treatments-xml/data/D9/00/05/D9000532704058FD9CFB0A7BC46BFC43.xml

<document ID-DOI="http://dx.doi.org/10.3897/jhr.95.96796" ID-Pensoft-Pub="1314-2607-95-143" ID-Pensoft-UUID="DE2EC0A557FC5F3D9AD2D4435824AA7B" ID-ZooBank="0AD4F90A9A41492D84C3C0AA1B8C275B" ModsDocID="1314-2607-95-143" checkinTime="1676724294143" checkinUser="pensoft" docAuthor="Praz, Christophe J. &amp; Benon, Dimitri" docDate="2023" docId="D9000532704058FD9CFB0A7BC46BFC43" docLanguage="en" docName="JourHymenoptRes 95: 143-198" docOrigin="Journal of Hymenoptera Research 95" docPubDate="2023-02-17" docSource="http://dx.doi.org/10.3897/jhr.95.96796" docTitle="Megachile argentata subsp. argentata argentata (Fabricius 1793" docType="treatment" docVersion="1" id="DE2EC0A557FC5F3D9AD2D4435824AA7B" lastPageNumber="143" masterDocId="DE2EC0A557FC5F3D9AD2D4435824AA7B" masterDocTitle="Revision of the leachella group of Megachile subgenus Eutricharaea in the Western Palaearctic (Hymenoptera, Apoidea, Megachilidae): A renewed plea for DNA barcoding type material" masterLastPageNumber="198" masterPageNumber="143" pageNumber="143" updateTime="1676724294143" updateUser="pensoft">
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<mods:title>Revision of the leachella group of Megachile subgenus Eutricharaea in the Western Palaearctic (Hymenoptera, Apoidea, Megachilidae): A renewed plea for DNA barcoding type material</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Praz, Christophe J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2649-3141</mods:nameIdentifier>
<mods:affiliation>University of Neuchatel, Neuchatel, Switzerland &amp; InfoFauna - Swiss Zoological Records Center, Neuchatel, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">christophe.praz@unine.ch</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Benon, Dimitri</mods:namePart>
<mods:affiliation>University of Neuchatel, Neuchatel, Switzerland</mods:affiliation>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-17</mods:number>
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<mods:number>95</mods:number>
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<mods:start>143</mods:start>
<mods:end>198</mods:end>
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<mods:url>http://dx.doi.org/10.3897/jhr.95.96796</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.95.96796</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:D9000532704058FD9CFB0A7BC46BFC43" httpUri="http://treatment.plazi.org/id/D9000532704058FD9CFB0A7BC46BFC43" lastPageNumber="143" pageId="0" pageNumber="143">
<subSubSection pageId="0" pageNumber="143" type="nomenclature">
<paragraph pageId="0" pageNumber="143">
<taxonomicName ID-CoL="8SFVF" LSID="D9000532-7040-58FD-9CFB-0A7BC46BFC43" authority="argentata (Fabricius, 1793)" authorityName="argentata (Fabricius" authorityYear="1793" baseAuthorityName="Fabricius" baseAuthorityYear="1793" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile argentata subsp. argentata" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="subSpecies" species="argentata" subSpecies="argentata">Megachile argentata argentata (Fabricius, 1793)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="reference_group">
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Fabricius" authorityYear="1793" class="Insecta" family="Megachilidae" genus="Apis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apis argentata" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="argentata">Apis argentata</taxonomicName>
Fabricius, 1793: 336 [sex not indicated], &quot;in Barbaria&quot; [Algeria or Tunisia]. Lectotype ♀ (NHMD), by designation of 
<bibRefCitation author="Hurd, PD" journalOrPublisher="Entomologiske Meddelelser" pageId="0" pageNumber="143" pagination="3 - 10" refId="B37" refString="Hurd, PD, 1967. The identity of Megachile rotundata (Fabricius) and M. argentata (Fabricius) (Hymenoptera: Apoidea). Entomologiske Meddelelser 35: 3 - 10" title="The identity of Megachile rotundata (Fabricius) and M. argentata (Fabricius) (Hymenoptera: Apoidea)." volume="35" year="1967">Hurd (1967)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Smith" authorityYear="1879" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile compacta" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="compacta">Megachile compacta</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
, 1895: 24, ♀, [Algeria]. Preoccupied, not 
<taxonomicName authorityName="Smith" authorityYear="1879" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile compacta" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="compacta">Megachile compacta</taxonomicName>
Smith, 1879. Lectotype ♀ (MNHN), by present designation (see below). New Synonymy.
</paragraph>
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Perez" authorityYear="1896" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile crassula" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="crassula">Megachile crassula</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
, 1896: 1. 
<emphasis italics="true" pageId="0" pageNumber="143">Nomen novum</emphasis>
for 
<taxonomicName family="Megachilidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="M. compacta" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="compacta">M. compacta</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
. New synonymy.
</paragraph>
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Ferton" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Perezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perezia maura" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="maura">Perezia maura</taxonomicName>
Ferton, 1914: 233, ♀ [gynandromorph specimen], &quot;Cimetière de Djidjelli&quot; [Jijel, Algeria]. Holotype intersex (MNHN). Preoccupied, not 
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile maura" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="maura">Megachile maura</taxonomicName>
Cresson, 1865. Synonymy with 
<taxonomicName family="Megachilidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="M. centuncularis" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="centuncularis">M. centuncularis</taxonomicName>
in 
<bibRefCitation DOI="https://doi.org/10.3406/bsef.1969.21096" author="Pasteels, JJ" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="0" pageNumber="143" pagination="248 - 248" refId="B53" refString="Pasteels, JJ, 1969. Perezia maura Ferton: une forme intersexuee de Megachile centuncularis L. [Hym. Apidae]. Bulletin de la Societe entomologique de France 74: 248 - 248, DOI: https://doi.org/10.3406/bsef.1969.21096" title="Perezia maura Ferton: une forme intersexuee de Megachile centuncularis L. [Hym. Apidae]." url="https://doi.org/10.3406/bsef.1969.21096" volume="74" year="1969">Pasteels (1969</bibRefCitation>
: 248). Synonymy with 
<taxonomicName family="Megachilidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="M. leachella" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="leachella">M. leachella</taxonomicName>
in 
<bibRefCitation author="van der Zanden, G" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="143" pagination="55 - 64" refId="B105" refString="van der Zanden, G, 1988. Nomenklatorische und taxonomische Bemerkungen zu einigen palaearktischen Arten der Familie Megachilidae (Insecta, Hymenoptera, Apoidea: Megachilidae). Reichenbachia 26: 55 - 64" title="Nomenklatorische und taxonomische Bemerkungen zu einigen palaearktischen Arten der Familie Megachilidae (Insecta, Hymenoptera, Apoidea: Megachilidae)." volume="26" year="1988">van der Zanden (1988</bibRefCitation>
: 56). New synonymy.
</paragraph>
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Alfken" authorityYear="1924" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile pilidens" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="pilidens">Megachile pilidens</taxonomicName>
Alfken, 1924: 88, ♀ ♂, &quot;Triest [Trieste, Italy]&quot;. Lectotype ♀ (ZMHB), by designation of 
<normalizedToken originalValue="Tkalců">Tkalcu</normalizedToken>
1967: 100. New synonymy.
</paragraph>
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Cockerell" authorityYear="1931" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile argyrea" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="argyrea">Megachile argyrea</taxonomicName>
Cockerell, 1931: 275, ♀ ♂, &quot;Asni [Morocco]&quot;. Holotype ♀ (USNM), paratypes ♀ ♂ (BMNH). New synonymy. Synonymy with 
<taxonomicName family="Megachilidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="M. pilidens" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="pilidens">M. pilidens</taxonomicName>
in Hedicke, 1933: 42.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="materials_examined">
<paragraph pageId="0" pageNumber="143">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" pageId="0" pageNumber="143">
Type material. 
<emphasis italics="true" pageId="0" pageNumber="143">
<typeStatus>Lectotype</typeStatus>
</emphasis>
</emphasis>
<specimenCount>♀</specimenCount>
of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata</emphasis>
</taxonomicName>
(NHMD) (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Vertex of the lectotype female of Megachile argentata (Fabricius, 1793), showing the ocelloccipital distance OOD (yellow line), two tangentials placed behind the lateral ocellus and along the inner margin of the compound eye (red lines), and the white rectangle with dimensions 1.1 x OOD x 0.55 x OOD within which the punctures were measured in our morphometric analyses." figureDoi="10.3897/jhr.95.96796.figure1" httpUri="https://binary.pensoft.net/fig/810381" pageId="0" pageNumber="143">1</figureCitation>
- 
<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Lectotype female of Megachile argentata 2 dorsal view 3 head in front view." figureDoi="10.3897/jhr.95.96796.figures2-3" httpUri="https://binary.pensoft.net/fig/811587" pageId="0" pageNumber="143">3</figureCitation>
; see above, identity of type specimens).
</paragraph>
<paragraph pageId="0" pageNumber="143">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Museum Paris Coll. J. Perez, Coll. J. Perez, C. Praz" country="Algeria" location="Algeria" specimenCount="♀" typeStatus="Lectotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="143">
<typeStatus>Lectotype</typeStatus>
</emphasis>
<specimenCount>♀</specimenCount>
of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. crassula" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="crassula">
<emphasis italics="true" pageId="0" pageNumber="143">M. crassula</emphasis>
</taxonomicName>
(MNHN), by present designation, a female in good condition labeled as follows: 1. 
<normalizedToken originalValue="“Algeria”">
&quot; 
<collectingCountry name="Algeria">Algeria</collectingCountry>
&quot;
</normalizedToken>
[printed]; 2. 
<taxonomicName authorityName="Perez" authorityYear="1896" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile crassula" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="crassula">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile crassula</emphasis>
</taxonomicName>
JP [handwritten, presumably by 
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
]; 3. &quot; 
<collectorName>
Museum Paris 
<collectorName>
Coll. J. 
<normalizedToken originalValue="Pérez”">Perez&quot;</normalizedToken>
</collectorName>
</collectorName>
[printed on blue paper]; 4. &quot; 
<typeStatus>Holotype</typeStatus>
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. crassula" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="crassula">
<emphasis italics="true" pageId="0" pageNumber="143">M. crassula</emphasis>
</taxonomicName>
des. Baker 1991&quot; [printed and handwritten on red paper]; 5. &quot; 
<typeStatus>Lectotype</typeStatus>
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. crassula" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="crassula">
<emphasis italics="true" pageId="0" pageNumber="143">M. crassula</emphasis>
</taxonomicName>
des. 
<collectorName>C. Praz</collectorName>
2022&quot;. Although the original description does not mention the number of specimens, we prefer to designate this specimen as the 
<typeStatus>lectotype</typeStatus>
.
</materialsCitation>
</paragraph>
<caption doi="10.3897/jhr.95.96796.figures15-24" httpUri="https://binary.pensoft.net/fig/810386" pageId="0" pageNumber="143" start="Figures 15–24" startId="F7">
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" pageId="0" pageNumber="143">Figures 15-24.</emphasis>
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile argentata" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile argentata</emphasis>
</taxonomicName>
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="143">15</emphasis>
female
</specimenCount>
metasoma of northwestern African populations, 
<taxonomicName authority="argentata" family="Megachilidae" kingdom="Animalia" lsidName="M. argentata subsp. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="argentata" subSpecies="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata argentata</emphasis>
</taxonomicName>
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="143">16</emphasis>
female
</specimenCount>
metasoma of European, Levant, 
<collectingCountry name="Turkey">Turkey</collectingCountry>
and Central Asian populations, 
<taxonomicName authority="argentata" family="Megachilidae" kingdom="Animalia" lsidName="M. argentata subsp. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="argentata" subSpecies="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata argentata</emphasis>
</taxonomicName>
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="143">17</emphasis>
female
</specimenCount>
metasoma of 
<collectingCountry name="Malta">Malta</collectingCountry>
populations, 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata subsp. schmiedeknechti" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="argentata" subSpecies="schmiedeknechti">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata schmiedeknechti</emphasis>
</taxonomicName>
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="143">18</emphasis>
female
</specimenCount>
metasoma of Corsican and Sardinian populations, 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata subsp. schmiedeknechti" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="argentata" subSpecies="schmiedeknechti">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata schmiedeknechti</emphasis>
</taxonomicName>
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="143">19</emphasis>
female
</specimenCount>
metasomal tergum 4 
<specimenCount type="female">
<emphasis bold="true" pageId="0" pageNumber="143">20</emphasis>
female
</specimenCount>
vertex 
<emphasis bold="true" pageId="0" pageNumber="143">21</emphasis>
apex of female clypeus 
<specimenCount type="male">
<emphasis bold="true" pageId="0" pageNumber="143">22</emphasis>
male
</specimenCount>
front tarsi 
<specimenCount type="male">
<emphasis bold="true" pageId="0" pageNumber="143">23</emphasis>
male
</specimenCount>
metasomal sterna 3-5 
<specimenCount type="male">
<emphasis bold="true" pageId="0" pageNumber="143">24</emphasis>
male
</specimenCount>
genitalia.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" italics="true" pageId="0" pageNumber="143">
<typeStatus>Holotype</typeStatus>
</emphasis>
[gynandromorph] of 
<taxonomicName authorityName="Ferton" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Perezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perezia maura" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="maura">
<emphasis italics="true" pageId="0" pageNumber="143">Perezia maura</emphasis>
</taxonomicName>
(MNHN), labeled as follows: 1. &quot;Djidjelli 17-8/ 
<specimenCount type="generic">13 p.</specimenCount>
315&quot; [handwritten by Ferton]; 2. &quot; 
<taxonomicName authorityName="Ferton" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Perezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perezia maura" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="maura">
<emphasis italics="true" pageId="0" pageNumber="143">Perezia maura</emphasis>
</taxonomicName>
<specimenCount>♀</specimenCount>
Fert&quot; [handwritten by Ferton]; 3. 
<typeStatus>Holotype</typeStatus>
[printed, red]; 4. 
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
Paris Ch. Ferton 1902 [printed, blue paper]; 5. 
<taxonomicName authorityName="Alfken" authorityYear="1924" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile pilidens" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="pilidens">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile pilidens</emphasis>
</taxonomicName>
Alfken [symbol for gynandromorph] det. G.v.d. Zanden 1986; 6. &quot; 
<taxonomicName authorityName="Ferton" authorityYear="1914" class="Insecta" family="Megachilidae" genus="Perezia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perezia maura" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="maura">
<emphasis italics="true" pageId="0" pageNumber="143">Perezia maura</emphasis>
</taxonomicName>
Ferton = gynandromorph de 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile centuncularis" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="centuncularis">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile centuncularis</emphasis>
</taxonomicName>
J. Pasteels det., 1969&quot;; 7. 
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile argentata" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile argentata</emphasis>
</taxonomicName>
intersex det C. Praz 2022. It is unclear why 
<bibRefCitation author="van der Zanden, G" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="143" pagination="55 - 64" refId="B105" refString="van der Zanden, G, 1988. Nomenklatorische und taxonomische Bemerkungen zu einigen palaearktischen Arten der Familie Megachilidae (Insecta, Hymenoptera, Apoidea: Megachilidae). Reichenbachia 26: 55 - 64" title="Nomenklatorische und taxonomische Bemerkungen zu einigen palaearktischen Arten der Familie Megachilidae (Insecta, Hymenoptera, Apoidea: Megachilidae)." volume="26" year="1988">van der Zanden (1988)</bibRefCitation>
placed this taxon in synonymy with 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. leachella" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="leachella">
<emphasis italics="true" pageId="0" pageNumber="143">M. leachella</emphasis>
</taxonomicName>
since the label on the specimen suggests he correctly identified the 
<typeStatus>holotype</typeStatus>
as 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. pilidens" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="pilidens">
<emphasis italics="true" pageId="0" pageNumber="143">M. pilidens</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" italics="true" pageId="0" pageNumber="143">
<typeStatus>Lectotype</typeStatus>
</emphasis>
<specimenCount>♀</specimenCount>
of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. pilidens" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="pilidens">
<emphasis italics="true" pageId="0" pageNumber="143">M. pilidens</emphasis>
</taxonomicName>
(ZMHB). A female without metasoma, but clearly identified as 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata</emphasis>
</taxonomicName>
(see 
<normalizedToken originalValue="Tkalců">Tkalcu</normalizedToken>
1967).
</paragraph>
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" italics="true" pageId="0" pageNumber="143">
<typeStatus>Paratypes</typeStatus>
</emphasis>
<specimenCount>♀</specimenCount>
and 
<specimenCount>♂</specimenCount>
of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argyrea" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argyrea">
<emphasis italics="true" pageId="0" pageNumber="143">M. argyrea</emphasis>
</taxonomicName>
(BMNH), all in good condition and clearly corresponding to 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata</emphasis>
</taxonomicName>
. We did not examine the 
<typeStatus>holotype</typeStatus>
<specimenCount>♀</specimenCount>
of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argyrea" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argyrea">
<emphasis italics="true" pageId="0" pageNumber="143">M. argyrea</emphasis>
</taxonomicName>
, but pictures are available at https://collections.nmnh.si.edu/search/ento/; based on these pictures, the 
<typeStatus>holotype</typeStatus>
agrees with 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata</emphasis>
</taxonomicName>
in the dark vestiture of the lateral parts of T5 (absent in 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. leachella subsp. albipila" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="leachella" subSpecies="albipila">
<emphasis italics="true" pageId="0" pageNumber="143">M. leachella albipila</emphasis>
</taxonomicName>
and 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. inexspectata" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="inexspectata">
<emphasis italics="true" pageId="0" pageNumber="143">M. inexspectata</emphasis>
</taxonomicName>
). Punctation of the vertex is not visible but the dense punctation of the terga appears to exclude 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. leachella subsp. albipila" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="leachella" subSpecies="albipila">
<emphasis italics="true" pageId="0" pageNumber="143">M. leachella albipila</emphasis>
</taxonomicName>
. This character and the identity of examined male and female 
<typeStatus>paratypes</typeStatus>
(BMNH) collected in the same site confirm synonymy with 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata." order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argentata.">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata.</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="notes">
<paragraph pageId="0" pageNumber="143">Note.</paragraph>
<paragraph pageId="0" pageNumber="143">
<taxonomicName authorityName="Benoist" authorityYear="1951" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile beaumonti" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile beaumonti</emphasis>
</taxonomicName>
Benoist, 1951, is currently treated as a synonym of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. crassula" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="crassula">
<emphasis italics="true" pageId="0" pageNumber="143">M. crassula</emphasis>
</taxonomicName>
(
<bibRefCitation author="van der Zanden, G" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="143" pagination="51 - 54" refId="B106" refString="van der Zanden, G, 1990. Nomenklatorische Aenderungen fuer einige palaearktische Arten der Familie Megachilidae (Insecta, Hymenoptera, Apoidea). Reichenbachia 28: 51 - 54" title="Nomenklatorische Aenderungen fuer einige palaearktische Arten der Familie Megachilidae (Insecta, Hymenoptera, Apoidea)." volume="28" year="1990">van der Zanden 1990</bibRefCitation>
: 53). We have examined the holotype ♀ of this taxon (MZL); 
<taxonomicName authorityName="Benoist" authorityYear="1951" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile beaumonti" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile beaumonti</emphasis>
</taxonomicName>
is not conspecific with 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="argentata">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata</emphasis>
</taxonomicName>
, but is a valid species of the 
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile rotundata" order="Hymenoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="rotundata">
<emphasis italics="true" pageId="0" pageNumber="143">Megachile rotundata</emphasis>
</taxonomicName>
group (stat. rev.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="materials_examined">
<paragraph pageId="0" pageNumber="143">Other material.</paragraph>
<paragraph pageId="0" pageNumber="143">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Croatia, France, Germany, Greece, Iran, Italy, Kyrgyzstan, Lebanon" location="Northern Macedonia" specimenCount="216">
<specimenCount type="generic">216 specimens</specimenCount>
from the following countries: 
<collectingCountry isEnumeration="true">Croatia, France, Germany, Greece, Iran, Italy, Kyrgyzstan, Lebanon</collectingCountry>
, 
<collectingCountry name="Morocco">Morocco</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:D9000532704058FD9CFB0A7BC46BFC43:F6EFA67B931F2A9897368D242316DAF2" country="Croatia, France, Germany, Greece, Iran, Italy, Kyrgyzstan, Lebanon" name="Northern Macedonia">Northern Macedonia</location>
, 
<collectingCountry isEnumeration="true">Portugal, Russia, Serbia, Spain, Switzerland, Tunisia, Turkey</collectingCountry>
, 
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
(Suppl. material 1)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="distribution">
<paragraph pageId="0" pageNumber="143">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="143">
Widespread and abundant in southern Europe; in expansion in central and northern Europe, e.g., in northern Switzerland (C. Praz et al., in prep.), the Netherlands (
<bibRefCitation author="Peeters, TMJ" journalOrPublisher="Nederlandse Faunistische Mededelingen" pageId="0" pageNumber="143" pagination="11 - 18" refId="B54" refString="Peeters, TMJ, Raemakers, I, van der Nieuwegiessen, J, Kuper, JT, 2006. De rotsbehangersbij Megachile pilidens, nieuw voor de Nederlandse fauna (Hymenoptera: Apoidea: Megachilidae). Nederlandse Faunistische Mededelingen 25: 11 - 18" title="De rotsbehangersbij Megachile pilidens, nieuw voor de Nederlandse fauna (Hymenoptera: Apoidea: Megachilidae)." volume="25" year="2006">Peeters et al. 2006</bibRefCitation>
) and Germany (LUBW 2007; 
<bibRefCitation author="Schweitzer, L" journalOrPublisher="Linzer Biologische Beitraege" pageId="0" pageNumber="143" refId="B84" refString="Schweitzer, L, Theunert, R, 2019. Zum Vorkommen von Halictus scabiosae (Rossi, 1790) und Megachile pilidens (Alfken, 1924) in Niedersachsen (Hymenoptera: Apidae). Peiner Biologische Arbeitsgemeinschaft - Online 2019-01: 1-9." title="Zum Vorkommen von Halictus scabiosae (Rossi, 1790) und Megachile pilidens (Alfken, 1924) in Niedersachsen (Hymenoptera: Apidae). Peiner Biologische Arbeitsgemeinschaft - Online 2019 - 01: 1 - 9." year="2019">Schweitzer and Theunert 2019</bibRefCitation>
). Present in the Levant (Turkey, Lebanon), but so far its presence has not been confirmed in Israel and Egypt (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5032.3.1" author="Praz, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="143" pagination="301 - 330" refId="B64" refString="Praz, C, Al-Shahat, AM, Gadallah, NS, 2021. Taxonomic revision of the subgenus Eutricharaea Thomson in Egypt, with a key to the species and the description of two new species (Hymenoptera, Anthophila, Megachilidae, genus Megachile Latreille). Zootaxa 5032: 301 - 330, DOI: https://doi.org/10.11646/zootaxa.5032.3.1" title="Taxonomic revision of the subgenus Eutricharaea Thomson in Egypt, with a key to the species and the description of two new species (Hymenoptera, Anthophila, Megachilidae, genus Megachile Latreille)." url="https://doi.org/10.11646/zootaxa.5032.3.1" volume="5032" year="2021">Praz et al. 2021</bibRefCitation>
). We have examined specimens from Rhodos, Lesvos, Samos, Karpathos, Patmos, Chios, but neither from Cyprus [the record of 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. pilidens" order="Hymenoptera" pageId="0" pageNumber="143" rank="species" species="pilidens">
<emphasis italics="true" pageId="0" pageNumber="143">M. pilidens</emphasis>
</taxonomicName>
by 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.924.38328" author="Varnava, AI" journalOrPublisher="ZooKeys" pageId="0" pageNumber="143" pagination="1 - 114" refId="B95" refString="Varnava, AI, Roberts, SPM, Michez, D, Ascher, JS, Petanidou, T, Dimitriou, S, Devalez, J, Pittara, M, Stavrinides, MC, 2020. The wild bees (Hymenoptera, Apoidea) of the island of Cyprus. ZooKeys 924: 1 - 114, DOI: https://doi.org/10.3897/zookeys.924.38328" title="The wild bees (Hymenoptera, Apoidea) of the island of Cyprus." url="https://doi.org/10.3897/zookeys.924.38328" volume="924" year="2020">Varnava et al. (2020)</bibRefCitation>
is probably an identification error and likely refers to 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. leachella subsp. densipunctata" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="leachella" subSpecies="densipunctata">
<emphasis italics="true" pageId="0" pageNumber="143">M. leachella densipunctata</emphasis>
</taxonomicName>
ssp. nov.] nor from Crete. Further east, the species is present in Iran, Armenia and Kyrgyzstan.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="geographic variation">
<paragraph pageId="0" pageNumber="143">Geographic variation.</paragraph>
<paragraph pageId="0" pageNumber="143">
Geographic variation has been discussed above in detail. The northwestern African populations, as well as populations from the Island of Pantelleria, have overall snow-white vestiture in the female sex (Fig. 
<figureCitation captionStart="Figures 15–24" captionStartId="F7" captionText="Figures 15 - 24. Megachile argentata 15 female metasoma of northwestern African populations, M. argentata argentata 16 female metasoma of European, Levant, Turkey and Central Asian populations, M. argentata argentata 17 female metasoma of Malta populations, M. argentata schmiedeknechti 18 female metasoma of Corsican and Sardinian populations, M. argentata schmiedeknechti 19 female metasomal tergum 4 20 female vertex 21 apex of female clypeus 22 male front tarsi 23 male metasomal sterna 3 - 5 24 male genitalia." figureDoi="10.3897/jhr.95.96796.figures15-24" httpUri="https://binary.pensoft.net/fig/810386" pageId="0" pageNumber="143">15</figureCitation>
) and dark vestiture on the mesonotum and vertex, and in the male sex a continuous fringe of hairs on the margin of T6. On the Island of Karpathos, the female scopa is orange on S5 and S6, approaching the condition observed in 
<taxonomicName family="Megachilidae" kingdom="Animalia" lsidName="M. argentata subsp. schmiedeknechti" order="Hymenoptera" pageId="0" pageNumber="143" rank="subSpecies" species="argentata" subSpecies="schmiedeknechti">
<emphasis italics="true" pageId="0" pageNumber="143">M. argentata schmiedeknechti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>