the two species in
|
||
<collectingRegion id="2931F8765F68FFC353B5968EB2DDF831" box="[655,750,1977,2003]" country="Philippines" name="Palawan" pageId="12" pageNumber="93">Palawan</collectingRegion>
|
||
are distinguished from
|
||
<taxonomicName id="2CF54D175F68FFC352C6968DB456F831" authorityName="Gill" authorityYear="1859" box="[1020,1125,1977,2003]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="93" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="D981EA865F68FFC352C6968DB456F831" box="[1020,1125,1977,2003]" italics="true" pageId="12" pageNumber="93">R. similis</emphasis>
|
||
</taxonomicName>
|
||
in having fewer rays in the pectoral fin (14–17 vs 18–19), predorsal squamation composed of cycloid scales (vs ctenoid scales), anterior-most dorsal scale located around a vertical line through posterior margin of the preopercle or more posteriorly (vs dorsal squamation reaching anteriorly near the posterior margin of the eye), and proximal part of the caudal fin covered by cycloid scales (vs by ctenoid scales).
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F69FFC251FD9034B49BFD67" blockId="13.[151,1437,151,1113]" pageId="13" pageNumber="94">
|
||
<taxonomicName id="2CF54D175F69FFC251FD9034B189FEFF" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[199,442,259,285]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F69FFC251FD9034B189FEFF" box="[199,442,259,285]" italics="true" pageId="13" pageNumber="94">Rhinogobius estrellae</emphasis>
|
||
</taxonomicName>
|
||
shares many characters with
|
||
<emphasis id="D981EA865F69FFC252399034B3B8FEFF" box="[771,907,259,285]" italics="true" pageId="13" pageNumber="94">
|
||
<taxonomicName id="2CF54D175F69FFC252399034B3B4FEFF" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[771,903,259,285]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">R. tandikan</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but the former differs from the latter by having fewer pectoral-fin rays (usually 15 vs 16–17;
|
||
<tableCitation id="A677032F5F69FFC253A89010B2D5FEA3" box="[658,742,295,321]" captionStart="TABLE 2" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 2. Pectoral-fin ray counts of Rhinogobius estrellae and Rhinogobius tandikan." httpUri="http://table.plazi.org/id/BF8A661C5F61FFCA51AD91A0B406FF4D" pageId="13" pageNumber="94" tableUuid="BF8A661C5F61FFCA51AD91A0B406FF4D">Table 2</tableCitation>
|
||
), fewer scale counts (longitudinal scales 27–31 vs 29–34, predorsal scales 3–11 vs 5–13;
|
||
<tableCitation id="A677032F5F69FFC250E8907CB202FE87" box="[466,561,331,357]" captionStart="TABLE 3" captionStartId="5.[151,239,675,699]" captionTargetPageId="5" captionText="TABLE 3. Longitudinal scale counts of Rhinogobius estrellae and Rhinogobius tandikan." pageId="13" pageNumber="94">Tables 3</tableCitation>
|
||
,
|
||
<tableCitation id="A677032F5F69FFC25305907DB27DFE87" box="[575,590,330,357]" captionStart="TABLE 5" captionStartId="5.[151,239,1063,1087]" captionTargetPageId="5" captionText="TABLE 5. Predorsal scale counts of Rhinogobius estrellae and Rhinogobius tandikan." pageId="13" pageNumber="94">5</tableCitation>
|
||
), and colourations of the body. In preservative,
|
||
<taxonomicName id="2CF54D175F69FFC25553907CB4D4FE87" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[1129,1255,331,357]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F69FFC25553907CB4D4FE87" box="[1129,1255,331,357]" italics="true" pageId="13" pageNumber="94">R. estrellae</emphasis>
|
||
</taxonomicName>
|
||
has dark brown mesh-like markings on the lateral and dorsal sides of the body (vs rather uniformly brown in
|
||
<taxonomicName id="2CF54D175F69FFC255AD9058B524FE6B" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[1175,1303,367,393]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F69FFC255AD9058B524FE6B" box="[1175,1303,367,393]" italics="true" pageId="13" pageNumber="94">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
), three dark brown stripes on the snout and cheek (vs often obscure), and a dark brown vertical band on the proximal part of the upper and middle pectoral-fin rays except lower 2–6 rays (vs this band restricted to upper half of the fin). In life,
|
||
<taxonomicName id="2CF54D175F69FFC254BE9080B0C5FE17" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F69FFC254BE9080B0C5FE17" italics="true" pageId="13" pageNumber="94">R. estrellae</emphasis>
|
||
</taxonomicName>
|
||
has a more reddish body than the yellowish or olive green
|
||
<taxonomicName id="2CF54D175F69FFC252AF90ECB425FE17" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[917,1046,475,501]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F69FFC252AF90ECB425FE17" box="[917,1046,475,501]" italics="true" pageId="13" pageNumber="94">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
, and the former has a bright white vertical band restricted to the upper and middle parts of the pectoral-fin base while this band reaches nearly the bottom of the fin in
|
||
<taxonomicName id="2CF54D175F69FFC250699314B1E7FDDF" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[339,468,547,573]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F69FFC250699314B1E7FDDF" box="[339,468,547,573]" italics="true" pageId="13" pageNumber="94">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
. Arrangements of the cephalic sensory pore system are basically the same between the two species, but
|
||
<taxonomicName id="2CF54D175F69FFC2506B9370B1FCFD83" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[337,463,583,609]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F69FFC2506B9370B1FCFD83" box="[337,463,583,609]" italics="true" pageId="13" pageNumber="94">R. estrellae</emphasis>
|
||
</taxonomicName>
|
||
more often lacks pore E on one or both sides than in
|
||
<taxonomicName id="2CF54D175F69FFC2551E9370B496FD83" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[1060,1189,583,609]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F69FFC2551E9370B496FD83" box="[1060,1189,583,609]" italics="true" pageId="13" pageNumber="94">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
(12/24 vs 5/29) while
|
||
<taxonomicName id="2CF54D175F69FFC251AD935CB12BFD67" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[151,280,619,645]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F69FFC251AD935CB12BFD67" box="[151,280,619,645]" italics="true" pageId="13" pageNumber="94">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
more often lacks pore C on one or both sides than in
|
||
<taxonomicName id="2CF54D175F69FFC2524A935CB3DDFD67" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[880,1006,619,645]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F69FFC2524A935CB3DDFD67" box="[880,1006,619,645]" italics="true" pageId="13" pageNumber="94">R. estrellae</emphasis>
|
||
</taxonomicName>
|
||
(16/29 vs 7/24).
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F69FFC251FD93B8B407FC44" blockId="13.[151,1437,151,1113]" pageId="13" pageNumber="94">
|
||
In addition to the transverse rows of sensory papillae on cheek (present vs absent), there are many differences between the two new species from
|
||
<collectingRegion id="2931F8765F69FFC253279384B24FFD2F" box="[541,636,691,717]" country="Philippines" name="Palawan" pageId="13" pageNumber="94">Palawan</collectingRegion>
|
||
and three other Philippine species,
|
||
<taxonomicName id="2CF54D175F69FFC255329384B514FD2F" authority="Seale, 1910" authorityName="Seale" authorityYear="1910" box="[1032,1319,691,717]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="D981EA865F69FFC255329384B4AFFD2E" box="[1032,1180,691,716]" italics="true" pageId="13" pageNumber="94">R. carpenteri</emphasis>
|
||
<bibRefCitation id="8F644B655F69FFC255989384B514FD2F" author="Seale, A." box="[1186,1319,691,717]" pageId="13" pageNumber="94" pagination="491 - 543" refId="ref11257" refString="Seale, A. (1910) New species of Philippine fishes. The Philippine Journal of Science, Section A, 4, 491 - 543, pls. 1 - 13." type="journal article" year="1910">Seale, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2CF54D175F69FFC2540B9384B148FD13" authority="(Herre, 1927)" baseAuthorityName="Herre" baseAuthorityYear="1927" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="bucculentus">
|
||
<emphasis id="D981EA865F69FFC2540B9384B0E9FD13" italics="true" pageId="13" pageNumber="94">R. bucculentus</emphasis>
|
||
(
|
||
<bibRefCitation id="8F644B655F69FFC251D293E0B140FD13" author="Herre, A. W. C. T." box="[232,371,727,753]" pageId="13" pageNumber="94" pagination="1 - 352" refId="ref9810" refString="Herre, A. W. C. T. (1927) Gobies of the Philippines and the China Sea. The Philippine Bureau of Science Monographic Publications of Fishes, 23, 1 - 352, pls. 1 - 30." type="journal article" year="1927">Herre, 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2CF54D175F69FFC2508F93E0B2C1FD13" authority="(Herre, 1927)" baseAuthorityName="Herre" baseAuthorityYear="1927" box="[437,754,727,753]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="philippinus">
|
||
<emphasis id="D981EA865F69FFC2508F93E0B262FD13" box="[437,593,727,753]" italics="true" pageId="13" pageNumber="94">R. philippinus</emphasis>
|
||
(
|
||
<bibRefCitation id="8F644B655F69FFC2536593E0B2D9FD13" author="Herre, A. W. C. T." box="[607,746,727,753]" pageId="13" pageNumber="94" pagination="1 - 352" refId="ref9810" refString="Herre, A. W. C. T. (1927) Gobies of the Philippines and the China Sea. The Philippine Bureau of Science Monographic Publications of Fishes, 23, 1 - 352, pls. 1 - 30." type="journal article" year="1927">Herre, 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, such as numbers of the vertebrae (26 vs 27–29), first-dorsalfin pterygiophores (usually 6 vs usually 7), first-dorsal-fin spines (usually 6 vs usually
|
||
<quantity id="2C0D9B715F69FFC2555993CCB4BDFCF7" box="[1123,1166,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="13" pageNumber="94" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="2CF54D175F69FFC255AF93CCB509FCF7" baseAuthorityName="Herre" baseAuthorityYear="1927" box="[1173,1338,763,789]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="bucculentus">
|
||
<emphasis id="D981EA865F69FFC255AF93CCB509FCF7" box="[1173,1338,763,789]" italics="true" pageId="13" pageNumber="94">R. bucculentus</emphasis>
|
||
</taxonomicName>
|
||
and 6 or
|
||
<quantity id="2C0D9B715F69FFC251AD9228B0F0FCDB" box="[151,195,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="13" pageNumber="94" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="2CF54D175F69FFC251F19228B153FCDA" authorityName="Seale" authorityYear="1910" box="[203,352,799,824]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="D981EA865F69FFC251F19228B153FCDA" box="[203,352,799,824]" italics="true" pageId="13" pageNumber="94">R. carpenteri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2CF54D175F69FFC250A29228B205FCDB" baseAuthorityName="Herre" baseAuthorityYear="1927" box="[408,566,799,825]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="philippinus">
|
||
<emphasis id="D981EA865F69FFC250A29228B205FCDB" box="[408,566,799,825]" italics="true" pageId="13" pageNumber="94">R. philippinus</emphasis>
|
||
</taxonomicName>
|
||
), pectoral-fin rays (14–17 vs 17–20), having fewer scales in the longitudinal scales (27–34 vs 35–43), in the transverse scales (8–10 vs 12–18), and in the transverse scales in caudal peduncle (7 vs 9–11), and presence of scales on belly behind the pelvic fins (vs absent in
|
||
<taxonomicName id="2CF54D175F69FFC252D79250B4B2FC62" authorityName="Seale" authorityYear="1910" box="[1005,1153,871,896]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="D981EA865F69FFC252D79250B4B2FC62" box="[1005,1153,871,896]" italics="true" pageId="13" pageNumber="94">R. carpenteri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2CF54D175F69FFC2558C9250B568FC63" baseAuthorityName="Herre" baseAuthorityYear="1927" box="[1206,1371,871,897]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="bucculentus">
|
||
<emphasis id="D981EA865F69FFC2558C9250B568FC63" box="[1206,1371,871,897]" italics="true" pageId="13" pageNumber="94">R. bucculentus</emphasis>
|
||
</taxonomicName>
|
||
). The latter three species are distributed in Luzon, northern
|
||
<collectingCountry id="93E276045F69FFC253D592BCB35EFC47" box="[751,877,907,933]" name="Philippines" pageId="13" pageNumber="94">Philippines</collectingCountry>
|
||
(see Discussion).
|
||
</paragraph>
|
||
<paragraph id="EB4A36945F69FFC251FD9298B2FBFBBB" blockId="13.[151,1437,151,1113]" pageId="13" pageNumber="94">
|
||
<emphasis id="D981EA865F69FFC251FD9298B27CFC2A" bold="true" box="[199,591,942,969]" pageId="13" pageNumber="94">Molecular phylogenetic analysis.</emphasis>
|
||
In the ML phylogenetic tree using the aligned 16,409 bp of mitochondrial genomes (
|
||
<figureCitation id="73CE2A115F69FFC2503392E4B17EFC0F" box="[265,333,979,1005]" captionStart="FIGURE 9" captionStartId="14.[151,250,1593,1617]" captionTargetBox="[158,1426,606,1562]" captionTargetId="figure-241@14.[151,1436,597,1570]" captionTargetPageId="14" captionText="FIGURE 9. Maximum likelihood phylogenetic tree with 100 bootstraps using the aligned 16,409 bp of mitochondrial genomes in Rhinogobius including the two new species, R. estrellae and R. tandikan, with Tridentiger kuroiwae as an outgroup taxon. Material sequenced in the present study are shown with the catalogue numbers of the vouchers (beginning with NSMT-P, URM- P, or WPU-PPC-P) and sequences from the International Nucleotide Sequence Database are shown with the accession numbers (R. cliffordpopei, R. duospilus, and R. leavelli). The scale bar indicates 0.02 substitutions per site. Pictures shown along with the species names are of the specimens with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5702122" httpUri="https://zenodo.org/record/5702122/files/figure.png" pageId="13" pageNumber="94">Fig. 9</figureCitation>
|
||
), the species in
|
||
<taxonomicName id="2CF54D175F69FFC250C792E4B2B8FC0F" box="[509,651,979,1005]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="genus">
|
||
<emphasis id="D981EA865F69FFC250C792E4B2B8FC0F" box="[509,651,979,1005]" italics="true" pageId="13" pageNumber="94">Rhinogobius</emphasis>
|
||
</taxonomicName>
|
||
were divided into two robust clades. One clade included
|
||
<taxonomicName id="2CF54D175F69FFC2543C92E4B55EFC0F" authorityName="Gill" authorityYear="1859" box="[1286,1389,979,1005]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="D981EA865F69FFC2543C92E4B55EFC0F" box="[1286,1389,979,1005]" italics="true" pageId="13" pageNumber="94">R. similis</emphasis>
|
||
</taxonomicName>
|
||
and the two new species from
|
||
<collectingRegion id="2931F8765F69FFC2508692C0B22DFBF3" box="[444,542,1015,1041]" country="Philippines" name="Palawan" pageId="13" pageNumber="94">Palawan</collectingRegion>
|
||
, while all other species belonged to another clade. Six specimens of
|
||
<taxonomicName id="2CF54D175F69FFC2542492C0B5AFFBF3" authorityName="Maeda, Kunishima & Palla" authorityYear="2021" box="[1310,1436,1015,1041]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="estrellae">
|
||
<emphasis id="D981EA865F69FFC2542492C0B5AFFBF3" box="[1310,1436,1015,1041]" italics="true" pageId="13" pageNumber="94">R. estrellae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<specimenCount id="FDF3FD1D5F69FFC251FD952CB141FBD7" box="[199,370,1051,1077]" pageId="13" pageNumber="94" type="generic">four specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="2CF54D175F69FFC250AD952CB22BFBD7" authorityName="Maeda, Kobayashi & Palla" authorityYear="2021" box="[407,536,1051,1077]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="tandikan">
|
||
<emphasis id="D981EA865F69FFC250AD952CB22BFBD7" box="[407,536,1051,1077]" italics="true" pageId="13" pageNumber="94">R. tandikan</emphasis>
|
||
</taxonomicName>
|
||
formed each well-supported monophyletic clade and were sister species to each other.
|
||
<taxonomicName id="2CF54D175F69FFC251E79508B188FBBB" authorityName="Gill" authorityYear="1859" box="[221,443,1087,1113]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="94" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis id="D981EA865F69FFC251E79508B188FBBB" box="[221,443,1087,1113]" italics="true" pageId="13" pageNumber="94">Rhinogobius similis</emphasis>
|
||
</taxonomicName>
|
||
was a sister to this pair.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |