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<document id="1B28B15E0102C1B6F16DBC7E6B743A1D" ID-CLB-Dataset="7470" ID-DOI="10.11646/zootaxa.4948.3.4" ID-GBIF-Dataset="6bfd846c-0304-4330-8941-95c406630230" ID-ISSN="1175-5326" ID-Zenodo-Dep="4629100" ID-ZooBank="8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616485698777" checkinUser="plazi" docAuthor="Xu, Deliang, Webb, Michael D., Shang, Suqin &amp; Zhang, Yalin" docDate="2021" docId="597A878EFFB9FF98FF66F1E285B0F26A" docLanguage="en" docName="zootaxa.4948.3.4.pdf" docOrigin="Zootaxa 4948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hecaloidella caudospina Zhang &amp; Webb 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="389" masterDocId="A543FFF6FFBCFF90FFF1F728870FF30F" masterDocTitle="Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species" masterLastPageNumber="403" masterPageNumber="381" pageNumber="386" updateTime="1698934487255" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CD716D7378FF5638D1FB8BFB9B1B8D26">Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species</mods:title>
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<mods:namePart id="7871509287BA6FCE1D06A41A299386A7">Xu, Deliang</mods:namePart>
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<mods:affiliation id="030E42326C122D558BFD7D5EC57B57EE">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570</mods:affiliation>
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<mods:namePart id="E96BA4E67AA25BAF7303BA9E2384096F">Webb, Michael D.</mods:namePart>
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<mods:affiliation id="81F848E98B3130806127DAD5CF62AD0B">Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
<mods:nameIdentifier id="7565AD7EF98F201153B8200E23FB079E" type="email">mdwebb04@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1AB401FC933B827DC9F425203CA4D5B4">Shang, Suqin</mods:namePart>
<mods:nameIdentifier id="9DE7F822C6577251502D7147D14B74C5" type="ORCID">0000-0001-9376-0735</mods:nameIdentifier>
<mods:affiliation id="F71789AC7155E09651B392753007286B">Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China &amp; shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735</mods:affiliation>
<mods:nameIdentifier id="FE3643279BD3AE0308B2B547E5660E9D" type="email">shangsq@gsau.edu.cn</mods:nameIdentifier>
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<mods:namePart id="9D85EE3972135024675117648C26E431">Zhang, Yalin</mods:namePart>
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<mods:affiliation id="30FA490FA0F465FA30CF5E1F4095EFC7">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181</mods:affiliation>
<mods:nameIdentifier id="AEBAB53027E49A69BDC47D24BD464352" type="email">yalinzh@nwsuaf.edu.cn</mods:nameIdentifier>
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<mods:date id="1D134E2FE8A64634BD6E1B08D79ED84F">2021</mods:date>
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<treatment id="597A878EFFB9FF98FF66F1E285B0F26A" ID-DOI="http://doi.org/10.5281/zenodo.4629120" ID-GBIF-Taxon="180877863" ID-Zenodo-Dep="4629120" LSID="urn:lsid:plazi:treatment:597A878EFFB9FF98FF66F1E285B0F26A" httpUri="http://treatment.plazi.org/id/597A878EFFB9FF98FF66F1E285B0F26A" lastPageId="8" lastPageNumber="389" pageId="5" pageNumber="386">
<subSubSection id="99C96513FFB9FF95FF66F1E2857AF5EC" box="[151,629,1737,1764]" pageId="5" pageNumber="386" type="nomenclature">
<paragraph id="D16C3698FFB9FF95FF66F1E2857AF5EC" blockId="5.[151,629,1737,1799]" box="[151,629,1737,1764]" pageId="5" pageNumber="386">
<heading id="8A2481F4FFB9FF95FF66F1E2857AF5EC" bold="true" box="[151,629,1737,1764]" fontSize="11" level="1" pageId="5" pageNumber="386" reason="1">
<emphasis id="E3A7EA8AFFB9FF95FF66F1E2857AF5EC" bold="true" box="[151,629,1737,1764]" pageId="5" pageNumber="386">
<taxonomicName id="16D34D1BFFB9FF95FF66F1E2851FF5EC" ID-CoL="8KMST" authority="Zhang &amp; Webb" authorityName="Zhang &amp; Webb" authorityYear="2021" box="[151,528,1737,1764]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="caudospina" status="sp. nov.">
<emphasis id="E3A7EA8AFFB9FF95FF66F1E287A1F5EB" bold="true" box="[151,174,1738,1764]" italics="true" pageId="5" pageNumber="386">H</emphasis>
.
<emphasis id="E3A7EA8AFFB9FF95FF4CF1E18647F5EB" bold="true" box="[189,328,1737,1764]" italics="true" pageId="5" pageNumber="386">caudospina</emphasis>
Zhang &amp; Webb
</taxonomicName>
<taxonomicNameLabel id="F89457F1FFB9FF95FDE6F1E1857AF5EC" box="[535,629,1737,1763]" pageId="5" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB9FF95FF66F1C586F1F4D0" pageId="5" pageNumber="386" type="description">
<paragraph id="D16C3698FFB9FF95FF66F1C5864AF408" blockId="5.[151,629,1737,1799]" box="[151,325,1773,1799]" pageId="5" pageNumber="386">
<figureCitation id="49E82A1DFFB9FF95FF66F1C587F7F408" box="[151,248,1773,1799]" captionStart="FIGURE 1" captionStartId="6.[151,250,1870,1894]" captionTargetBox="[223,1380,183,1845]" captionTargetId="figure-19@6.[205,1382,181,1846]" captionTargetPageId="6" captionText="FIGURE 1. Habitus images of Hecaloidella species; A, H. bimaculata (holotype); B, H. caudospina (holotype); C, H. nitida (holotype); D, H. nudapenis (holotype); E–G, H. pacifica (paratypes and immature); H, H. borneoensis (holotype); I, H. brevipenis (holotype); J, H. brevispina (holotype); K, H. longipenis (holotype); L, H. quadrispina (holotype); M, H. spinosa (holotype); N, H. brevis (holotype); O, H. brunnifacialis (topotype); P, H. knighti (holotype); Q, H. quadripicta (paratype); R, H. sandakanensis (holotype); S, H. singaporensis (paratype); T, H. brunifasciata (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4629102" httpUri="https://zenodo.org/record/4629102/files/figure.png" pageId="5" pageNumber="386">Figs. 1B</figureCitation>
,
<figureCitation id="49E82A1DFFB9FF95FEF5F1C5864AF408" box="[260,325,1773,1799]" captionStart="FIGURE 4" captionStartId="12.[151,250,1895,1919]" captionTargetBox="[198,1374,189,1861]" captionTargetId="figure-19@12.[196,1391,181,1871]" captionTargetPageId="12" captionText="FIGURE 4. Male genitalia of H. nitida species group; A–E, H. caudospina; A–B, aedeagus, lateral and caudal view; C–D, connective, lateral and dorsal view; E, style; F–J, H. nudapenis; F–G, aedeagus, lateral and caudal view; H–I, connective, lateral and dorsal view; J, style; K–O, H. pacifica; K–L, aedeagus, lateral and caudal view; M–N, connective, lateral and dorsal view; O, style." figureDoi="http://doi.org/10.5281/zenodo.4629108" httpUri="https://zenodo.org/record/4629108/files/figure.png" pageId="5" pageNumber="386">4A–E</figureCitation>
</paragraph>
<paragraph id="D16C3698FFB9FF95FF66F01D8477F45F" blockId="5.[151,1436,1845,2016]" box="[151,888,1845,1872]" pageId="5" pageNumber="386">
<emphasis id="E3A7EA8AFFB9FF95FF66F01D8626F440" bold="true" box="[151,297,1845,1871]" pageId="5" pageNumber="386">Description.</emphasis>
Body length (including tegmina), male:
<quantity id="162B9B7DFFB9FF95FD00F01D847BF45F" box="[753,884,1845,1872]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" pageId="5" pageNumber="386" unit="mm" value="5.3" valueMax="5.8" valueMin="4.8">4.8–5.8mm</quantity>
.
</paragraph>
<paragraph id="D16C3698FFB9FF95FF36F0718434F498" blockId="5.[151,1436,1845,2016]" pageId="5" pageNumber="386">Stramineous to sordid yellow. Vertex anteriorly orange with a small brown spot each side of midline. Face with variable brown marking dorsally. Forewing veins yellowish.</paragraph>
<paragraph id="D16C3698FFB9FF95FF36F08986F1F4D0" blockId="5.[151,1436,1845,2016]" pageId="5" pageNumber="386">
Male genitalia similar to
<taxonomicName id="16D34D1BFFB9FF95FE11F08A857AF4B4" box="[480,629,1953,1980]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="nudapenis">
<emphasis id="E3A7EA8AFFB9FF95FE11F08A86FBF4B4" box="[480,500,1954,1979]" italics="true" pageId="5" pageNumber="386">H</emphasis>
.
<emphasis id="E3A7EA8AFFB9FF95FDF3F089857AF4B4" box="[514,629,1953,1979]" italics="true" pageId="5" pageNumber="386">nudapenis</emphasis>
</taxonomicName>
but caudal extension of connective with a long ventral process and ventral process of aedeagus shorter.
</paragraph>
</subSubSection>
<caption id="85AC6610FFBAFF96FF66F066832EF4F9" ID-DOI="http://doi.org/10.5281/zenodo.4629102" ID-Zenodo-Dep="4629102" httpUri="https://zenodo.org/record/4629102/files/figure.png" pageId="6" pageNumber="387" startId="6.[151,250,1870,1894]" targetBox="[223,1380,183,1845]" targetPageId="6">
<paragraph id="D16C3698FFBAFF96FF66F066832EF4F9" blockId="6.[151,1436,1870,2038]" pageId="6" pageNumber="387">
<emphasis id="E3A7EA8AFFBAFF96FF66F066861BF469" bold="true" box="[151,276,1870,1894]" pageId="6" pageNumber="387">FIGURE 1.</emphasis>
Habitus images of
<taxonomicName id="16D34D1BFFBAFF96FE2CF066856FF469" authorityName="Osborn" authorityYear="1934" box="[477,608,1870,1894]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="E3A7EA8AFFBAFF96FE2CF066856FF469" box="[477,608,1870,1894]" italics="true" pageId="6" pageNumber="387">Hecaloidella</emphasis>
</taxonomicName>
species; A,
<taxonomicName id="16D34D1BFFBAFF96FD2FF067847FF469" box="[734,880,1870,1894]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="E3A7EA8AFFBAFF96FD2FF06785FFF469" box="[734,752,1871,1894]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FD0FF066847FF469" box="[766,880,1870,1894]" italics="true" pageId="6" pageNumber="387">bimaculata</emphasis>
</taxonomicName>
(holotype); B,
<taxonomicName id="16D34D1BFFBAFF96FBFFF06783ACF469" box="[1038,1187,1870,1894]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="caudospina">
<emphasis id="E3A7EA8AFFBAFF96FBFFF067832FF469" box="[1038,1056,1871,1894]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FBDFF06683ACF469" box="[1070,1187,1870,1894]" italics="true" pageId="6" pageNumber="387">caudospina</emphasis>
</taxonomicName>
(holotype); C,
<taxonomicName id="16D34D1BFFBAFF96FAB0F0678293F469" authorityName="Osborn" authorityYear="1934" box="[1345,1436,1870,1894]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="E3A7EA8AFFBAFF96FAB0F067825CF469" box="[1345,1363,1871,1894]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FA90F0668293F469" box="[1377,1436,1870,1894]" italics="true" pageId="6" pageNumber="387">nitida</emphasis>
</taxonomicName>
(holotype); D,
<taxonomicName id="16D34D1BFFBAFF96FEDEF05B86B7F485" box="[303,440,1906,1930]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="nudapenis">
<emphasis id="E3A7EA8AFFBAFF96FEDEF05B864EF485" box="[303,321,1907,1930]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FEA1F05A86B7F485" box="[336,440,1906,1930]" italics="true" pageId="6" pageNumber="387">nudapenis</emphasis>
</taxonomicName>
(holotype); E–G,
<taxonomicName id="16D34D1BFFBAFF96FD85F05B85E9F485" box="[628,742,1906,1930]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="E3A7EA8AFFBAFF96FD85F05B8589F485" box="[628,646,1907,1930]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FD65F05A85E9F485" box="[660,742,1906,1930]" italics="true" pageId="6" pageNumber="387">pacifica</emphasis>
</taxonomicName>
(paratypes and immature); H,
<taxonomicName id="16D34D1BFFBAFF96FBD3F05B83B3F485" box="[1058,1212,1906,1930]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFBAFF96FBD3F05B833BF485" box="[1058,1076,1907,1930]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FBB3F05A83B3F485" box="[1090,1212,1906,1930]" italics="true" pageId="6" pageNumber="387">borneoensis</emphasis>
</taxonomicName>
(holotype); I,
<taxonomicName id="16D34D1BFFBAFF96FAA2F05B87D0F4A1" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="brevipenis">
<emphasis id="E3A7EA8AFFBAFF96FAA2F05B826AF485" box="[1363,1381,1907,1930]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FA82F05A87D0F4A1" italics="true" pageId="6" pageNumber="387">brevipenis</emphasis>
</taxonomicName>
(holotype); J,
<taxonomicName id="16D34D1BFFBAFF96FE84F0BF86F1F4A1" box="[373,510,1942,1966]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="E3A7EA8AFFBAFF96FE84F0BF8688F4A1" box="[373,391,1943,1966]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FE65F0BE86F1F4A1" box="[404,510,1942,1966]" italics="true" pageId="6" pageNumber="387">brevispina</emphasis>
</taxonomicName>
(holotype); K,
<taxonomicName id="16D34D1BFFBAFF96FD6CF0BF8429F4A1" box="[669,806,1942,1966]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis id="E3A7EA8AFFBAFF96FD6CF0BF85A0F4A1" box="[669,687,1943,1966]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FD4DF0BE8429F4A1" box="[700,806,1942,1966]" italics="true" pageId="6" pageNumber="387">longipenis</emphasis>
</taxonomicName>
(holotype); L,
<taxonomicName id="16D34D1BFFBAFF96FC30F0BF8354F4A1" box="[961,1115,1942,1966]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="quadrispina">
<emphasis id="E3A7EA8AFFBAFF96FC30F0BF84DCF4A1" box="[961,979,1943,1966]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FC10F0BE8354F4A1" box="[993,1115,1942,1966]" italics="true" pageId="6" pageNumber="387">quadrispina</emphasis>
</taxonomicName>
(holotype); M,
<taxonomicName id="16D34D1BFFBAFF96FB0FF0BF8264F4A1" box="[1278,1387,1943,1966]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E3A7EA8AFFBAFF96FB0FF0BF821FF4A1" box="[1278,1296,1943,1966]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FAEFF0BF8264F4A1" box="[1310,1387,1943,1966]" italics="true" pageId="6" pageNumber="387">spinosa</emphasis>
</taxonomicName>
(holotype); N,
<taxonomicName id="16D34D1BFFBAFF96FEFCF0938665F4DD" box="[269,362,1978,2002]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="E3A7EA8AFFBAFF96FEFCF0938610F4DD" box="[269,287,1979,2002]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FEDCF0928665F4DD" box="[301,362,1978,2002]" italics="true" pageId="6" pageNumber="387">brevis</emphasis>
</taxonomicName>
(holotype); O,
<taxonomicName id="16D34D1BFFBAFF96FDF9F09385B9F4DD" box="[520,694,1978,2002]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="brunnifacialis">
<emphasis id="E3A7EA8AFFBAFF96FDF9F0938515F4DD" box="[520,538,1979,2002]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FDD9F09285B9F4DD" box="[552,694,1978,2002]" italics="true" pageId="6" pageNumber="387">brunnifacialis</emphasis>
</taxonomicName>
(topotype); P,
<taxonomicName id="16D34D1BFFBAFF96FCBCF09384BCF4DD" box="[845,947,1978,2002]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="knighti">
<emphasis id="E3A7EA8AFFBAFF96FCBCF0938450F4DD" box="[845,863,1979,2002]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FC9CF09284BCF4DD" box="[877,947,1978,2002]" italics="true" pageId="6" pageNumber="387">knighti</emphasis>
</taxonomicName>
(holotype); Q,
<taxonomicName id="16D34D1BFFBAFF96FBA3F09383E7F4DD" box="[1106,1256,1978,2002]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="quadripicta">
<emphasis id="E3A7EA8AFFBAFF96FBA3F093836BF4DD" box="[1106,1124,1979,2002]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FB83F09283E7F4DD" box="[1138,1256,1978,2002]" italics="true" pageId="6" pageNumber="387">quadripicta</emphasis>
</taxonomicName>
(paratype); R,
<taxonomicName id="16D34D1BFFBAFF96FA75F0938624F4F9" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="sandakanensis">
<emphasis id="E3A7EA8AFFBAFF96FA75F0938299F4DD" box="[1412,1430,1979,2002]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FF66F0F68624F4F9" box="[151,299,2014,2038]" italics="true" pageId="6" pageNumber="387">sandakanensis</emphasis>
</taxonomicName>
(holotype); S,
<taxonomicName id="16D34D1BFFBAFF96FE32F0F78561F4F9" box="[451,622,2015,2038]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="E3A7EA8AFFBAFF96FE32F0F786DAF4F9" box="[451,469,2015,2038]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FE13F0F78561F4F9" box="[482,622,2015,2038]" italics="true" pageId="6" pageNumber="387">singaporensis</emphasis>
</taxonomicName>
(paratype); T,
<taxonomicName id="16D34D1BFFBAFF96FCF5F0F784A5F4F9" box="[772,938,2014,2038]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="brunifasciata">
<emphasis id="E3A7EA8AFFBAFF96FCF5F0F78419F4F9" box="[772,790,2015,2038]" italics="true" pageId="6" pageNumber="387">H</emphasis>
.
<emphasis id="E3A7EA8AFFBAFF96FCD2F0F684A5F4F9" box="[803,938,2014,2038]" italics="true" pageId="6" pageNumber="387">brunifasciata</emphasis>
</taxonomicName>
(holotype).
</paragraph>
</caption>
<caption id="85AC6610FFBBFF97FF66F0BF8484F4F7" ID-DOI="http://doi.org/10.5281/zenodo.4629104" ID-Zenodo-Dep="4629104" httpUri="https://zenodo.org/record/4629104/files/figure.png" pageId="7" pageNumber="388" startId="7.[151,250,1943,1967]" targetBox="[191,1403,188,1905]" targetPageId="7">
<paragraph id="D16C3698FFBBFF97FF66F0BF8484F4F7" blockId="7.[151,1436,1943,2040]" pageId="7" pageNumber="388">
<emphasis id="E3A7EA8AFFBBFF97FF66F0BF861BF4A0" bold="true" box="[151,276,1943,1967]" pageId="7" pageNumber="388">FIGURE 2.</emphasis>
<taxonomicName id="16D34D1BFFBBFF97FEEAF0B086A2F4BF" box="[283,429,1944,1968]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="E3A7EA8AFFBBFF97FEEAF0B086A2F4BF" box="[283,429,1944,1968]" italics="true" pageId="7" pageNumber="388">H. bimaculata</emphasis>
</taxonomicName>
; A–B, habitus, dorsal and lateral view; C, face, D, head and thorax, anterior view; E, male genital capsule, lateral view; F, style; G, valve, subgenital plates, styles and connective, dorsal view; H, aedeagus, lateral view; I aedeagus, caudal view; J, anal tube, lateral view; K, connective, lateral view.
</paragraph>
</caption>
<subSubSection id="99C96513FFB4FF98FF36F7BE84B4F212" pageId="8" pageNumber="389" type="materials_examined">
<paragraph id="D16C3698FFB4FF98FF36F7BE839DF3DA" blockId="8.[151,1436,150,357]" pageId="8" pageNumber="389">
<emphasis id="E3A7EA8AFFB4FF98FF36F7BE86BEF3BF" bold="true" box="[199,433,150,176]" pageId="8" pageNumber="389">Material examined.</emphasis>
<materialsCitation id="61BB3CC5FFB4FF98FE49F7BF839DF3DA" ID-GBIF-Occurrence="3068500304" collectingDate="1985-04" collectionCode="BMNH" collectorName="J. H. Martin &amp; Wallace" country="Indonesia" elevation="1140" location="Clark's Camp" municipality="Dumoga-Bone National Park" pageId="8" pageNumber="389" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<typeStatus id="0E68883AFFB4FF98FE49F7BF852AF3BE" box="[440,549,151,177]" pageId="8" pageNumber="389" type="holotype">Holotype</typeStatus>
:
<specimenCount id="C7D5FD11FFB4FF98FDC1F7BF8548F3BE" box="[560,583,151,177]" pageId="8" pageNumber="389" type="male">♂</specimenCount>
,
<collectingCountry id="A9C47608FFB4FF98FDA2F7BF85CCF3BE" box="[595,707,151,177]" name="Indonesia" pageId="8" pageNumber="389">Indonesia</collectingCountry>
,
<collectingRegion id="1317F87AFFB4FF98FD3EF7BF8473F3BE" box="[719,892,151,177]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="389">Sulawesi Utara</collectingRegion>
,
<collectingMunicipality id="3108ACE2FFB4FF98FC79F7BF83DDF3BE" box="[904,1234,151,177]" pageId="8" pageNumber="389">Dumoga-Bone National Park</collectingMunicipality>
,
<location id="D40C6043FFB4FF98FB2FF7BF8272F3BE" LSID="urn:lsid:plazi:treatment:597A878EFFB9FF98FF66F1E285B0F26A:D40C6043FFB4FF98FB2FF7BF8272F3BE" box="[1246,1405,151,177]" country="Indonesia" municipality="Dumoga-Bone National Park" name="Clark's Camp" pageId="8" pageNumber="389" stateProvince="Sulawesi Utara">Clark’s Camp</location>
, at light,
<quantity id="162B9B7DFFB4FF98FF28F7938627F3DA" box="[217,296,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="8" pageNumber="389" unit="m" value="1140.0">
<elevation id="5AFED1ABFFB4FF98FF28F7938627F3DA" box="[217,296,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="8" pageNumber="389" unit="m" value="1140.0">1140m</elevation>
</quantity>
,
<collectorName id="7C26534EFFB4FF98FEC2F79386B9F3DA" box="[307,438,187,213]" pageId="8" pageNumber="389">J.H. Martin</collectorName>
,
<date id="A56D1058FFB4FF98FE33F7928518F3DA" box="[450,535,186,213]" pageId="8" pageNumber="389" value="1985-04">
<collectingDate id="B529E9B0FFB4FF98FE33F7928518F3DA" box="[450,535,186,213]" pageId="8" pageNumber="389" value="1985-04">iv.1985</collectingDate>
</date>
, RESL-Project
<collectorName id="7C26534EFFB4FF98FD37F793842DF3DA" box="[710,802,187,213]" pageId="8" pageNumber="389">Wallace</collectorName>
, NHMUK 010591767 (
<collectionCode id="B7C2AE5DFFB4FF98FBDAF793838AF3DA" box="[1067,1157,187,213]" pageId="8" pageNumber="389">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D16C3698FFB4FF98FF36F7F784B4F212" blockId="8.[151,1436,150,357]" pageId="8" pageNumber="389">
<materialsCitation id="61BB3CC5FFB4FF98FF36F7F78430F3F6" ID-GBIF-Occurrence="3068500312" box="[199,831,222,249]" collectingDate="1985-05" collectionCode="BMNH" collectorName="J. H. Martin &amp; Wallace" country="Indonesia" elevation="1140" location="Clark's Camp" municipality="Dumoga-Bone National Park" pageId="8" pageNumber="389" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
Other material.
<specimenCount id="C7D5FD11FFB4FF98FE87F7F78696F3F6" box="[374,409,223,249]" pageId="8" pageNumber="389" type="female">1♀</specimenCount>
, same data as holotype except
<date id="A56D1058FFB4FF98FD00F7F68430F3F6" box="[753,831,222,249]" pageId="8" pageNumber="389" value="1985-05">
<collectingDate id="B529E9B0FFB4FF98FD00F7F68430F3F6" box="[753,831,222,249]" pageId="8" pageNumber="389" value="1985-05">v.1985</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="61BB3CC5FFB4FF98FCBBF7F784B8F212" ID-GBIF-Occurrence="3068500319" collectingDate="1985-02" collectionCode="BMNH" country="Indonesia" location="Wallace" municipality="Dumoga-Bone National Park" pageId="8" pageNumber="389" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
<specimenCount id="C7D5FD11FFB4FF98FCBBF7F78464F3F6" box="[842,875,223,249]" pageId="8" pageNumber="389" type="female">1♀</specimenCount>
1? (abdomen missing),
<collectingCountry id="A9C47608FFB4FF98FB80F7F783EEF3F6" box="[1137,1249,223,249]" name="Indonesia" pageId="8" pageNumber="389">Indonesia</collectingCountry>
,
<collectingRegion id="1317F87AFFB4FF98FB1DF7F78297F3F6" box="[1260,1432,223,249]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="389">Sulawesi Utara</collectingRegion>
,
<collectingMunicipality id="3108ACE2FFB4FF98FF66F62B86EFF212" box="[151,480,259,285]" pageId="8" pageNumber="389">Dumoga-Bone National Park</collectingMunicipality>
,
<date id="A56D1058FFB4FF98FE1DF62A8533F212" box="[492,572,258,285]" pageId="8" pageNumber="389" value="1985-02">
<collectingDate id="B529E9B0FFB4FF98FE1DF62A8533F212" box="[492,572,258,285]" pageId="8" pageNumber="389" value="1985-02">ii.1985</collectingDate>
</date>
, RESL-Project
<location id="D40C6043FFB4FF98FD1DF62B844AF212" LSID="urn:lsid:plazi:treatment:597A878EFFB9FF98FF66F1E285B0F26A:D40C6043FFB4FF98FD1DF62B844AF212" box="[748,837,259,285]" country="Indonesia" municipality="Dumoga-Bone National Park" name="Wallace" pageId="8" pageNumber="389" stateProvince="Sulawesi Utara">Wallace</location>
(
<collectionCode id="B7C2AE5DFFB4FF98FCA5F62B84A1F212" box="[852,942,259,285]" pageId="8" pageNumber="389">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB4FF98FF36F60F8371F24E" box="[199,1150,295,321]" pageId="8" pageNumber="389" type="etymology">
<paragraph id="D16C3698FFB4FF98FF36F60F8371F24E" blockId="8.[151,1436,150,357]" box="[199,1150,295,321]" pageId="8" pageNumber="389">
<emphasis id="E3A7EA8AFFB4FF98FF36F60F8641F24E" bold="true" box="[199,334,295,321]" pageId="8" pageNumber="389">Etymology.</emphasis>
Named after the ventral spine on the caudal extension of the connective.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB4FF98FF36F66385B0F26A" box="[199,703,331,357]" pageId="8" pageNumber="389" type="discussion">
<paragraph id="D16C3698FFB4FF98FF36F66385B0F26A" blockId="8.[151,1436,150,357]" box="[199,703,331,357]" pageId="8" pageNumber="389">
<emphasis id="E3A7EA8AFFB4FF98FF36F6638634F26A" bold="true" box="[199,315,331,357]" pageId="8" pageNumber="389">Remarks.</emphasis>
See Remarks under
<taxonomicName id="16D34D1BFFB4FF98FDD2F66385B7F26A" box="[547,696,331,357]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="389" phylum="Arthropoda" rank="species" species="nudapenis">
<emphasis id="E3A7EA8AFFB4FF98FDD2F6638538F26B" box="[547,567,331,356]" italics="true" pageId="8" pageNumber="389">H</emphasis>
.
<emphasis id="E3A7EA8AFFB4FF98FDB4F66385B7F26A" box="[581,696,331,357]" italics="true" pageId="8" pageNumber="389">nudapenis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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