418 lines
30 KiB
XML
418 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. & Marek, Paul E." docDate="2021" docId="7ABBB4AFFE4D5FD68BFD5E2F456842C3" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria sheari Means & Hennen & Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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||
<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1030</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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||
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
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||
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-GBIF-Taxon="181004735" LSID="urn:lsid:zoobank.org:act:2A2E7202-5E4E-4915-9533-5FD5235834E8" httpUri="http://treatment.plazi.org/id/7ABBB4AFFE4D5FD68BFD5E2F456842C3" lastPageNumber="1" pageId="0" pageNumber="1">
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||
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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||
<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="8RVWT" LSID="http://zoobank.org/2A2E7202-5E4E-4915-9533-5FD5235834E8" authority="Means & Hennen & Marek, 2021" authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria sheari" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sheari" status="sp. nov.">Nannaria sheari</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Nannaria sheari sp. nov. paratype ♂ (VMNH, NAN 0012) left gonopod A anterior view; red arrow indicates lateral flange B medial view C posterior view; red triangle indicates reduced, ridge-like prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure67" httpUri="https://binary.pensoft.net/fig/531170" pageId="0" pageNumber="1">Figs 67</figureCitation>
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<figureCitation captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Nannaria sheari sp. nov. holotype ♂ (VTEC, MPE 01684) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure68" httpUri="https://binary.pensoft.net/fig/531171" pageId="0" pageNumber="1">
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, 68 Vernacular name: "William
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<normalizedToken originalValue="Shear’s">Shear's</normalizedToken>
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Twisted-Claw Millipede"
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355465" accessionNumber="MPE01684" collectingDate="2016-06-14" collectorName="J. Means, D. Hennen" country="United States" county="Mercer County" elevation="628" latitude="37.4647" location="Brush Creek" longLatPrecision="6" longitude="-81.0623" municipality="Brush Creek Preserve" specimenCount="♂" stateProvince="West Virginia" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingCounty>Mercer County</collectingCounty>
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,
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<collectingMunicipality>
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<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:3DCB0A4A28E22AE3F8BAE80B76C4BD22" country="United States" county="Mercer County" latitude="37.4647" longLatPrecision="6" longitude="-81.0623" municipality="Brush Creek Preserve" name="Brush Creek" stateProvince="West Virginia">Brush Creek</location>
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Preserve
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</collectingMunicipality>
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, along trail to waterfall;
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<geoCoordinate degrees="37.4647" direction="north" orientation="latitude" precision="5" value="37.4647">37.4647°N</geoCoordinate>
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, -
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<geoCoordinate degrees="81.0623" direction="west" orientation="longitude" precision="5" value="-81.0623">81.0623°W</geoCoordinate>
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; elev.
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<quantity metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">628 m</elevation>
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</quantity>
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;
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<collectingDate value="2016-06-14">14 June 2016</collectingDate>
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; hand collected;
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<collectorName>J. Means</collectorName>
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,
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<collectorName>D. Hennen</collectorName>
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leg.; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE01684">MPE01684</accessionNumber>
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.
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355658" accessionNumber="MPE01685" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Mercer County" elevation="628" latitude="37.4647" location="West Virginia" longLatPrecision="6" longitude="-81.0623" municipality="Brush Creek Preserve" specimenCount="1" specimenCount-female="1" stateProvince="West Virginia" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount type="female">1 ♀</specimenCount>
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; same collection data as holotype; VMNH
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE01685">MPE01685</accessionNumber>
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</materialsCitation>
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•
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<materialsCitation ID-GBIF-Occurrence="3071355407" collectingDate="1967-10-05" collectorName="W. Shear" country="United States" county="Mercer County" latitude="37.4671" location="Tullgren" longLatPrecision="6" longitude="-81.0597" municipality="Brush Creek Falls" specimenCount="1" specimenCount-male="1" stateProvince="West Virginia" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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;
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<collectingCounty>Mercer County</collectingCounty>
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,
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<collectingMunicipality>Brush Creek Falls</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:4768E7B2E93B083A3EDC8E2E17C1F12D" country="United States" county="Mercer County" latitude="37.4671" longLatPrecision="6" longitude="-81.0597" municipality="Brush Creek Falls" name="Tullgren" stateProvince="West Virginia">Tullgren</location>
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, sort of
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rhododendron" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">Rhododendron</taxonomicName>
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litter;
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<geoCoordinate degrees="37.4671" direction="north" orientation="latitude" precision="5" value="37.4671">37.4671°N</geoCoordinate>
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, -
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<geoCoordinate degrees="81.0597" direction="west" orientation="longitude" precision="5" value="-81.0597">81.0597°W</geoCoordinate>
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;
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<collectingDate value="1967-10-05">5 Oct.1967</collectingDate>
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;
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<collectorName>W. Shear</collectorName>
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leg.; VMNH NAN0012
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355355" collectingDate="1967-04-04" collectorName="W. Shear" country="United States" county="Mercer County" latitude="37.4564" location="Speedway Hemlock Grove" longLatPrecision="6" longitude="-81.0105" specimenCount="1" specimenCount-female="1" stateProvince="West Virginia">
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount type="female">1 ♀</specimenCount>
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;
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<collectingCounty>Mercer County</collectingCounty>
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,
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<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:C2F195F1816E79B3884211C234DFC89F" country="United States" county="Mercer County" latitude="37.4564" longLatPrecision="6" longitude="-81.0105" name="Speedway Hemlock Grove" stateProvince="West Virginia">Speedway Hemlock Grove</location>
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, oak-pine litter;
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<geoCoordinate degrees="37.4564" direction="north" orientation="latitude" precision="5" value="37.4564">37.4564°N</geoCoordinate>
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, -
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<geoCoordinate degrees="81.0105" direction="west" orientation="longitude" precision="5" value="-81.0105">81.0105°W</geoCoordinate>
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;
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<collectingDate value="1967-04-04">4 Apr. 1967</collectingDate>
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;
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<collectorName>W. Shear</collectorName>
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leg.; VMNH NAN0002
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</materialsCitation>
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•
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<materialsCitation ID-GBIF-Occurrence="3071355310" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="W. Shear" country="United States" county="Mercer County" latitude="37.4564" location="Speedway Hemlock Grove" longLatPrecision="6" longitude="-81.0105" specimenCount="2" specimenCount-male="2" stateProvince="West Virginia">
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<specimenCount type="male">2 ♂♂</specimenCount>
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; same collection data as preceding; VMNH NAN0048
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</materialsCitation>
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•
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<materialsCitation ID-GBIF-Occurrence="3071355702" collectingDate="1968-03-09" collectorName="W. Shear" country="United States" county="Mercer County" latitude="37.4646" location="Athens" longLatPrecision="6" longitude="-81.0112" municipality="Speedway Roadside Park" specimenCount="13" specimenCount-male="13" stateProvince="West Virginia">
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<specimenCount type="male">13 ♂</specimenCount>
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<specimenCount>♀</specimenCount>
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;
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<collectingCounty>Mercer County</collectingCounty>
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,
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<collectingMunicipality>Speedway Roadside Park</collectingMunicipality>
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, route 20, 3 mi. north of
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<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:7D490229E2B8A796DD4AC3CCE4F243C1" country="United States" county="Mercer County" latitude="37.4646" longLatPrecision="6" longitude="-81.0112" municipality="Speedway Roadside Park" name="Athens" stateProvince="West Virginia">Athens</location>
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, hibernating aggregation of dead animals under log;
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||
<geoCoordinate degrees="37.4646" direction="north" orientation="latitude" precision="5" value="37.4646">37.4646°N</geoCoordinate>
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, -
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<geoCoordinate degrees="81.0112" direction="west" orientation="longitude" precision="5" value="-81.0112">81.0112°W</geoCoordinate>
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;
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<collectingDate value="1968-03-09">9 Mar. 1968</collectingDate>
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||
;
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||
<collectorName>W. Shear</collectorName>
|
||
leg.; VMNH NAN0020
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||
</materialsCitation>
|
||
•
|
||
<materialsCitation ID-GBIF-Occurrence="3071355486" accessionNumber="MPE01665" collectingDate="2016-06-14" collectorName="J. Means, D. Hennen" country="United States" county="Mercer County" elevation="628" latitude="37.4647" location="Brush Creek" longLatPrecision="6" longitude="-81.0623" specimenCount="3" specimenCount-male="3" stateProvince="West Virginia">
|
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<specimenCount type="male">3 ♂♂</specimenCount>
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||
;
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||
<collectingCounty>Mercer County</collectingCounty>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:3F05A740434A60177078B226AA74FBC3" country="United States" county="Mercer County" latitude="37.4647" longLatPrecision="6" longitude="-81.0623" name="Brush Creek" stateProvince="West Virginia">Brush Creek</location>
|
||
Preserve, along trail to waterfall;
|
||
<geoCoordinate degrees="37.4647" direction="north" orientation="latitude" precision="5" value="37.4647">37.4647°N</geoCoordinate>
|
||
, -
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||
<geoCoordinate degrees="81.0623" direction="west" orientation="longitude" precision="5" value="-81.0623">81.0623°W</geoCoordinate>
|
||
; elev.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" unit="m" value="628.0">628 m</elevation>
|
||
</quantity>
|
||
;
|
||
<collectingDate value="2016-06-14">14 June 2016</collectingDate>
|
||
; hand collected;
|
||
<collectorName>J. Means</collectorName>
|
||
,
|
||
<collectorName>D. Hennen</collectorName>
|
||
leg.; VTEC
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE01665">MPE01665</accessionNumber>
|
||
</materialsCitation>
|
||
-7 •
|
||
<materialsCitation ID-GBIF-Occurrence="3071355592" collectingDate="1968-03-30" collectorName="W. Shear" country="United States" county="Mercer County" latitude="37.4222" location="Jackson's Park" longLatPrecision="6" longitude="-81.0163" municipality="Athens" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="West Virginia">
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingCounty>Mercer County</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Athens</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:321F21E507160CE3FD0A7DE315862F1F" country="United States" county="Mercer County" latitude="37.4222" longLatPrecision="6" longitude="-81.0163" municipality="Athens" name="Jackson's Park" stateProvince="West Virginia">
|
||
<normalizedToken originalValue="Jackson’s">Jackson's</normalizedToken>
|
||
Park
|
||
</location>
|
||
;
|
||
<geoCoordinate degrees="37.4222" direction="north" orientation="latitude" precision="5" value="37.4222">37.4222°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate degrees="81.0163" direction="west" orientation="longitude" precision="5" value="-81.0163">81.0163°W</geoCoordinate>
|
||
;
|
||
<collectingDate value="1968-03-30">30 Mar. 1968</collectingDate>
|
||
;
|
||
<collectorName>W. Shear</collectorName>
|
||
leg.; VMNH NAN0013
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation ID-GBIF-Occurrence="3071355466" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Athens" country="United States" county="Mercer County" location="Mercer County" specimenCount="1" specimenCount-male="1" stateProvince="West Virginia">
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingCounty>Mercer County</collectingCounty>
|
||
,
|
||
<collectorName>Athens</collectorName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation ID-GBIF-Occurrence="3071355585" collectingDate="1973-03-29" collectorName="W. Shear, For, Suppl." country="United States" latitude="37.4254" location="Broadway" longLatPrecision="6" longitude="-81.0211" specimenCount="218" stateProvince="West Virginia">
|
||
<specimenCount type="generic">218 W.</specimenCount>
|
||
<location LSID="urn:lsid:plazi:treatment:7ABBB4AFFE4D5FD68BFD5E2F456842C3:FE1E71C5954A986854F1DDB4028537CD" country="United States" latitude="37.4254" longLatPrecision="6" longitude="-81.0211" name="Broadway" stateProvince="West Virginia">Broadway</location>
|
||
;
|
||
<geoCoordinate degrees="37.4254" direction="north" orientation="latitude" precision="5" value="37.4254">37.4254°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate degrees="81.0211" direction="west" orientation="longitude" precision="5" value="-81.0211">81.0211°W</geoCoordinate>
|
||
;
|
||
<collectingDate value="1973-03-29">29 Mar. 1973</collectingDate>
|
||
;
|
||
<collectorName>W. Shear</collectorName>
|
||
leg.; VMNH NAN0046.
|
||
<collectorName>For</collectorName>
|
||
detailed collection data see
|
||
<collectorName>Suppl.</collectorName>
|
||
material 7
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Adult males of
|
||
<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria sheari" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sheari">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria sheari</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are distinct from other
|
||
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
|
||
</taxonomicName>
|
||
and the nearby
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
|
||
</taxonomicName>
|
||
, based on the following combination of characters:
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
|
||
.
|
||
</emphasis>
|
||
Gonopodal acropodite continually curving medially, not relatively straight before curving medially at half-way point as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castra" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. castra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Distal zone curving medially, not dorsally as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castra" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. castra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Acropodite tip directed caudally (Fig.
|
||
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Nannaria sheari sp. nov. paratype ♂ (VMNH, NAN 0012) left gonopod A anterior view; red arrow indicates lateral flange B medial view C posterior view; red triangle indicates reduced, ridge-like prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure67" httpUri="https://binary.pensoft.net/fig/531170" pageId="0" pageNumber="1">67B</figureCitation>
|
||
), not medially as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
|
||
</taxonomicName>
|
||
. Acropodite tip with lateral flange (Fig.
|
||
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Nannaria sheari sp. nov. paratype ♂ (VMNH, NAN 0012) left gonopod A anterior view; red arrow indicates lateral flange B medial view C posterior view; red triangle indicates reduced, ridge-like prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure67" httpUri="https://binary.pensoft.net/fig/531170" pageId="0" pageNumber="1">67</figureCitation>
|
||
, red arrow). Acropodite simple in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. sheari" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="sheari">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. sheari</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., lacking slight twist and swelling at midpoint as found in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castra" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. castra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., and lateral flange as found in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
|
||
</taxonomicName>
|
||
. Telopodite basal zone with slight lateral bulge and wider than space between prefemoral process and acropodite, not thinner as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castra" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. castra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Prefemur with dorsomedially curving prefemoral process (Fig.
|
||
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Nannaria sheari sp. nov. paratype ♂ (VMNH, NAN 0012) left gonopod A anterior view; red arrow indicates lateral flange B medial view C posterior view; red triangle indicates reduced, ridge-like prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure67" httpUri="https://binary.pensoft.net/fig/531170" pageId="0" pageNumber="1">67B</figureCitation>
|
||
), not straight as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
|
||
</taxonomicName>
|
||
. Prefemoral spine fused with prefemoral process, reduced to shelf-like ridge at base of prefemoral process (Fig.
|
||
<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Nannaria sheari sp. nov. paratype ♂ (VMNH, NAN 0012) left gonopod A anterior view; red arrow indicates lateral flange B medial view C posterior view; red triangle indicates reduced, ridge-like prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure67" httpUri="https://binary.pensoft.net/fig/531170" pageId="0" pageNumber="1">67C</figureCitation>
|
||
, red triangle).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tergites with orange paranotal spots (Fig.
|
||
<figureCitation captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Nannaria sheari sp. nov. holotype ♂ (VTEC, MPE 01684) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure68" httpUri="https://binary.pensoft.net/fig/531171" pageId="0" pageNumber="1">68</figureCitation>
|
||
). Dark brown background. Dorsum of collum smooth with orange and white margin.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1030.62544.figure67" httpUri="https://binary.pensoft.net/fig/531170" pageId="0" pageNumber="1" start="Figure 67" startId="F67">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 67.</emphasis>
|
||
<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria sheari" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sheari">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria sheari</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. paratype ♂ (VMNH, NAN0012) left gonopod
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
anterior view; red arrow indicates lateral flange
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
medial view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
posterior view; red triangle indicates reduced, ridge-like prefemoral spine. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1030.62544.figure68" httpUri="https://binary.pensoft.net/fig/531171" pageId="0" pageNumber="1" start="Figure 68" startId="F68">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 68.</emphasis>
|
||
<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria sheari" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sheari">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria sheari</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype ♂ (VTEC, MPE01684) coloration. Scale bar: 4.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">♂ holotype (VTEC, MPE01684): BL = 30.7, CW = 3.5, IW = 1.9, ISW = 0.7, B11W = 4.1, B11H = 2.4. ♀ paratype (VMNH, MPE01685): BL = 30.2, CW = 3.2, IW = 1.9, ISW = 0.8, B11W = 4.5, B11H = 3.3.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="variation">
|
||
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Known from a small area in and around Athens, West Virginia (West Virginia: Mercer County, Suppl. material 7; Fig.
|
||
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
|
||
). Distribution area: 12 km2; status: MRE.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="ecology">
|
||
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Individuals of
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. sheari" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="sheari">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. sheari</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. have been collected from mesic hardwood forests composed of oak, tuliptree, birch, maple, buckeye, rhododendron, and some pine and hemlock. Individuals were found under deciduous leaf litter by the side of hiking trails.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">This species is named after its original collector, and a longtime mentor to the authors, Dr. William Shear. The specific name is a genitive noun derived as a patronym.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type locality">
|
||
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
United States, West Virginia, Mercer County, Brush Creek Preserve, along trail to waterfall,
|
||
<geoCoordinate degrees="37.4647" direction="north" orientation="latitude" precision="5" value="37.4647">37.4647°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate degrees="81.0623" direction="west" orientation="longitude" precision="5" value="-81.0623">81.0623°W</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |