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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="8A8901D5441CE154B3818344035A592F" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Dasiosoma Britton 1937" docType="treatment" docVersion="5" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="64" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="62" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>284</mods:number>
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<mods:start>1</mods:start>
<mods:end>129</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
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<treatment ID-GBIF-Taxon="152042248" LSID="urn:lsid:plazi:treatment:8A8901D5441CE154B3818344035A592F" httpUri="http://treatment.plazi.org/id/8A8901D5441CE154B3818344035A592F" lastPageId="64" lastPageNumber="64" pageId="61" pageNumber="62">
<subSubSection pageId="61" pageNumber="62" type="nomenclature">
<paragraph pageId="61" pageNumber="62">
Genus
<taxonomicName LSID="8A8901D5-441C-E154-B381-8344035A592F" authority="Britton, 1937" authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Dasiosoma" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">Dasiosoma Britton, 1937</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="reference_group">
<paragraph pageId="61" pageNumber="62">
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">Dasiosoma</taxonomicName>
<bibRefCitation author="Britton, EB" journalOrPublisher="Proceedings of the Royal Entomological Society of London, Series B" pageId="86" pageNumber="87" pagination="233 - 237" refId="B30" refString="Britton, EB, 1937. New African Carabidae (Col.) including the description of a new genus. Proceedings of the Royal Entomological Society of London, Series B 6: 233 - 237" title="New African Carabidae (Col.) including the description of a new genus." volume="6" year="1937">Britton 1937</bibRefCitation>
: 233;
<bibRefCitation author="Basilewsky, P" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="85" pageNumber="86" pagination="323 - 342" refId="B21" refString="Basilewsky, P, 1949. Coleopteres Carabidae africains nouveaux. II, III. Revue de Zoologie et de Botanique Africaines 41: 323 - 342" title="Coleopteres Carabidae africains nouveaux. II, III." volume="41" year="1949">Basilewsky 1949</bibRefCitation>
: 221 (key to species);
<bibRefCitation author="Basilewsky, P" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="85" pageNumber="86" pagination="142 - 147" refId="B22" refString="Basilewsky, P, 1968. Descriptions de trois carabides nouveaux de Cote d'Ivoire. (Coleoptera Carabidae). Revue de Zoologie et de Botanique Africaines 77: 142 - 147" title="Descriptions de trois carabides nouveaux de Cote d'Ivoire. (Coleoptera Carabidae)." volume="77" year="1968">Basilewsky 1968</bibRefCitation>
: 145.
</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">Teradaia</taxonomicName>
<bibRefCitation author="Habu, A" journalOrPublisher="Proceedings of the Japanese Society of Systematic Zoology" pageId="87" pageNumber="88" pagination="65 - 69" refId="B49" refString="Habu, A, 1979a. A new genus and species of the Lebiini from Formosa (Coleoptera, Carabidae). Proceedings of the Japanese Society of Systematic Zoology 16: 65 - 69" title="A new genus and species of the Lebiini from Formosa (Coleoptera, Carabidae)." volume="16" year="1979 a">Habu 1979a</bibRefCitation>
: 65. Type-species:
<taxonomicName authorityName="Habu" authorityYear="1979" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="61" pageNumber="62">Teradaia bella</emphasis>
</taxonomicName>
Habu, 1979a, by original designation.
<emphasis bold="true" pageId="61" pageNumber="62">Syn. n.</emphasis>
[Synonym]
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="62" type="type-species">
<paragraph pageId="61" pageNumber="62">Type-species:</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="testaceum">
<emphasis italics="true" pageId="61" pageNumber="62">Dasiosoma testaceum</emphasis>
</taxonomicName>
Britton, 1937, by original designation.
</paragraph>
</subSubSection>
<subSubSection lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="diagnosis">
<paragraph pageId="61" pageNumber="62">Diagnosis.</paragraph>
<paragraph pageId="61" pageNumber="62">Dorsal side evenly pubescent, elytra with accessory setae on all intervals; labrum and outer scrobe of mandibles nearly glabrous; pronotum with basal foveae deep and narrow, forming a pair of deep grooves; elytral striae distinct, finely punctate; aedeagal internal sac with main flagellum developed, apex reaching the apical orifice, not projecting out from it.</paragraph>
<paragraph lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
Some species of the genus were placed in
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="61" pageNumber="62">Lachnoderma</emphasis>
</taxonomicName>
by mistake (
<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="85" pageNumber="86" pagination="205 - 290" refId="B25" refString="Bates, HW, 1883. Supplement to the Geodephagous Coleoptera of Japan, chiefly from the collection of George Lewis, made during his second visit, from February, 1880, to September, 1881. The Transactions of the Entomological Society of London 1883: 205 - 290" title="Supplement to the Geodephagous Coleoptera of Japan, chiefly from the collection of George Lewis, made during his second visit, from February, 1880, to September, 1881." volume="1883" year="1883">Bates 1883</bibRefCitation>
;
<bibRefCitation author="Tian, M" journalOrPublisher="Nouvelle Revue d'Entomologie" pageId="89" pageNumber="90" pagination="123 - 133" refId="B84" refString="Tian, M, Deuve, T, 2001. Notes on the genus Lachnoderma Macleay, 1873, with descriptions of six new species from China and Vietnam (Coleoptera, Carabidae). Nouvelle Revue d'Entomologie 18 (2): 123 - 133" title="Notes on the genus Lachnoderma Macleay, 1873, with descriptions of six new species from China and Vietnam (Coleoptera, Carabidae)." volume="18" year="2001">Tian and Deuve 2001</bibRefCitation>
). These two genera may be closely allied, but
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="61" pageNumber="62">Dasiosoma</emphasis>
</taxonomicName>
can be distinguished in having: (1) labrum and outer scrobe of mandibles nearly glabrous, at most labrum with a few very short fine accessory setae, but in
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="61" pageNumber="62">Lachnoderma</emphasis>
</taxonomicName>
, labrum and mandibles always with long accessory setae, accessory setae on labrum as long as the six primary setae near apex; (2) mentum tooth simple, but bifid in
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="61" pageNumber="62">Lachnoderma</emphasis>
</taxonomicName>
;
<pageBreakToken pageId="62" pageNumber="63" start="start">(</pageBreakToken>
3) pronotum basal foveae much deeper and groove-like, but shallower and wide in
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Lachnoderma</emphasis>
</taxonomicName>
; (4) punctures in striae much finer than in
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Lachnoderma</emphasis>
</taxonomicName>
; (5) median lobe of aedeagus slender, apical bursa present, main flagellum not projected out from apical orifice, but
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="62" pageNumber="63">Lachnoderma</emphasis>
</taxonomicName>
with median lobe of aedeagus stout, apical bursa absent, main flagellum projected out from apical orifice.
</paragraph>
</subSubSection>
<subSubSection lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="generic characters">
<paragraph pageId="62" pageNumber="63">Generic characters.</paragraph>
<paragraph lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
Dorsal side yellow, brown or piceous, elytra unicolored or bicolored, usually with weak metallic luster.
<emphasis bold="true" pageId="62" pageNumber="63">Head</emphasis>
densely pubescent, occiput and central area of clypeus nearly glabrous; eyes hemispherical, strongly prominent; tempora shorter or slightly longer than half length of eyes, abruptly or gradually narrowed behind eyes; vertex slightly or strongly tumid. Antennae extended to basal one-fifth of elytra; 1st antennomere slightly curved, 3rd slightly longer than 4th. Labrum slightly widened to apex, anterior margin straight or slightly curved, with faint microsculpture, glabrous or with a few very fine and short secondary setae; mandibles distinctly widened, outer margins rounded, glabrous on outer scrobe, with some setae arranged along dorsal ridge; terminal maxillary palpomeres fusiform in both sexes; terminal labial palpomeres strongly securiform, truncate apically in males, less widened in females; ligula with apex slightly projected, with four long setae and some short setae; paraglossae membranous, not longer than ligula, narrow, adnate; mentum tooth simple, short and rounded, with two long setae near base, sometimes with a few additional short setae; submentum with two long setae; genae with long setae beneath eyes; gula glabrous except apex.
<emphasis bold="true" pageId="62" pageNumber="63">Pronotum</emphasis>
wider than head; disc, lateral explanate areas and lateral margins densely and evenly pubescent; mid-lateral primary setae present, distinctly longer than accessory ones; basal foveae deep, forming a pair of deep grooves, extended anteriorly or anteriomedially; median line deep, reaching apical and basal margins; pronotal base more or less lobed; lateral margins rounded or narrowed in middle, more or less sinuate before hind angles; hind angles sharp, rectangular or acute, projected or not.
<emphasis bold="true" pageId="62" pageNumber="63">Elytra</emphasis>
slightly narrow, lateral margins parallel or expanded to apex; apex truncate, sutural angles not projected, outer angles completely rounded; basal margination reaching 3rd interval; basal pores large; striae shallow, distinct, finely punctate; intervals slightly convex; all intervals evenly pubescent, pubescence fine and long, erect; primary setigerous pores small, indistinct, with setae longer than erect pubescence, three or four pores on 3rd interval, one or two pores on base of 5th; 7th and 8th intervals slightly tumid near apex.
<emphasis bold="true" pageId="62" pageNumber="63">Ventral side</emphasis>
with long dense pubescence except posterior area of proepisterna, sparser on mesosternum and central area of metasternum; males with apex of terminal sternum moderately emarginate, with two pairs of setae, females straight or slightly emarginate, with two pairs of setae.
<emphasis bold="true" pageId="62" pageNumber="63">Legs</emphasis>
short; protibiae with cleaning spur well developed, distant from inner margin; tarsi widened, 4th tarsomere bifid, claws pectinate; males with adhesive hairs absent on all tarsomeres.
<emphasis bold="true" pageId="62" pageNumber="63">Male genitalia</emphasis>
with median lobe of aedeagus not twisted, strongly expanded and bent at base; strongly bent to right side in dorsal view; apical orifice opened apically; dorsal surface with a few fine setae subapically; internal sac with main flagellum slender, apex nearly reaching apical orifice, not projected out from it, trumpet-form expansion small; apical bursa present; secondary flagellum distinct.
<emphasis bold="true" pageId="62" pageNumber="63">Female genitalia</emphasis>
. Spermatheca tubular, with ring-sculpture, inserted on bursa copulatrix; spermathecal
<pageBreakToken pageId="63" pageNumber="64" start="start">gland</pageBreakToken>
very short or longer than spermatheca, inserted near middle of spermatheca; spermatheca not bent. Apical segment of ovipositor scimitar-shaped, curved to outer side; apex sharp; apex with membranous extension short but sharp.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="distribution">
<paragraph pageId="63" pageNumber="64">Distribution.</paragraph>
<paragraph pageId="63" pageNumber="64">South China, Indo-China Peninsula, South Asia, African mainland. Not discovered in the Philippines, Malay Archipelago or Madagascar.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="monophyly and relationships">
<paragraph pageId="63" pageNumber="64">Monophyly and relationships.</paragraph>
<paragraph pageId="63" pageNumber="64">
Relationship between
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Lachnoderma</emphasis>
</taxonomicName>
is discussed under
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Lachnoderma</emphasis>
</taxonomicName>
. Monophyly of
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
is suggested by the following apomorphic character states: (1) basal foveae of pronotum deep and narrow; (2) median lobe of aedeagus strongly expanded and bent at base.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="taxonomic comments">
<paragraph pageId="63" pageNumber="64">Taxonomic comments.</paragraph>
<paragraph pageId="63" pageNumber="64">
<bibRefCitation author="Habu, A" journalOrPublisher="Proceedings of the Japanese Society of Systematic Zoology" pageId="87" pageNumber="88" pagination="65 - 69" refId="B49" refString="Habu, A, 1979a. A new genus and species of the Lebiini from Formosa (Coleoptera, Carabidae). Proceedings of the Japanese Society of Systematic Zoology 16: 65 - 69" title="A new genus and species of the Lebiini from Formosa (Coleoptera, Carabidae)." volume="16" year="1979 a">Habu (1979a)</bibRefCitation>
described the genus
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Teradaia</emphasis>
</taxonomicName>
for his new species
<taxonomicName authorityName="Habu" authorityYear="1979" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="63" pageNumber="64">Teradaia bella</emphasis>
</taxonomicName>
, and indicated that it is allied with
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Lachnoderma</emphasis>
</taxonomicName>
. But the Palaearctic catalogue (
<bibRefCitation author="Loebl, I" journalOrPublisher="Linzer Biologische Beitraege" pageId="88" pageNumber="89" refId="B65" refString="Loebl, I, Smetana, A, 2003. Apollo Books, Denmark, 819 pp." title="Apollo Books, Denmark, 819 pp." year="2003">
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2003
</bibRefCitation>
) included
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Teradaia</emphasis>
</taxonomicName>
in
<taxonomicName authorityName="Hope" authorityYear="1838" class="Insecta" family="Carabidae" genus="Pericalina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Pericalina</taxonomicName>
without explanation. We examined types of all four species from Africa belonging to
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
in the former concept, and dissected a paratype of
<taxonomicName authorityName="Basilewsky" authorityYear="1968" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ivorense">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma ivorense</emphasis>
</taxonomicName>
. The male genitalia (
<figureCitation captionStart="Figure 86" captionStartId="F31" captionText="Figure 86. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire." httpUri="https://binary.pensoft.net/fig/29704" pageId="63" pageNumber="64">Fig. 86</figureCitation>
) are not significantly different from those of
<taxonomicName authorityName="Habu" authorityYear="1979" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="63" pageNumber="64">Teradaia bella</emphasis>
</taxonomicName>
and its allied species (
<figureCitation captionStart="Figure 87" captionStartId="F32" captionText="Figure 87. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Dasiosoma bellum (Habu), a specimen from Vietnam." httpUri="https://binary.pensoft.net/fig/29705" pageId="63" pageNumber="64">Figs 87</figureCitation>
<figureCitation captionStart="Figure 90" captionStartId="F35" captionText="Figure 90. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Dasiosoma quadraticolle sp. n., holotype." httpUri="https://binary.pensoft.net/fig/29708" pageId="63" pageNumber="64">-90</figureCitation>
), and the small differences of external morphological characters between species from the two continents (see the key, first item) are not enough to be considered of genericlevel difference. So we herein synonymize
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Teradaia</emphasis>
</taxonomicName>
with
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
, and include
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
in
<taxonomicName genus="Physoderina" lsidName="" pageId="63" pageNumber="64" rank="genus">Physoderina</taxonomicName>
, not
<taxonomicName authorityName="Hope" authorityYear="1838" class="Insecta" family="Carabidae" genus="Pericalina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Pericalina</taxonomicName>
.
</paragraph>
<paragraph pageId="63" pageNumber="64">
Moreover, two Asian species (
<taxonomicName authorityName="Bates" authorityYear="1873" class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis italics="true" pageId="63" pageNumber="64">Singilis hirsutus</emphasis>
</taxonomicName>
Bates and
<taxonomicName authorityName="Tian &amp; Deuve" authorityYear="2001" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis italics="true" pageId="63" pageNumber="64">Lachnoderma maindroni</emphasis>
</taxonomicName>
Tian &amp; Deuve) that used to be placed in
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Lachnoderma</emphasis>
</taxonomicName>
, and also
<taxonomicName class="Insecta" family="Carabidae" genus="Diamella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="indica">
<emphasis italics="true" pageId="63" pageNumber="64">Diamella indica</emphasis>
</taxonomicName>
Kirschenhofer, are closer to
<taxonomicName authorityName="Habu" authorityYear="1979" class="Insecta" family="Carabidae" genus="Teradaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="bella">
<emphasis italics="true" pageId="63" pageNumber="64">Teradaia bella</emphasis>
</taxonomicName>
Habu than the other species of
<taxonomicName authorityName="W. J. Macleay" authorityYear="1873" class="Insecta" family="Carabidae" genus="Lachnoderma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Lachnoderma</emphasis>
</taxonomicName>
or
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Diamella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Diamella</emphasis>
</taxonomicName>
. So we combine these three species with
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
and describe a new species from Asia in the present paper. Thus the current concept of
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
includes nine species, with four of them from Africa, and the other five from Asia. The definitive characters of
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
are in the diagnosis.
</paragraph>
<paragraph pageId="63" pageNumber="64">
The four Africa species (
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="testaceum">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma testaceum</emphasis>
</taxonomicName>
Britton, 1937,
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma basilewskyi</emphasis>
</taxonomicName>
Shi &amp; Liang nom. n.,
<taxonomicName authorityName="Basilewsky" authorityYear="1949" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="sudanicum">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma sudanicum</emphasis>
</taxonomicName>
Basilewsky, 1949 and
<taxonomicName authorityName="Basilewsky" authorityYear="1968" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ivorense">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma ivorense</emphasis>
</taxonomicName>
Basilewsky, 1968) are very similar to each other. Two of them were described based each on a single female.
<bibRefCitation author="Basilewsky, P" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="85" pageNumber="86" pagination="323 - 342" refId="B21" refString="Basilewsky, P, 1949. Coleopteres Carabidae africains nouveaux. II, III. Revue de Zoologie et de Botanique Africaines 41: 323 - 342" title="Coleopteres Carabidae africains nouveaux. II, III." volume="41" year="1949">Basilewsky (1949</bibRefCitation>
,
<bibRefCitation author="Basilewsky, P" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="85" pageNumber="86" pagination="142 - 147" refId="B22" refString="Basilewsky, P, 1968. Descriptions de trois carabides nouveaux de Cote d'Ivoire. (Coleoptera Carabidae). Revue de Zoologie et de Botanique Africaines 77: 142 - 147" title="Descriptions de trois carabides nouveaux de Cote d'Ivoire. (Coleoptera Carabidae)." volume="77" year="1968">1968</bibRefCitation>
) provided a key and diagnosis to these four species, but when we examined types of all four of these species, it is still difficult for us to clearly distinguish them all. So we leave this part of review work for future study when more material becomes available. In the present paper, we just revise the Asian fauna of
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="key to species of dasiosoma britton">
<paragraph pageId="63" pageNumber="64">
Key to species of
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma</emphasis>
</taxonomicName>
Britton
</paragraph>
<paragraph lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<table inLine="true" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<tr pageId="63" pageNumber="64">
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">1</td>
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">Hind angles of pronotum acute, strongly projected outward; pronotal base strongly lobed; elytra with 3rd to 5th intervals distinctly depressed subapically</td>
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">
African species (
<taxonomicName authorityName="Britton" authorityYear="1937" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="testaceum">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma testaceum</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma basilewskyi</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Basilewsky" authorityYear="1949" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="sudanicum">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma sudanicum</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Basilewsky" authorityYear="1968" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ivorense">
<emphasis italics="true" pageId="63" pageNumber="64">Dasiosoma ivorense</emphasis>
</taxonomicName>
)
</td>
</tr>
<tr pageId="63" pageNumber="64">
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">-</td>
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">Hind angles nearly rectangular, not strongly projected outward; pronotal base briefly lobed; elytral disc not depressed (Oriental species)</td>
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">2</td>
</tr>
<tr lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<td colspan="1" pageId="63" pageNumber="64" rowspan="1">2</td>
<td colspan="1" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" rowspan="1">
Basal foveae of pronotum straight, subparallel with median line, disc with distinct elongate depression on each side; body slender, ratio EL/EW more
<pageBreakToken pageId="64" pageNumber="65" start="start">than</pageBreakToken>
1.55; elytra dark blue, usually with a large reddish yellow patch occupying inner 3-4 intervals
</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Habu" baseAuthorityYear="1979" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bellum">
<emphasis italics="true" pageId="64" pageNumber="65">Dasiosoma bellum</emphasis>
</taxonomicName>
(Habu)
</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">-</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">Basal foveae of pronotum curved anteromedially, disc without or with barely visible depression; body stouter, ratio EL/EW less than 1.50; elytra reddish brown to dark brown, without distinct large patch on the center</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">3</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">3</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
Vertex strongly tumid; pronotum wider, lateral margins strongly expanded and completely rounded in middle; elytral color much darker than head and pronotum (
<figureCitation captionStart="Figures 55–60" captionStartId="F10" captionText="Figures 55 - 60. Type materials, scale bars = 2.0 mm: 55 Holotype of Dasiosoma sudanicum Basilewsky 56 Holotype of Dasiosoma ivorense Basilewsky 57 Holotype of Diamella indica Kirschenhofer [= Dasiosoma indicum (Kirschenhofer)] 58 Paratype of Lachnoderma maindroni Tian &amp; Deuve [= Dasiosoma maindroni (Tian &amp; Deuve)] 59 Lectotype of Singlis hirsutus Bates [= Dasiosoma hirsutum (Bates)] 60 The &quot; lectotype &quot; designated by Tian &amp; Deuve, 2001 for Singlis hirsutus Bates." httpUri="https://binary.pensoft.net/fig/29683" pageId="64" pageNumber="65">Figs 57, 58</figureCitation>
)
</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">4</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">-</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
Vertex slightly tumid; pronotum narrower, lateral margins slightly expanded and less rounded in middle; elytral color close to (only slightly darker than) head and pronotum (
<figureCitation captionStart="Figures 55–60" captionStartId="F10" captionText="Figures 55 - 60. Type materials, scale bars = 2.0 mm: 55 Holotype of Dasiosoma sudanicum Basilewsky 56 Holotype of Dasiosoma ivorense Basilewsky 57 Holotype of Diamella indica Kirschenhofer [= Dasiosoma indicum (Kirschenhofer)] 58 Paratype of Lachnoderma maindroni Tian &amp; Deuve [= Dasiosoma maindroni (Tian &amp; Deuve)] 59 Lectotype of Singlis hirsutus Bates [= Dasiosoma hirsutum (Bates)] 60 The &quot; lectotype &quot; designated by Tian &amp; Deuve, 2001 for Singlis hirsutus Bates." httpUri="https://binary.pensoft.net/fig/29683" pageId="64" pageNumber="65">Figs 59</figureCitation>
,
<figureCitation captionStart="Figures 61–66" captionStartId="F11" captionText="Figures 61 - 66. Type materials, scale bars = 2.0 mm: 61 Holotype of Dasiosoma quadraticolle sp. n. 62 Lectotype of Endynomena lewisii Bates [= Orionella lewisii (Bates)] 63 Holotype of Orionella obenbergi Jedlicka [= Orionella lewisii (Bates)] 64 Holotype of Endynomena discoidalis Bates [= Orionella discoidalis (Bates)] 65 Lectotype of Plochionus pradieri Fairmaire [= Endynomena pradieri (Fairmaire)] 66 Lectotype of Saronychium inconspicuum Blackburn [= Endynomena pradieri (Fairmaire)]." httpUri="https://binary.pensoft.net/fig/29684" pageId="64" pageNumber="65">61</figureCitation>
)
</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">5</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">4</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
Elytra dark brown, each side with a yellowish spot subapically, spots nearly joining at elytral suture; tempora gradually narrowed behind eyes; elytra wider, ratio EL/EW 1.30; body length 6.4 mm; species from India (
<figureCitation captionStart="Figures 55–60" captionStartId="F10" captionText="Figures 55 - 60. Type materials, scale bars = 2.0 mm: 55 Holotype of Dasiosoma sudanicum Basilewsky 56 Holotype of Dasiosoma ivorense Basilewsky 57 Holotype of Diamella indica Kirschenhofer [= Dasiosoma indicum (Kirschenhofer)] 58 Paratype of Lachnoderma maindroni Tian &amp; Deuve [= Dasiosoma maindroni (Tian &amp; Deuve)] 59 Lectotype of Singlis hirsutus Bates [= Dasiosoma hirsutum (Bates)] 60 The &quot; lectotype &quot; designated by Tian &amp; Deuve, 2001 for Singlis hirsutus Bates." httpUri="https://binary.pensoft.net/fig/29683" pageId="64" pageNumber="65">Fig. 57</figureCitation>
)
</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Kirschenhofer" baseAuthorityYear="2011" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="indicum">
<emphasis italics="true" pageId="64" pageNumber="65">Dasiosoma indicum</emphasis>
</taxonomicName>
(Kirschenhofer)
</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">-</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
Elytra uniformly dark brown, without spots; tempora abruptly narrowed behind eyes; elytra narrower, ratio EL/EW more than 1.35; body longer than 7.0 mm; species from Tonkin (
<figureCitation captionStart="Figures 55–60" captionStartId="F10" captionText="Figures 55 - 60. Type materials, scale bars = 2.0 mm: 55 Holotype of Dasiosoma sudanicum Basilewsky 56 Holotype of Dasiosoma ivorense Basilewsky 57 Holotype of Diamella indica Kirschenhofer [= Dasiosoma indicum (Kirschenhofer)] 58 Paratype of Lachnoderma maindroni Tian &amp; Deuve [= Dasiosoma maindroni (Tian &amp; Deuve)] 59 Lectotype of Singlis hirsutus Bates [= Dasiosoma hirsutum (Bates)] 60 The &quot; lectotype &quot; designated by Tian &amp; Deuve, 2001 for Singlis hirsutus Bates." httpUri="https://binary.pensoft.net/fig/29683" pageId="64" pageNumber="65">Fig. 58</figureCitation>
)
</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Tian &amp; Deuve" baseAuthorityYear="2001" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis italics="true" pageId="64" pageNumber="65">Dasiosoma maindroni</emphasis>
</taxonomicName>
(Tian &amp; Deuve)
</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">5</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">Pronotum quadrate, lateral margins nearly straight, slightly sinuate before hind angles; elytral striae shallower, with finer punctures, intervals slightly convex; head and pronotum brown to dark brown, elytra piceous, sometimes disc with a large indistinct brownish patch; Yunnan, Laos</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="quadraticolle">
<emphasis italics="true" pageId="64" pageNumber="65">Dasiosoma quadraticolle</emphasis>
</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="64" pageNumber="65">
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">-</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">Pronotum weakly cordiform, lateral margins slightly arcuate, strongly sinuate before hind angles; elytral striae deeper, with coarser punctures, intervals distinctly convex; dorsal side uniformly reddish brown; Hong Kong</td>
<td colspan="1" pageId="64" pageNumber="65" rowspan="1">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Dasiosoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="hirsutum">
<emphasis italics="true" pageId="64" pageNumber="65">Dasiosoma hirsutum</emphasis>
</taxonomicName>
(Bates)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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