347 lines
45 KiB
XML
347 lines
45 KiB
XML
<document id="9B2E99D93AFE764A0C465BBEE0E581DB" ID-CLB-Dataset="45928" ID-DOI="10.11646/zootaxa.3793.2.3" ID-GBIF-Dataset="cec96d21-19fa-4e66-8305-9e70166f04f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="230581" ID-ZooBank="5E6DB00C-E98F-4D9D-958B-95B634E6D9E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460412815762" checkinUser="plazi" docAuthor="Baca, Stephen M., Gustafson, Grey T., Toledo, Mario & Miller, Kelly B." docDate="2014" docId="03D9AD22FFBFFFFF75A8FAC0FAFC93EB" docLanguage="en" docName="zt03793p246.pdf" docOrigin="Zootaxa 3793 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liocanthydrus octoguttatus Zimmermann 1921" docType="treatment" docVersion="9" lastPageNumber="245" masterDocId="FFE0D55AFFB2FFF1753FFF9FFF65972F" masterDocTitle="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species" masterLastPageNumber="246" masterPageNumber="231" pageNumber="244" updateTime="1698301202102" updateUser="ExternalLinkService">
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<mods:title id="650F0DC143986C279B339AF75CE44F5A">Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species</mods:title>
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<mods:name id="2A3385137DBA8E9619308839309DC7AF" type="personal">
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<mods:namePart id="BAE009A63EAB3A1339D5284D5180B12F">Baca, Stephen M.</mods:namePart>
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<mods:namePart id="9149A73E7F06F58CCDE57AC68A195119">Gustafson, Grey T.</mods:namePart>
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<mods:namePart id="0B2F0A950EC8B080C887982DE0112061">Toledo, Mario</mods:namePart>
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<mods:name id="A40D4644A3FEAD7BBF7DC04586C33DED" type="personal">
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<mods:namePart id="589F23AA544255BEDC45CB43ADE16329">Miller, Kelly B.</mods:namePart>
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<mods:title id="3D24BECAB0E08E984F10272D32A80E2A">Zootaxa</mods:title>
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<mods:date id="583520DC44273761B3A166D405E41A0F">2014</mods:date>
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<mods:number id="CB2525DCDD162188D138ABEA82B02D7B">3793</mods:number>
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<treatment id="03D9AD22FFBFFFFF75A8FAC0FAFC93EB" ID-DOI="http://doi.org/10.5281/zenodo.6131943" ID-GBIF-Taxon="119423979" ID-Zenodo-Dep="6131943" LSID="urn:lsid:plazi:treatment:03D9AD22FFBFFFFF75A8FAC0FAFC93EB" httpUri="http://treatment.plazi.org/id/03D9AD22FFBFFFFF75A8FAC0FAFC93EB" lastPageId="14" lastPageNumber="245" pageId="13" pageNumber="244">
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<subSubSection id="C36A4FBFFFBFFFFC75A8FAC0FDBC9257" box="[151,729,1375,1400]" pageId="13" pageNumber="244" type="nomenclature">
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<paragraph id="8BCF1C34FFBFFFFC75A8FAC0FDBC9257" blockId="13.[151,729,1375,1400]" box="[151,729,1375,1400]" pageId="13" pageNumber="244">
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<heading id="D087AB58FFBFFFFC75A8FAC0FDBC9257" bold="true" box="[151,729,1375,1400]" fontSize="10" level="2" pageId="13" pageNumber="244" reason="2">
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<taxonomicName id="4C7067B7FFBFFFFC75A8FAC0FDBC9257" authority="Zimmermann, 1921" authorityName="Zimmermann" authorityYear="1921" box="[151,729,1375,1400]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus">
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<emphasis id="B904C026FFBFFFFC75A8FAC0FDBC9257" bold="true" box="[151,729,1375,1400]" pageId="13" pageNumber="244">
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<emphasis id="B904C026FFBFFFFC75A8FAC0FEB69258" bold="true" box="[151,467,1375,1400]" italics="true" pageId="13" pageNumber="244">Liocanthydrus octoguttatus</emphasis>
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(
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<bibRefCitation id="EFE161C5FFBFFFFC74DEFAC0FDB49257" author="Zimmermann" box="[481,721,1375,1400]" pageId="13" pageNumber="244" refString="Zimmermann, A. (1921) Beitrage zur Kenntnis der sudamerikanischen Schwimmkaferfauna nebst 41 Neubeschreibungen. Archiv fur Naturgeschichte, 87 (3), 181 - 206." type="journal article" year="1921">Zimmermann, 1921</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36A4FBFFFBFFFFC75A8FA37FBA192D4" pageId="13" pageNumber="244" type="reference_group">
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<paragraph id="8BCF1C34FFBFFFFC75A8FA37FC3F92F3" blockId="13.[151,1220,1448,1531]" pageId="13" pageNumber="244">
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<treatmentCitationGroup id="AB603B1AFFBFFFFC75A8FA37FC3F92F3" pageId="13" pageNumber="244">
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<taxonomicName id="4C7067B7FFBFFFFC75A8FA37FDF89291" authority="Zimmermann (1921: 189)" authorityName="Zimmermann" authorityPageNumber="189" authorityYear="1921" box="[151,669,1448,1470]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus">
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<emphasis id="B904C026FFBFFFFC75A8FA37FEF59291" box="[151,400,1448,1470]" italics="true" pageId="13" pageNumber="244">Canthydrus octoguttatus</emphasis>
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<treatmentCitation id="0AD13A25FFBFFFFC74A8FA37FDF89291" author="Zimmermann" box="[407,669,1448,1470]" page="189" pageId="13" pageNumber="244" year="1921">
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<bibRefCitation id="EFE161C5FFBFFFFC74A8FA37FDF89291" author="Zimmermann" box="[407,669,1448,1470]" pageId="13" pageNumber="244" refString="Zimmermann, A. (1921) Beitrage zur Kenntnis der sudamerikanischen Schwimmkaferfauna nebst 41 Neubeschreibungen. Archiv fur Naturgeschichte, 87 (3), 181 - 206." type="journal article" year="1921">Zimmermann (1921: 189)</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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(orig. descr.);
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<treatmentCitation id="0AD13A25FFBFFFFC760DFA37FB409291" author="Blackwelder" box="[818,1061,1448,1470]" page="73" pageId="13" pageNumber="244" year="1944">
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<bibRefCitation id="EFE161C5FFBFFFFC760DFA37FB409291" author="Blackwelder" box="[818,1061,1448,1470]" pageId="13" pageNumber="244" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. United States National Museum Bulletin, 185, 73. http: // dx. doi. org / 10.5479 / si. 03629236.185. i" type="journal article" year="1944">Blackwelder (1944: 73)</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
<taxonomicName id="4C7067B7FFBFFFFC75A8FA59FD3492F3" authority="Zimmermann" authorityName="Zimmermann" box="[151,593,1478,1501]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus">
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||
<emphasis id="B904C026FFBFFFFC75A8FA59FEC992F2" box="[151,428,1478,1501]" italics="true" pageId="13" pageNumber="244">Liocanthydrus octoguttatus</emphasis>
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(Zimmermann)
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</taxonomicName>
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;
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<treatmentCitation id="0AD13A25FFBFFFFC7764FA59FC7292F3" author="Nilsson" box="[603,791,1478,1500]" page="28" pageId="13" pageNumber="244" year="2011">
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<bibRefCitation id="EFE161C5FFBFFFFC7764FA59FC7292F3" author="Nilsson" box="[603,791,1478,1500]" pageId="13" pageNumber="244" refString="Nilsson, A. N. (2011) A world catalogue of the family Noteridae, or the burrowing water beetles (Coleoptera, Adephaga). Version 16. VIII. 2011, 1 - 54 pp. Available from: http: // www 2. emg. umu. se / projects / biginst / andersn / WCN / WCN _ 20110816. pdf (accessed 9 January 2014)" type="journal article" year="2011">Nilsson (2011: 28)</bibRefCitation>
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||
</treatmentCitation>
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||
(cat.).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BCF1C34FFBFFFFC75A8FA7AFBA192D4" blockId="13.[151,1220,1448,1531]" box="[151,1220,1509,1531]" pageId="13" pageNumber="244">
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||
<treatmentCitationGroup id="AB603B1AFFBFFFFC75A8FA7AFBA192D4" box="[151,1220,1509,1531]" pageId="13" pageNumber="244">
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<taxonomicName id="4C7067B7FFBFFFFC75A8FA7AFDBA92D4" authority="Zimmermann" authorityName="Zimmermann" box="[151,735,1509,1531]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus" subGenus="Liocanthydrus">
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||
<emphasis id="B904C026FFBFFFFC75A8FA7AFE6892D4" box="[151,269,1509,1531]" italics="true" pageId="13" pageNumber="244">Canthydrus</emphasis>
|
||
(
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||
<emphasis id="B904C026FFBFFFFC7423FA7AFECB92D4" box="[284,430,1509,1531]" italics="true" pageId="13" pageNumber="244">Liocanthydrus</emphasis>
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||
)
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||
<emphasis id="B904C026FFBFFFFC7482FA79FD5F92D4" box="[445,570,1510,1531]" italics="true" pageId="13" pageNumber="244">octoguttatus</emphasis>
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||
(Zimmermann)
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||
</taxonomicName>
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||
;
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||
<treatmentCitation id="0AD13A25FFBFFFFC77D7FA7AFCCB92D4" author="Guignot" box="[744,942,1509,1531]" page="43" pageId="13" pageNumber="244" year="1957">
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||
<bibRefCitation id="EFE161C5FFBFFFFC77D7FA7AFCCB92D4" author="Guignot" box="[744,942,1509,1531]" pageId="13" pageNumber="244" refString="Guignot, F. (1957) Contribution a la connaissance des dytiscides Sud-Americains. Revue Francaise d'Entomologie, 24, 33 - 45." type="journal article" year="1957">Guignot (1957: 43)</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
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||
<treatmentCitation id="0AD13A25FFBFFFFC7686FA7AFBE792D4" author="Nilsson" box="[953,1154,1509,1531]" page="110" pageId="13" pageNumber="244" year="2005">
|
||
<bibRefCitation id="EFE161C5FFBFFFFC7686FA7AFBE792D4" author="Nilsson" box="[953,1154,1509,1531]" pageId="13" pageNumber="244" refString="Nilsson, A. N. (2005) Family Noteridae (Coleoptera, Adephaga). In: Nilsson, A. N. & van Vondel, B. J., (Eds.), Amphizoidae, Aspidytidae, Haliplidae, Noteridae and Paelobiidae (Coleoptera, Adephaga). World Catalogue of Insects. Vol. 7. Apollo Books, Stenstrup, 87 - 153. http: // dx. doi. org / 10.1649 / 0010 - 065 x (2006) 60 [305: aahnap] 2.0. co; 2" type="book chapter" year="2005">Nilsson (2005: 110)</bibRefCitation>
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||
</treatmentCitation>
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||
(cat.).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36A4FBFFFBFFFFC75A8F9B8FEE59087" pageId="13" pageNumber="244" type="materials_examined">
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||
<paragraph id="8BCF1C34FFBFFFFC75A8F9B8FCD7916F" blockId="13.[151,1436,1575,2033]" box="[151,946,1575,1600]" pageId="13" pageNumber="244">
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<emphasis id="B904C026FFBFFFFC75A8F9B8FE50916F" bold="true" box="[151,309,1575,1600]" pageId="13" pageNumber="244">
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||
<typeStatus id="54CBA296FFBFFFFC75A8F9B8FFB7916F" box="[151,210,1575,1600]" pageId="13" pageNumber="244">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<collectingCountry id="F3675CA4FFBFFFFC7403F9B8FEE6916F" box="[316,387,1575,1600]" name="Brazil" pageId="13" pageNumber="244">Brazil</collectingCountry>
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||
, São Paulo State, São Paulo, Sierra da Cantareira.
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</paragraph>
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||
<paragraph id="8BCF1C34FFBFFFFC75F8F9D3FEE59087" blockId="13.[151,1436,1575,2033]" pageId="13" pageNumber="244">
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<emphasis id="B904C026FFBFFFFC75F8F9D3FD67914A" bold="true" box="[199,514,1612,1637]" pageId="13" pageNumber="244">
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<typeStatus id="54CBA296FFBFFFFC75F8F9D3FE67914A" box="[199,258,1612,1637]" pageId="13" pageNumber="244">Type</typeStatus>
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||
material examined.
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||
</emphasis>
|
||
<typeStatus id="54CBA296FFBFFFFC772DF9D2FDE7914A" box="[530,642,1613,1637]" pageId="13" pageNumber="244" type="lectotype">Lectotype</typeStatus>
|
||
(male) (designated here): “Riedel” [printed] “
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||
<collectingCountry id="F3675CA4FFBFFFFC71FFF9D3FA4F914B" box="[1216,1322,1612,1636]" name="Brazil" pageId="13" pageNumber="244">BRASIL</collectingCountry>
|
||
/S. Paulo/ Umgebung v. S. Paulo/Sierra da Cantareira/
|
||
<date id="FFCE3AF4FFBFFFFC77AAF9F0FC4B91A7" box="[661,814,1647,1672]" pageId="13" pageNumber="244" value="1900-10-19">19. Oct. 1900</date>
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||
” [printed], “
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<typeStatus id="54CBA296FFBFFFFC7680F9EFFC9E91A7" box="[959,1019,1648,1672]" pageId="13" pageNumber="244">Type</typeStatus>
|
||
” [small round label, handwriting A. Zimmermann], “
|
||
<taxonomicName id="4C7067B7FFBFFFFC7466F90BFD169183" box="[345,627,1684,1709]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBFFFFC7466F90BFD169183" box="[345,627,1684,1709]" italics="true" pageId="13" pageNumber="244">Canthydrus octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
” [printed], “Samml. A. Zimmermann” [printed], plus an additional, unwritten, red label and our
|
||
<typeStatus id="54CBA296FFBFFFFC74DAF928FD2991FF" box="[485,588,1719,1744]" pageId="13" pageNumber="244" type="lectotype">lectotype</typeStatus>
|
||
designation label: “
|
||
<typeStatus id="54CBA296FFBFFFFC760CF927FCB091FF" box="[819,981,1720,1744]" pageId="13" pageNumber="244" type="lectotype">LECTOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBFFFFC76E2F928FB9091FF" box="[989,1269,1719,1744]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBFFFFC76E2F928FB9091FF" box="[989,1269,1719,1744]" italics="true" pageId="13" pageNumber="244">Canthydrus octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EFE161C5FFBFFFFC71C2F928FFAB91DA" author="Zimmermann" pageId="13" pageNumber="244" refString="Zimmermann, A. (1921) Beitrage zur Kenntnis der sudamerikanischen Schwimmkaferfauna nebst 41 Neubeschreibungen. Archiv fur Naturgeschichte, 87 (3), 181 - 206." type="journal article" year="1921">Zimmermann, 1921</bibRefCitation>
|
||
/det. Baca, Gustafson, Toledo, Miller 2014” [red; printed]. (
|
||
<collectionCode id="ED6184F1FFBFFFFC764FF943FCCA91DB" box="[880,943,1756,1780]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="13" pageNumber="244">ZSM</collectionCode>
|
||
).
|
||
<typeStatus id="54CBA296FFBFFFFC76FDF943FB3F91DA" box="[962,1114,1756,1781]" pageId="13" pageNumber="244" type="paralectotype">Paralectotype</typeStatus>
|
||
(female): “Riedel” [printed] “
|
||
<collectingCountry id="F3675CA4FFBFFFFC759EF89FFE6E9037" box="[161,267,1792,1816]" name="Brazil" pageId="13" pageNumber="244">BRASIL</collectingCountry>
|
||
/S. Paulo/Umgebung v. S. Paulo/Sierra da Cantareira/
|
||
<date id="FFCE3AF4FFBFFFFC765EF960FC919037" box="[865,1012,1791,1816]" pageId="13" pageNumber="244" value="1900-10-19">19. Oct. 1900</date>
|
||
” [printed], “
|
||
<typeStatus id="54CBA296FFBFFFFC7141F89FFBDF9037" box="[1150,1210,1792,1816]" pageId="13" pageNumber="244">Type</typeStatus>
|
||
” [small round label, handwriting A. Zimmermann], “Samml. A. Zimmermann” [printed], “
|
||
<typeStatus id="54CBA296FFBFFFFC76F5F8BBFB589013" box="[970,1085,1828,1852]" pageId="13" pageNumber="244" type="paratype">Paratypus</typeStatus>
|
||
” [red, unauthorized curatorial label; printed], plus our yellow
|
||
<typeStatus id="54CBA296FFBFFFFC74C6F8D8FDF5904F" box="[505,656,1863,1888]" pageId="13" pageNumber="244" type="paralectotype">paralectotype</typeStatus>
|
||
label: “
|
||
<typeStatus id="54CBA296FFBFFFFC77D8F8D7FCB4904F" box="[743,977,1864,1888]" pageId="13" pageNumber="244" type="paralectotype">PARALECTOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBFFFFC76E6F8D8FB8B904F" box="[985,1262,1863,1888]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBFFFFC76E6F8D8FB8B904F" box="[985,1262,1863,1888]" italics="true" pageId="13" pageNumber="244">Canthydrus octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EFE161C5FFBFFFFC71C3F8D8FFAB90AA" author="Zimmermann" pageId="13" pageNumber="244" refString="Zimmermann, A. (1921) Beitrage zur Kenntnis der sudamerikanischen Schwimmkaferfauna nebst 41 Neubeschreibungen. Archiv fur Naturgeschichte, 87 (3), 181 - 206." type="journal article" year="1921">Zimmermann, 1921</bibRefCitation>
|
||
/det. Baca, Gustafson, Toledo, Miller 2014” [yellow; printed] (
|
||
<collectionCode id="ED6184F1FFBFFFFC76FAF8F3FB6190AB" box="[965,1028,1900,1924]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="13" pageNumber="244">ZSM</collectionCode>
|
||
).
|
||
<emphasis id="B904C026FFBFFFFC7121F8F3FB3A90AA" bold="true" box="[1054,1119,1900,1925]" pageId="13" pageNumber="244">Note:</emphasis>
|
||
All label descriptions are included in brackets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBFFFFF75F8F82BFC969537" lastPageId="14" lastPageNumber="245" pageId="13" pageNumber="244" type="diagnosis">
|
||
<paragraph id="8BCF1C34FFBFFFFF75F8F82BFC109633" blockId="13.[151,1436,1575,2033]" lastBlockId="14.[151,1437,151,1221]" lastPageId="14" lastPageNumber="245" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8F82BFE2490E2" bold="true" box="[199,321,1972,1997]" pageId="13" pageNumber="244">Diagnosis.</emphasis>
|
||
Elytra maculate, distinguishable from other members of
|
||
<taxonomicName id="4C7067B7FFBFFFFC7126F82BFBDE90E2" box="[1049,1211,1972,1997]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBFFFFC7126F82BFBDE90E2" box="[1049,1211,1972,1997]" italics="true" pageId="13" pageNumber="244">Liocanthydrus</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) small eye size relative to head capsule, HW/EW = 1.55 (
|
||
<figureCitation id="134B00B1FFBFFFFC715DF847FBC290DF" box="[1122,1191,2008,2033]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="13" pageNumber="244">Fig. 9</figureCitation>
|
||
); b) elytral patterning as in
|
||
<figureCitation id="134B00B1FFBCFFFF75EDFF08FE73979F" box="[210,278,151,176]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 9</figureCitation>
|
||
, with posterior bands oblique and nearly reduced to spots; anterior bands reduced to spots; c) aedeagus as in
|
||
<figureCitation id="134B00B1FFBCFFFF75E8FF23FE4A97FB" box="[215,303,188,213]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 22</figureCitation>
|
||
a–e; median lobe slightly sinuate in dorsal aspect (
|
||
<figureCitation id="134B00B1FFBCFFFF764AFF23FCAE97FA" box="[885,971,188,213]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 22</figureCitation>
|
||
b), distally very elongate and subparallel with rounded apex in lateral aspect, not attenuate (
|
||
<figureCitation id="134B00B1FFBCFFFF77EDFF40FC4C97D7" box="[722,809,223,248]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 22</figureCitation>
|
||
a, c); left lateral lobe narrowed distally (
|
||
<figureCitation id="134B00B1FFBCFFFF71D0FF40FA2697D7" box="[1263,1347,223,248]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 22</figureCitation>
|
||
d); right lateral lobe triangular, distal angle acute, not projected (
|
||
<figureCitation id="134B00B1FFBCFFFF7637FE9BFC3E9633" box="[776,859,260,285]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 22</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FEB8FC969537" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="245">
|
||
Externally
|
||
<taxonomicName id="4C7067B7FFBCFFFF747CFEB6FE8F966F" box="[323,490,297,320]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBCFFFF747CFEB6FE8F966F" box="[323,490,297,320]" italics="true" pageId="14" pageNumber="245">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
appears very similar to
|
||
<taxonomicName id="4C7067B7FFBCFFFF77C6FEB6FC02966F" box="[761,871,297,320]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBCFFFF77C6FEB6FC02966F" box="[761,871,297,320]" italics="true" pageId="14" pageNumber="245">L. nanops</emphasis>
|
||
</taxonomicName>
|
||
and also, though less so, to
|
||
<emphasis id="B904C026FFBCFFFF719CFEB8FAF9966F" box="[1187,1436,295,320]" italics="true" pageId="14" pageNumber="245">
|
||
<taxonomicName id="4C7067B7FFBCFFFF719CFEB8FAFC966F" box="[1187,1433,295,320]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="clayae">Liocanthydrus clayae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Externally
|
||
<taxonomicName id="4C7067B7FFBCFFFF7429FED2FEA4964B" box="[278,449,333,356]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBCFFFF7429FED2FEA4964B" box="[278,449,333,356]" italics="true" pageId="14" pageNumber="245">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C7067B7FFBCFFFF77F3FED2FC53964A" box="[716,822,332,357]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBCFFFF77F3FED2FC53964A" box="[716,822,332,357]" italics="true" pageId="14" pageNumber="245">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
by its small relative eye size and the more strongly obliquely oriented posterior elytral maculae that also extend anteriorly to the basal halves of the elytra.
|
||
<taxonomicName id="4C7067B7FFBCFFFF75A8FE0BFEAE9683" box="[151,459,404,429]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBCFFFF75A8FE0BFEAE9683" box="[151,459,404,429]" italics="true" pageId="14" pageNumber="245">Liocanthydrus octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from both
|
||
<taxonomicName id="4C7067B7FFBCFFFF763FFE0AFCCC9683" box="[768,937,405,428]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBCFFFF763FFE0AFCCC9683" box="[768,937,405,428]" italics="true" pageId="14" pageNumber="245">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7067B7FFBCFFFF76E0FE0AFB2B9683" box="[991,1102,405,428]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBCFFFF76E0FE0AFB2B9683" box="[991,1102,405,428]" italics="true" pageId="14" pageNumber="245">L. nanops</emphasis>
|
||
</taxonomicName>
|
||
by its distinct aedeagus, with a median lobe that is distally very elongate and subparallel in lateral aspect, not attenuate as in both
|
||
<taxonomicName id="4C7067B7FFBCFFFF703BFE26FA0E96FF" box="[1284,1387,439,464]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBCFFFF703BFE26FA0E96FF" box="[1284,1387,439,464]" italics="true" pageId="14" pageNumber="245">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7067B7FFBCFFFF75A8FE42FE2896DA" authority="Also" authorityName="Also" box="[151,333,476,501]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBCFFFF75A8FE42FE6896DB" box="[151,269,477,500]" italics="true" pageId="14" pageNumber="245">L. nanops,</emphasis>
|
||
Also
|
||
</taxonomicName>
|
||
, the right lateral lobe of the aedeagus is triangular with an acute distal angle, similar to
|
||
<taxonomicName id="4C7067B7FFBCFFFF700BFE42FAF996DA" box="[1332,1436,476,501]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBCFFFF700BFE42FAF996DA" box="[1332,1436,476,501]" italics="true" pageId="14" pageNumber="245">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
but not subtriangular with the distal angle strongly projected as in
|
||
<emphasis id="B904C026FFBCFFFF7642FD9EFC969537" box="[893,1011,513,536]" italics="true" pageId="14" pageNumber="245">
|
||
<taxonomicName id="4C7067B7FFBCFFFF7642FD9EFC959537" box="[893,1008,513,536]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="nanops">L. nanops</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBCFFFF75F8FDBBFDB194E7" pageId="14" pageNumber="245" type="description">
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FDBBFBE1954F" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FDBBFE3C9512" bold="true" box="[199,345,548,573]" pageId="14" pageNumber="245">Description.</emphasis>
|
||
Measurements. TL =
|
||
<quantity id="4C88B1D1FFBCFFFF776EFDBBFDD29513" box="[593,695,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="14" pageNumber="245" unit="mm" value="3.05">3.05 mm</quantity>
|
||
, GW =
|
||
<quantity id="4C88B1D1FFBCFFFF7630FDBBFC0D9513" box="[783,872,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="245" unit="mm" value="1.4">1.4 mm</quantity>
|
||
, TL/GW = 2.18, HW =
|
||
<quantity id="4C88B1D1FFBCFFFF7149FDBBFBB99513" box="[1142,1244,548,573]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="14" pageNumber="245" unit="mm" value="0.85">0.85 mm</quantity>
|
||
, EW =
|
||
<quantity id="4C88B1D1FFBCFFFF700EFDBBFAF29513" box="[1329,1431,548,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="245" unit="mm" value="0.55">0.55 mm</quantity>
|
||
, HW/EW = 1.55, PNWpost. =
|
||
<quantity id="4C88B1D1FFBCFFFF74D8FDD8FD289570" box="[487,589,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="14" pageNumber="245" unit="mm" value="1.35">1.35 mm</quantity>
|
||
, PNWant. =
|
||
<quantity id="4C88B1D1FFBCFFFF77E3FDD8FC279570" box="[732,834,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="14" pageNumber="245" unit="mm" value="0.85">0.85 mm</quantity>
|
||
, PNWpost./PNWant. = 1.59.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FDF3FF97943B" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FDF3FE2795AA" box="[199,322,620,645]" italics="true" pageId="14" pageNumber="245">Coloration</emphasis>
|
||
. Elytra maculate, with bands or spots; color of elytra dark reddish brown; color of head, pronotum and elytral maculae yellow; band pattern as in
|
||
<figureCitation id="134B00B1FFBCFFFF77F1FD10FC729587" box="[718,791,655,680]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 9</figureCitation>
|
||
, posterior elytral band strongly oblique and extending anterolaterally to basal halves of elytra, anterior bands reduced to spots and laterally oriented, with medial spots larger, more distinct than lateral spots; anteromedial portion of pronotal disc with distinct large dark maculation (
|
||
<figureCitation id="134B00B1FFBCFFFF75A0FD63FF81943B" box="[159,228,764,789]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FC80FB489417" blockId="14.[151,1437,151,1221]" box="[199,1069,799,824]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FC80FF9A9417" box="[199,255,799,824]" italics="true" pageId="14" pageNumber="245">Body</emphasis>
|
||
. Body forms as in
|
||
<figureCitation id="134B00B1FFBCFFFF74EBFC80FD729417" box="[468,535,799,824]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 9</figureCitation>
|
||
, oval, moderately narrow, attenuate posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FCDBFDB194E7" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FCDBFE549472" box="[199,305,836,861]" italics="true" pageId="14" pageNumber="245">Abdomen</emphasis>
|
||
. Aedeagus as in
|
||
<figureCitation id="134B00B1FFBCFFFF74C8FCDBFD2A9473" box="[503,591,836,861]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 22</figureCitation>
|
||
a–e; median lobe sinuate in dorsal aspect (
|
||
<figureCitation id="134B00B1FFBCFFFF7109FCDBFBEF9472" box="[1078,1162,836,861]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 22</figureCitation>
|
||
b), distally elongate and subparallel in lateral aspect, apex rounded. (
|
||
<figureCitation id="134B00B1FFBCFFFF77BEFCF8FDB294AF" box="[641,727,871,896]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Figs 22</figureCitation>
|
||
a, c); left lateral lobe broad and rounded near base but narrowed distally, setal fringe expanded to small field apically and barely extending basad of distal third (
|
||
<figureCitation id="134B00B1FFBCFFFF71D3FC13FA24948A" box="[1260,1345,908,933]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 22</figureCitation>
|
||
d); right lateral lobe triangular, distal angle acute.(
|
||
<figureCitation id="134B00B1FFBCFFFF7757FC30FDDF94E7" box="[616,698,943,968]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="14" pageNumber="245">Fig. 22</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBCFFFF75F8FC4BFD0A94C2" box="[199,623,980,1005]" pageId="14" pageNumber="245" type="biology_ecology">
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FC4BFD0A94C2" blockId="14.[151,1437,151,1221]" box="[199,623,980,1005]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FC4BFE4294C2" bold="true" box="[199,295,980,1005]" pageId="14" pageNumber="245">Biology.</emphasis>
|
||
No collection data available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBCFFFF75F8FC68FC0F933F" box="[199,874,1015,1040]" pageId="14" pageNumber="245" type="distribution">
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FC68FC0F933F" blockId="14.[151,1437,151,1221]" box="[199,874,1015,1040]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FC68FE05933F" bold="true" box="[199,352,1015,1040]" pageId="14" pageNumber="245">Distribution.</emphasis>
|
||
Known only from São Paulo,
|
||
<collectingCountry id="F3675CA4FFBCFFFF7789FC68FD9F933F" box="[694,762,1015,1040]" name="Brazil" pageId="14" pageNumber="245">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="134B00B1FFBCFFFF7636FC68FC3B933F" box="[777,862,1015,1040]" captionStart="FIGURE 23" captionStartId="7.[151,250,1491,1513]" captionText="FIGURE 23. Distribution map of Liocanthydrus species." pageId="14" pageNumber="245">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBCFFFF75F8FB83FAFC93EB" pageId="14" pageNumber="245" type="description">
|
||
<paragraph id="8BCF1C34FFBCFFFF75F8FB83FAFC93EB" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="245">
|
||
<emphasis id="B904C026FFBCFFFF75F8FB83FE5E931A" bold="true" box="[199,315,1052,1077]" pageId="14" pageNumber="245">Variation.</emphasis>
|
||
The variation exhibited by this species was difficult to assess due to the limited amount of material available for examination.
|
||
<collectingCountry id="F3675CA4FFBCFFFF74F1FBDFFE889377" box="[462,493,1088,1112]" name="American Samoa" pageId="14" pageNumber="245">As</collectingCountry>
|
||
mentioned, the
|
||
<typeStatus id="54CBA296FFBCFFFF7792FBDFFDB89377" box="[685,733,1088,1112]" pageId="14" pageNumber="245">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7067B7FFBCFFFF7634FBDEFCD29377" box="[779,951,1089,1112]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBCFFFF7634FBDEFCD29377" box="[779,951,1089,1112]" italics="true" pageId="14" pageNumber="245">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
has a large, very dark maculation on the anteromedial region of the pronotum (
|
||
<figureCitation id="134B00B1FFBCFFFF7750FBFBFDD99353" box="[623,700,1124,1149]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="14" pageNumber="245">Fig. 9</figureCitation>
|
||
), much larger and darker than any of the other species of
|
||
<taxonomicName id="4C7067B7FFBCFFFF75A8FB18FE5C938F" box="[151,313,1159,1184]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBCFFFF75A8FB18FE5C938F" box="[151,313,1159,1184]" italics="true" pageId="14" pageNumber="245">Liocanthydrus</emphasis>
|
||
</taxonomicName>
|
||
. However, the variation of this feature could not be assessed and may not be useful as a diagnostic character as similar though far less distinct markings were observed to vary within other species of
|
||
<taxonomicName id="4C7067B7FFBCFFFF71CFFB33FAF793EA" box="[1264,1426,1196,1221]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBCFFFF71CFFB33FAF793EA" box="[1264,1426,1196,1221]" italics="true" pageId="14" pageNumber="245">Liocanthydrus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |