109 lines
21 KiB
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109 lines
21 KiB
XML
<document id="74F095ABEB2391905C71155CC1431F50" ID-DOI="10.14446/FI.2016.161" ID-ISSN="2533-4069" ID-Zenodo-Dep="4773100" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621431429390" checkinUser="felipe" docAuthor="Laibl, Lukáš, Esteve, Jorge & Fatka, Oldřich" docDate="2016" docId="03D9472EFFC4FFC9068DFA4E0589FBA2" docLanguage="en" docName="FossilImprint.72.3-4.161-171.pdf" docOrigin="Fossil Imprint 72 (3 - 4)" docSource="http://dx.doi.org/10.14446/fi.2016.161" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Hydrocephalus BARRANDE 1846" docType="treatment" docVersion="3" lastPageNumber="166" masterDocId="FFE03F56FFC1FFCC0650FFF30605FFF6" masterDocTitle="Enrollment And Thoracic Morphology In Paradoxidid Trilobites From The Cambrian Of The Czech Republic" masterLastPageNumber="171" masterPageNumber="161" pageNumber="166" updateTime="1698921862362" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="9EB68781A58A543F7F17E09B3F74956C">Enrollment And Thoracic Morphology In Paradoxidid Trilobites From The Cambrian Of The Czech Republic</mods:title>
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<mods:namePart id="55351D29F385401CD0F2DB0299256915">Laibl, Lukáš</mods:namePart>
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<mods:namePart id="AB261FBD35CB8A3C2E79C42F27077EA8">Esteve, Jorge</mods:namePart>
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<mods:namePart id="267D3126FA39484CF8AB7BEC95CD4338">Fatka, Oldřich</mods:namePart>
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<mods:date id="297C72A29F70BB4CB9D84F3B40BD84AA">2016</mods:date>
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<subSubSection id="C36AA5B3FFC4FFC9068DFA4E0470FA22" box="[221,629,1469,1493]" pageId="5" pageNumber="166" type="nomenclature">
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<paragraph id="8BCFF638FFC4FFC9068DFA4E0470FA22" blockId="5.[221,629,1469,1493]" box="[221,629,1469,1493]" pageId="5" pageNumber="166">
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<heading id="D0874154FFC4FFC9068DFA4E0470FA22" box="[221,629,1469,1493]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="166" reason="2">
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<emphasis id="B9042A2AFFC4FFC9068DFA4E0470FA22" bold="true" box="[221,629,1469,1493]" pageId="5" pageNumber="166">
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<emphasis id="B9042A2AFFC4FFC9068DFA4E07CBFA23" bold="true" box="[221,462,1469,1493]" italics="true" pageId="5" pageNumber="166">
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<taxonomicName id="4C708DBBFFC4FFC9068DFA4E0781FA23" authorityName="BARRANDE" authorityYear="1846" box="[221,388,1469,1493]" class="Trilobita" family="Paradoxididae" genus="Hydrocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="5" pageNumber="166" phylum="Arthropoda" rank="genus">Hydrocephalus</taxonomicName>
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minor
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</emphasis>
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(
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<bibRefCitation id="EFE18BC9FFC4FFC9078CFA4E042FFA22" author="Boeck, C. P. B." box="[476,554,1469,1492]" pageId="5" pageNumber="166" pagination="11 - 44" refId="ref6124" refString="Boeck, C. P. B. (1827): Notiser til Laeren om Trilobiterne. - Magazin for Naturvidenskaberen, 8 (1): 11 - 44." type="journal article" year="1827">BOECK</bibRefCitation>
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, 1827)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36AA5B3FFC4FFC90766FA1E0589FBA2" pageId="5" pageNumber="166" type="description">
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<paragraph id="8BCFF638FFC4FFC90766FA1E0419F9F5" blockId="5.[310,540,1517,1539]" box="[310,540,1517,1539]" pageId="5" pageNumber="166">
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<figureCitation id="134BEABDFFC4FFC90766FA1E07B4F9F5" box="[310,433,1517,1539]" captionStart="Text-fig" captionStartId="4.[129,212,1963,1984]" captionTargetBox="[129,1482,151,1930]" captionTargetId="figure-14@4.[128,1482,151,1930]" captionTargetPageId="4" captionText="Text-fig. 3. Hydrocephalus minor (BOECK, 1827); Jince Formation, Hypagnostus parvifrons to Paradoxides paradoxissimus gracilis Biozones, Příbram-Jince Basin: a – specimen no. NM-L 46543, from dorsal (a1), oblique lateral (a2) and posterior (a3) view; b – specimen no. NM-L 46544, from dorsal (b1), and posterior (b2) view; c – specimen no. NM-L 46545, from dorsal (c1) and posterior (c2) view; d – specimen no. NM-L 46546, from dorsal (d1) and ventral (d2) view; e – specimen no. NM-L 41157, external (e1) and internal (e2) mold. Abbreviations: op – overlapping pleurae." figureDoi="http://doi.org/10.5281/zenodo.4773110" httpUri="https://zenodo.org/record/4773110/files/figure.png" pageId="5" pageNumber="166">Text-figs 3e</figureCitation>
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,
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<figureCitation id="134BEABDFFC4FFC907ECFA1E0419F9F5" box="[444,540,1517,1539]" captionStart="Text-fig" captionStartId="6.[130,214,1559,1580]" captionTargetBox="[129,1481,152,1512]" captionTargetId="figure-117@6.[129,1482,151,1512]" captionTargetPageId="6" captionText="Text-fig. 5. Thoracic structure, articulating and coaptative devices of H. minor (a, d, f–g), P. paradoxissimus gracilis (c, e, h–j) and A. cf. sacheri (b): a – specimen no. NM-L 46550, internal mold of pleurae; b – specimen no. NM-L 4776, internal mold of pleura; c – specimen no. NM-L 46551, internal mold of pleurae; d – specimen no. NM-L 46550, internal mold of axial rings and articulating half-rings; e – specimen no. NM-L 46552, internal mold of axial rings and articulating half-rings; f – specimen no. NM-L 46550, detail of pleural spine; g – specimen no. NM-L 41157, external mold of pleural spines; h – specimen no. NM-L 46547, external mold of pleural spines; i – specimen no. NM-L 46548, external mold of pleural spines; j – specimen no. NM-L 46549, pleural spines. Abbreviations: ahr – articulating half ring; alp – antero-lateral projection of flange; bp – backward projection at base of pleural spine; dtl – dorsal terrace lines; fl – flange; fu – fulcrum; ptl – petaloid-like terrace lines; vf – ventral furrow of pleural spine; vtl – ventral terrace lines." figureDoi="http://doi.org/10.5281/zenodo.4773118" httpUri="https://zenodo.org/record/4773118/files/figure.png" pageId="5" pageNumber="166">5a, d, f–g</figureCitation>
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</paragraph>
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<paragraph id="8BCFF638FFC4FFC906C0F9E80589FBA2" blockId="5.[106,745,1563,2098]" lastBlockId="5.[820,1459,829,1108]" pageId="5" pageNumber="166">
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The exoskeleton is micropygous and the overall outline of the trunk is parabolic (posterior width of the thorax ca. 50–55% of the anterior width). The pleural parts are wider (tr.) than in
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<emphasis id="B9042A2AFFC4FFC906BDF98F040EF965" box="[237,523,1659,1683]" italics="true" pageId="5" pageNumber="166">P. paradoxissimus gracilis</emphasis>
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, being ca. 1.6 times wider than the width of the corresponding axial ring (
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<figureCitation id="134BEABDFFC4FFC90621F94806F3F924" box="[113,246,1723,1746]" captionStart="Text-fig" captionStartId="4.[129,212,1963,1984]" captionTargetBox="[129,1482,151,1930]" captionTargetId="figure-14@4.[128,1482,151,1930]" captionTargetPageId="4" captionText="Text-fig. 3. Hydrocephalus minor (BOECK, 1827); Jince Formation, Hypagnostus parvifrons to Paradoxides paradoxissimus gracilis Biozones, Příbram-Jince Basin: a – specimen no. NM-L 46543, from dorsal (a1), oblique lateral (a2) and posterior (a3) view; b – specimen no. NM-L 46544, from dorsal (b1), and posterior (b2) view; c – specimen no. NM-L 46545, from dorsal (c1) and posterior (c2) view; d – specimen no. NM-L 46546, from dorsal (d1) and ventral (d2) view; e – specimen no. NM-L 41157, external (e1) and internal (e2) mold. Abbreviations: op – overlapping pleurae." figureDoi="http://doi.org/10.5281/zenodo.4773110" httpUri="https://zenodo.org/record/4773110/files/figure.png" pageId="5" pageNumber="166">Text-fig. 3e</figureCitation>
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). Several posterior-most pleurae are inclined backward. In the cross-section, the pleurae are considerably vaulted, without any prominent fulcrum, as was described by
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<bibRefCitation id="EFE18BC9FFC4FFC9063AF8E80747F8C4" author="Whittington, H. B." box="[106,322,1819,1842]" pageId="5" pageNumber="166" pagination="27 - 46" refId="ref8229" refString="Whittington, H. B. (1990): Articulation and Exuviation in Cambrian Trilobites. - Philosophical Transactions of the Royal Society of London, Series B, 329: 27 - 46. https: // doi. org / 10.1098 / rstb. 1990.0147" type="journal article" year="1990">Whittington (1990)</bibRefCitation>
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for the other Barrandian species,
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<emphasis id="B9042A2AFFC4FFC9049EF8EF06B4F8A5" italics="true" pageId="5" pageNumber="166">H. carens</emphasis>
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<bibRefCitation id="EFE18BC9FFC4FFC906E9F8C80771F8A4" author="Barrande, J." box="[185,372,1851,1874]" pageId="5" pageNumber="166" refId="ref5976" refString="Barrande, J. (1846): Notice preliminaire sur le Systeme Silurien et les trilobites de Boheme. - C. L. Hirschfeld, Leipsic [= Leipzig], vi + 97 pp." type="book" year="1846">BARRANDE, 1846</bibRefCitation>
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. The flange in the anterior part of each pleura is wide, concave (sag.) and more distinct than in
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<emphasis id="B9042A2AFFC4FFC9063AF88F078EF865" box="[106,395,1915,1939]" italics="true" pageId="5" pageNumber="166">P. paradoxissimus gracilis</emphasis>
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, with a prominent projection at the antero-lateral corner (
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<figureCitation id="134BEABDFFC4FFC90720F86807EBF844" box="[368,494,1947,1970]" captionStart="Text-fig" captionStartId="6.[130,214,1559,1580]" captionTargetBox="[129,1481,152,1512]" captionTargetId="figure-117@6.[129,1482,151,1512]" captionTargetPageId="6" captionText="Text-fig. 5. Thoracic structure, articulating and coaptative devices of H. minor (a, d, f–g), P. paradoxissimus gracilis (c, e, h–j) and A. cf. sacheri (b): a – specimen no. NM-L 46550, internal mold of pleurae; b – specimen no. NM-L 4776, internal mold of pleura; c – specimen no. NM-L 46551, internal mold of pleurae; d – specimen no. NM-L 46550, internal mold of axial rings and articulating half-rings; e – specimen no. NM-L 46552, internal mold of axial rings and articulating half-rings; f – specimen no. NM-L 46550, detail of pleural spine; g – specimen no. NM-L 41157, external mold of pleural spines; h – specimen no. NM-L 46547, external mold of pleural spines; i – specimen no. NM-L 46548, external mold of pleural spines; j – specimen no. NM-L 46549, pleural spines. Abbreviations: ahr – articulating half ring; alp – antero-lateral projection of flange; bp – backward projection at base of pleural spine; dtl – dorsal terrace lines; fl – flange; fu – fulcrum; ptl – petaloid-like terrace lines; vf – ventral furrow of pleural spine; vtl – ventral terrace lines." figureDoi="http://doi.org/10.5281/zenodo.4773118" httpUri="https://zenodo.org/record/4773118/files/figure.png" pageId="5" pageNumber="166">Text-fig. 5a</figureCitation>
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). Articulating half-rings are short (sag.), reaching ca. 40% of the length of axial rings (
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<figureCitation id="134BEABDFFC4FFC90621F82806F0F804" box="[113,245,2011,2034]" captionStart="Text-fig" captionStartId="6.[130,214,1559,1580]" captionTargetBox="[129,1481,152,1512]" captionTargetId="figure-117@6.[129,1482,151,1512]" captionTargetPageId="6" captionText="Text-fig. 5. Thoracic structure, articulating and coaptative devices of H. minor (a, d, f–g), P. paradoxissimus gracilis (c, e, h–j) and A. cf. sacheri (b): a – specimen no. NM-L 46550, internal mold of pleurae; b – specimen no. NM-L 4776, internal mold of pleura; c – specimen no. NM-L 46551, internal mold of pleurae; d – specimen no. NM-L 46550, internal mold of axial rings and articulating half-rings; e – specimen no. NM-L 46552, internal mold of axial rings and articulating half-rings; f – specimen no. NM-L 46550, detail of pleural spine; g – specimen no. NM-L 41157, external mold of pleural spines; h – specimen no. NM-L 46547, external mold of pleural spines; i – specimen no. NM-L 46548, external mold of pleural spines; j – specimen no. NM-L 46549, pleural spines. Abbreviations: ahr – articulating half ring; alp – antero-lateral projection of flange; bp – backward projection at base of pleural spine; dtl – dorsal terrace lines; fl – flange; fu – fulcrum; ptl – petaloid-like terrace lines; vf – ventral furrow of pleural spine; vtl – ventral terrace lines." figureDoi="http://doi.org/10.5281/zenodo.4773118" httpUri="https://zenodo.org/record/4773118/files/figure.png" pageId="5" pageNumber="166">Text-fig. 5d</figureCitation>
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), with the exception of the first one, which is slightly longer, similarly to
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<emphasis id="B9042A2AFFC4FFC907D8F80F07E9F7E5" box="[392,492,2044,2067]" italics="true" pageId="5" pageNumber="166">H. carens</emphasis>
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(see
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<bibRefCitation id="EFE18BC9FFC4FFC90471F80804DBF7E4" author="Whittington, H. B." box="[545,734,2043,2066]" pageId="5" pageNumber="166" pagination="27 - 46" refId="ref8229" refString="Whittington, H. B. (1990): Articulation and Exuviation in Cambrian Trilobites. - Philosophical Transactions of the Royal Society of London, Series B, 329: 27 - 46. https: // doi. org / 10.1098 / rstb. 1990.0147" type="journal article" year="1990">Whittington 1990</bibRefCitation>
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). Pleural spines are flat and claw-like, with a backward projection at their posterior bases (
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<figureCitation id="134BEABDFFC4FFC902FDFCCE0329FCA2" box="[1197,1324,829,852]" captionStart="Text-fig" captionStartId="6.[130,214,1559,1580]" captionTargetBox="[129,1481,152,1512]" captionTargetId="figure-117@6.[129,1482,151,1512]" captionTargetPageId="6" captionText="Text-fig. 5. Thoracic structure, articulating and coaptative devices of H. minor (a, d, f–g), P. paradoxissimus gracilis (c, e, h–j) and A. cf. sacheri (b): a – specimen no. NM-L 46550, internal mold of pleurae; b – specimen no. NM-L 4776, internal mold of pleura; c – specimen no. NM-L 46551, internal mold of pleurae; d – specimen no. NM-L 46550, internal mold of axial rings and articulating half-rings; e – specimen no. NM-L 46552, internal mold of axial rings and articulating half-rings; f – specimen no. NM-L 46550, detail of pleural spine; g – specimen no. NM-L 41157, external mold of pleural spines; h – specimen no. NM-L 46547, external mold of pleural spines; i – specimen no. NM-L 46548, external mold of pleural spines; j – specimen no. NM-L 46549, pleural spines. Abbreviations: ahr – articulating half ring; alp – antero-lateral projection of flange; bp – backward projection at base of pleural spine; dtl – dorsal terrace lines; fl – flange; fu – fulcrum; ptl – petaloid-like terrace lines; vf – ventral furrow of pleural spine; vtl – ventral terrace lines." figureDoi="http://doi.org/10.5281/zenodo.4773118" httpUri="https://zenodo.org/record/4773118/files/figure.png" pageId="5" pageNumber="166">Text-fig. 5f</figureCitation>
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). The dorsal surface of the pleural spines bears terrace lines, which are inclined in an angle of around 45 degrees to the sagittal axis of the exoskeleton (
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<figureCitation id="134BEABDFFC4FFC90243FC6E029EFC42" box="[1043,1179,925,948]" captionStart="Text-fig" captionStartId="6.[130,214,1559,1580]" captionTargetBox="[129,1481,152,1512]" captionTargetId="figure-117@6.[129,1482,151,1512]" captionTargetPageId="6" captionText="Text-fig. 5. Thoracic structure, articulating and coaptative devices of H. minor (a, d, f–g), P. paradoxissimus gracilis (c, e, h–j) and A. cf. sacheri (b): a – specimen no. NM-L 46550, internal mold of pleurae; b – specimen no. NM-L 4776, internal mold of pleura; c – specimen no. NM-L 46551, internal mold of pleurae; d – specimen no. NM-L 46550, internal mold of axial rings and articulating half-rings; e – specimen no. NM-L 46552, internal mold of axial rings and articulating half-rings; f – specimen no. NM-L 46550, detail of pleural spine; g – specimen no. NM-L 41157, external mold of pleural spines; h – specimen no. NM-L 46547, external mold of pleural spines; i – specimen no. NM-L 46548, external mold of pleural spines; j – specimen no. NM-L 46549, pleural spines. Abbreviations: ahr – articulating half ring; alp – antero-lateral projection of flange; bp – backward projection at base of pleural spine; dtl – dorsal terrace lines; fl – flange; fu – fulcrum; ptl – petaloid-like terrace lines; vf – ventral furrow of pleural spine; vtl – ventral terrace lines." figureDoi="http://doi.org/10.5281/zenodo.4773118" httpUri="https://zenodo.org/record/4773118/files/figure.png" pageId="5" pageNumber="166">
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Text-fig.
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). On the doublure of the pleural spines, the terrace lines are sub-parallel to the body axis, slightly turned adaxially in the posterior half of the spine (
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<figureCitation id="134BEABDFFC4FFC9056BFC0E05B0FBE2" box="[827,949,1021,1044]" captionStart="Text-fig" captionStartId="6.[130,214,1559,1580]" captionTargetBox="[129,1481,152,1512]" captionTargetId="figure-117@6.[129,1482,151,1512]" captionTargetPageId="6" captionText="Text-fig. 5. Thoracic structure, articulating and coaptative devices of H. minor (a, d, f–g), P. paradoxissimus gracilis (c, e, h–j) and A. cf. sacheri (b): a – specimen no. NM-L 46550, internal mold of pleurae; b – specimen no. NM-L 4776, internal mold of pleura; c – specimen no. NM-L 46551, internal mold of pleurae; d – specimen no. NM-L 46550, internal mold of axial rings and articulating half-rings; e – specimen no. NM-L 46552, internal mold of axial rings and articulating half-rings; f – specimen no. NM-L 46550, detail of pleural spine; g – specimen no. NM-L 41157, external mold of pleural spines; h – specimen no. NM-L 46547, external mold of pleural spines; i – specimen no. NM-L 46548, external mold of pleural spines; j – specimen no. NM-L 46549, pleural spines. Abbreviations: ahr – articulating half ring; alp – antero-lateral projection of flange; bp – backward projection at base of pleural spine; dtl – dorsal terrace lines; fl – flange; fu – fulcrum; ptl – petaloid-like terrace lines; vf – ventral furrow of pleural spine; vtl – ventral terrace lines." figureDoi="http://doi.org/10.5281/zenodo.4773118" httpUri="https://zenodo.org/record/4773118/files/figure.png" pageId="5" pageNumber="166">Text-fig. 5f</figureCitation>
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). The mutual orientation of the dorsal and ventral terrace lines forms a criss-cross pattern when the pleurae overlap.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |