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<document id="DC6FA6552EEBEC65D08315ECB66451CA" ID="10.11646/zootaxa.4170.2.7" ID-DOI="10.11646/zootaxa.4170.2.7" ID-GBIF-Dataset="54d2c6e9-9ad2-402e-8e4e-c569c4c5c027" ID-ISSN="1175-5326" ID-Zenodo-Dep="259264" ID-ZooBank="68799414-3E78-43A3-8A93-BE50F0F82407" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474614126190" checkinUser="plazi" docAuthor="Agarwal, Ishan, Mirza, Zeeshan A., Pal, Saunak, Maddock, Simon T., Mishra, Anurag &amp; Bauer, Aaron M." docDate="2016" docId="5F5C6110FF8CB911FF48FB99FAE0F858" docLanguage="en" docName="zootaxa.4170.2.7.pdf" docOrigin="Zootaxa 4170 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus collegalensis" docType="treatment" docVersion="9" lastPageNumber="344" masterDocId="A3651968FF88B914FFDFFF8EFFC2FFF0" masterDocTitle="A new species of the Cyrtodactylus (Geckoella) collegalensis (Beddome, 1870) complex (Squamata: Gekkonidae) from Western India" masterLastPageNumber="354" masterPageNumber="339" pageNumber="343" updateTime="1698387113481" updateUser="plazi">
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<mods:title id="EB08BB8D8539351D145371291CF97FAC">A new species of the Cyrtodactylus (Geckoella) collegalensis (Beddome, 1870) complex (Squamata: Gekkonidae) from Western India</mods:title>
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<mods:namePart id="A81E30BDC8E49A8C8FE860E0770A3E70">Agarwal, Ishan</mods:namePart>
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<mods:namePart id="DF41FE6E3F8BCA1CD743B6A7E6461AEC">Mirza, Zeeshan A.</mods:namePart>
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<mods:namePart id="0BA08C16CE033D9C89A4E4AE493F0C35">Pal, Saunak</mods:namePart>
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<mods:namePart id="8D4530034861723B4D0C186D1FA7F607">Maddock, Simon T.</mods:namePart>
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<mods:namePart id="FD51281241C1DD8EEC10CC6B4CBDABE0">Mishra, Anurag</mods:namePart>
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<mods:namePart id="2732D35E890F8B9168B5AB1C56B42D2D">Bauer, Aaron M.</mods:namePart>
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<mods:date id="BDC1893C1A582DFA1D4CCEE9B04A457D">2016</mods:date>
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<mods:number id="2C1887801001229A3C4909C6304C631B">4170</mods:number>
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<treatment id="5F5C6110FF8CB911FF48FB99FAE0F858" ID-DOI="http://doi.org/10.5281/zenodo.5670212" ID-GBIF-Taxon="124552800" ID-Zenodo-Dep="5670212" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858" httpUri="http://treatment.plazi.org/id/5F5C6110FF8CB911FF48FB99FAE0F858" lastPageId="5" lastPageNumber="344" pageId="4" pageNumber="343">
<subSubSection id="9FEF838DFF8CB910FF48FB99FF1BFBA2" pageId="4" pageNumber="343" type="nomenclature">
<paragraph id="D74AD006FF8CB910FF48FB99FE1DFBC1" blockId="4.[151,479,1047,1107]" box="[151,479,1047,1073]" pageId="4" pageNumber="343">
<heading id="8C02676AFF8CB910FF48FB99FE1DFBC1" bold="true" box="[151,479,1047,1073]" fontSize="11" level="1" pageId="4" pageNumber="343" reason="1">
<taxonomicName id="10F5AB85FF8CB910FF48FB99FE1DFBC1" box="[151,479,1047,1073]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="343" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8CB910FF48FB99FE1DFBC1" bold="true" box="[151,479,1047,1073]" italics="true" pageId="4" pageNumber="343">Cyrtodactylus collegalensis</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D74AD006FF8CB910FF48FBB4FF1BFBA2" blockId="4.[151,479,1047,1107]" box="[151,217,1082,1107]" pageId="4" pageNumber="343">
<figureCitation id="4FCECC83FF8CB910FF48FBB4FF1BFBA2" box="[151,217,1082,1107]" captionStart="FIGURE 3" captionStartId="5.[151,250,1135,1157]" captionTargetBox="[150,1432,227,1099]" captionTargetId="figure@5.[147,1432,227,1100]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of Cyrtodactylus collegalensis (top row, A F) and C. speciosus (bottom row, G L). A, holotype of C. collegalensis (BMNH 1946.8.25.28); B, CES / 09 / 1444; C, CES / 09 / 1463; D, CES / 09 / 1442; E, CES / 09 / 1443; F, C. collegalensis, CES / 09 / 1403. G, holotype of C. speciosus, BMNH 1946.9.4.88; H, presumed sketch of C. speciosus (Boulenger 1885); I, CES / 09 / 1405; J, CES / 09 / 1406; K, CES / 09 / 1407; L, CES / 09 / 1408." httpUri="https://zenodo.org/record/259266/files/figure.png" pageId="4" pageNumber="343">Fig. 3</figureCitation>
</paragraph>
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<subSubSection id="9FEF838DFF8CB910FF48FB0CFD52FB25" pageId="4" pageNumber="343" type="reference_group">
<paragraph id="D74AD006FF8CB910FF48FB0CFD9DFB68" blockId="4.[151,673,1154,1237]" box="[151,607,1154,1176]" pageId="4" pageNumber="343">
<taxonomicName id="10F5AB85FF8CB910FF48FB0CFD9DFB68" authority="Beddome 1870" authorityName="Beddome" authorityYear="1870" box="[151,607,1154,1176]" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="343" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8CB910FF48FB0CFE7DFB68" box="[151,447,1154,1176]" italics="true" pageId="4" pageNumber="343">Gymnodactylus collegalensis</emphasis>
<bibRefCitation id="B364ADF7FF8CB910FE1AFB0CFD9DFB68" author="Beddome" box="[453,607,1154,1176]" pageId="4" pageNumber="343" refString="Beddome, R. H. (1870) Description of new reptiles from Madras presidency. Madras Monthly Journal of Medical Science, 2, 169 - 176, 2 pls." type="journal article" year="1870">Beddome 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="D74AD006FF8CB910FF48FB2FFD52FB25" blockId="4.[151,673,1154,1237]" pageId="4" pageNumber="343">
<taxonomicName id="10F5AB85FF8CB910FF48FB2FFD8BFB47" authority="Boulenger 1885" authorityName="Boulenger" authorityYear="1885" box="[151,585,1185,1207]" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="343" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="E5810C14FF8CB910FF48FB2FFE62FB47" box="[151,416,1185,1207]" italics="true" pageId="4" pageNumber="343">Gymnodactylus nebulosus</emphasis>
<bibRefCitation id="B364ADF7FF8CB910FE79FB2FFD8BFB47" author="Boulenger" box="[422,585,1185,1207]" pageId="4" pageNumber="343" refString="Boulenger, J. A. (1885) Catalogue of the lizards in the British Museum (Natural History). I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. British Museum (Natural History), London, xii + 436 pp., 32 pls." type="book" year="1885">Boulenger 1885</bibRefCitation>
</taxonomicName>
(in part)
<taxonomicName id="10F5AB85FF8CB910FF48FB31FD52FB25" authority="Agarwal &amp; Karanth 2015" authorityName="Agarwal &amp; Karanth" authorityYear="2015" box="[151,656,1215,1237]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="343" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8CB910FF48FB31FE4AFB25" box="[151,392,1215,1237]" italics="true" pageId="4" pageNumber="343">Geckoella collegalensis</emphasis>
<bibRefCitation id="B364ADF7FF8CB910FE51FB31FD52FB25" author="Agarwal" box="[398,656,1215,1237]" pageId="4" pageNumber="343" refString="Agarwal, I. &amp; Karanth, K. P. (2015) A phylogeny of the only ground-dwelling radiation of Cyrtodactylus (Squamata, Gekkonidae): diversification of Geckoella across peninsular India and Sri Lanka. Molecular Phylogenetics and Evolution, 82, 193 - 199. http: // dx. doi. org / 10.1016 / j. ympev. 2014.09.016" type="journal article" year="2015">Agarwal &amp; Karanth 2015</bibRefCitation>
</taxonomicName>
</paragraph>
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<paragraph id="D74AD006FF8CB910FF48FA8CFADCFACF" blockId="4.[151,1437,1282,2030]" pageId="4" pageNumber="343">
<materialsCitation id="679DDA5BFF8CB910FF48FA8CFADCFACF" ID-GBIF-Occurrence="1314772787" collectionCode="BMNH" collectorName="BR Hills &amp; R. H. Beddome." country="India" county="Chamarajanagara District" location="Balarangans" municipality="Yelandur" pageId="4" pageNumber="343" specimenCode="BMNH 1946.8" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<emphasis id="E5810C14FF8CB910FF48FA8CFEC9FAEB" bold="true" box="[151,267,1282,1307]" pageId="4" pageNumber="343">
<typeStatus id="084E6EA4FF8CB910FF48FA8CFEC4FAEB" box="[151,262,1282,1307]" pageId="4" pageNumber="343" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="8753787DFF8CB910FEC6FA8DFE0EFAEB" box="[281,460,1282,1307]" collectionCode="BMNH" pageId="4" pageNumber="343">
<collectionCode id="B1E448C3FF8CB910FEC6FA8DFEAFFAEA" box="[281,365,1283,1306]" pageId="4" pageNumber="343">BMNH</collectionCode>
1946.8
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.25.28, adult male, “lower slopes of
<location id="D22A86DDFF8CB910FC51FA8CFBDBFAEB" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858:D22A86DDFF8CB910FC51FA8CFBDBFAEB" box="[910,1049,1282,1307]" country="India" county="Chamarajanagara District" municipality="Yelandur" name="Balarangans" pageId="4" pageNumber="343" stateProvince="Karnataka">Balarangans</location>
(
<collectorName id="7A00B5D0FF8CB910FBF0FA8CFB5CFAEB" box="[1071,1182,1282,1307]" pageId="4" pageNumber="343">BR Hills</collectorName>
), in dry forests near
<location id="D22A86DDFF8CB910FF48FAA8FECAFACF" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858:D22A86DDFF8CB910FF48FAA8FECAFACF" box="[151,264,1318,1343]" country="India" county="Chamarajanagara District" municipality="Yelandur" name="Yellundur" pageId="4" pageNumber="343" stateProvince="Karnataka">Yellundur</location>
” (today
<collectingMunicipality id="372E4A7CFF8CB910FEB8FAA8FE0DFACF" box="[359,463,1318,1343]" pageId="4" pageNumber="343">Yelandur</collectingMunicipality>
,
<collectingCounty id="3E2BA88AFF8CB910FE05FAA8FD3EFACF" box="[474,764,1318,1343]" pageId="4" pageNumber="343">Chamarajanagara District</collectingCounty>
,
<collectingRegion id="15311EE4FF8CB910FCD8FAA8FCBFFACF" box="[775,893,1318,1343]" country="India" name="Karnataka" pageId="4" pageNumber="343">Karnataka</collectingRegion>
,
<collectingCountry id="AFE29096FF8CB910FC57FAA8FC04FACF" box="[904,966,1318,1343]" name="India" pageId="4" pageNumber="343">India</collectingCountry>
), collected by
<collectorName id="7A00B5D0FF8CB910FBB7FAA9FADCFACF" box="[1128,1310,1318,1343]" pageId="4" pageNumber="343">R. H. Beddome.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="D74AD006FF8CB910FF18FAC4FCE4F9E7" blockId="4.[151,1437,1282,2030]" pageId="4" pageNumber="343">
<materialsCitation id="679DDA5BFF8CB910FF18FAC4FC37FA77" ID-GBIF-Occurrence="1314772789" collectingDate="2013-04-13" collectorName="MM Hills Wildlife Sanctuary &amp; Chintan Sheth &amp; Ishan Agarwal" country="India" location="Chamarajanagar District" pageId="4" pageNumber="343" specimenCount="1" stateProvince="Karnataka">
<emphasis id="E5810C14FF8CB910FF18FAC4FE7FFA93" bold="true" box="[199,445,1354,1379]" pageId="4" pageNumber="343">Additional material.</emphasis>
CES/09/1444, subadult male,
<collectorName id="7A00B5D0FF8CB910FCC1FAC5FBB3FA92" box="[798,1137,1354,1379]" pageId="4" pageNumber="343">MM Hills Wildlife Sanctuary</collectorName>
,
<location id="D22A86DDFF8CB910FBA0FAC4FA5AFA93" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858:D22A86DDFF8CB910FBA0FAC4FA5AFA93" box="[1151,1432,1354,1379]" country="India" name="Chamarajanagar District" pageId="4" pageNumber="343" stateProvince="Karnataka">Chamarajanagar District</location>
,
<collectingRegion id="15311EE4FF8CB910FF48FAE0FECFFA77" box="[151,269,1390,1415]" country="India" name="Karnataka" pageId="4" pageNumber="343">Karnataka</collectingRegion>
,
<collectingCountry id="AFE29096FF8CB910FEC8FAE0FE91FA77" box="[279,339,1390,1415]" name="India" pageId="4" pageNumber="343">India</collectingCountry>
, collected by
<collectorName id="7A00B5D0FF8CB910FE32FAE0FD4EFA77" box="[493,652,1390,1415]" pageId="4" pageNumber="343">Chintan Sheth</collectorName>
and
<collectorName id="7A00B5D0FF8CB910FD1DFAE0FCAAFA77" box="[706,872,1390,1415]" pageId="4" pageNumber="343">Ishan Agarwal</collectorName>
,
<date id="A34BF6C6FF8CB910FCACFAE1FC37FA77" box="[883,1013,1391,1415]" pageId="4" pageNumber="343" value="2013-04-13">
<collectingDate id="B30F0F2EFF8CB910FCACFAE1FC37FA77" box="[883,1013,1391,1415]" pageId="4" pageNumber="343" value="2013-04-13">13.04.2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="679DDA5BFF8CB910FC20FAE0FE0EFA3F" ID-GBIF-Occurrence="1314772796" collectingDate="2013-09-14" collectorName="Ishan Agarwal &amp; Saunak Pal &amp; Varun Torsekar &amp; Navendu Page" country="India" location="Kollegal Taluk" municipality="Chamarajanagar District" pageId="4" pageNumber="343" specimenCount="1" stateProvince="Karnataka">
CES/09/1463, adult female,
<location id="D22A86DDFF8CB910FAE3FAE0FF1BFA5B" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858:D22A86DDFF8CB910FAE3FAE0FF1BFA5B" country="India" municipality="Chamarajanagar District" name="Kollegal Taluk" pageId="4" pageNumber="343" stateProvince="Karnataka">Kollegal Taluk</location>
,
<collectingMunicipality id="372E4A7CFF8CB910FF33FA1CFDCAFA5B" box="[236,520,1426,1451]" pageId="4" pageNumber="343">Chamarajanagar District</collectingMunicipality>
,
<collectingRegion id="15311EE4FF8CB910FDC4FA1CFD53FA5B" box="[539,657,1426,1451]" country="India" name="Karnataka" pageId="4" pageNumber="343">Karnataka</collectingRegion>
,
<collectingCountry id="AFE29096FF8CB910FD7CFA1CFD1FFA5B" box="[675,733,1426,1451]" name="India" pageId="4" pageNumber="343">India</collectingCountry>
collected by
<collectorName id="7A00B5D0FF8CB910FC56FA1CFBF5FA5B" box="[905,1079,1426,1451]" pageId="4" pageNumber="343">Ishan Agarwal</collectorName>
,
<collectorName id="7A00B5D0FF8CB910FB95FA1CFB13FA5B" box="[1098,1233,1426,1451]" pageId="4" pageNumber="343">Saunak Pal</collectorName>
,
<collectorName id="7A00B5D0FF8CB910FB3CFA1DFA5AFA5B" box="[1251,1432,1426,1451]" pageId="4" pageNumber="343">Varun Torsekar</collectorName>
,
<collectorName id="7A00B5D0FF8CB910FF48FA38FEFFFA3F" box="[151,317,1462,1487]" pageId="4" pageNumber="343">Navendu Page</collectorName>
,
<date id="A34BF6C6FF8CB910FE95FA39FE0EFA3F" box="[330,460,1463,1487]" pageId="4" pageNumber="343" value="2013-09-14">
<collectingDate id="B30F0F2EFF8CB910FE95FA39FE0EFA3F" box="[330,460,1463,1487]" pageId="4" pageNumber="343" value="2013-09-14">14.09.2013</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="679DDA5BFF8CB910FE08FA38FCE1F9E7" ID-GBIF-Occurrence="1314772786" collectingDate="2013-04-13" collectorName="MM Hills Wildlife Sanctuary &amp; Chintan Sheth, V. &amp; Deepak &amp; Ishan Agarwal" country="India" location="Unvouchered" municipality="Chamarajanagar District" pageId="4" pageNumber="343" specimenCount="1" stateProvince="Karnataka">
<location id="D22A86DDFF8CB910FE08FA38FDACFA3F" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858:D22A86DDFF8CB910FE08FA38FDACFA3F" box="[471,622,1462,1487]" country="India" municipality="Chamarajanagar District" name="Unvouchered" pageId="4" pageNumber="343" stateProvince="Karnataka">Unvouchered</location>
photographic records and tissue collected for two additional specimens CES/09/
<date id="A34BF6C6FF8CB910FF26FA54FEB5FA02" box="[249,375,1498,1522]" pageId="4" pageNumber="343" value="1442" valueMax="1443">14421443</date>
;
<collectorName id="7A00B5D0FF8CB910FE54FA55FD2CFA02" box="[395,750,1498,1523]" pageId="4" pageNumber="343">MM Hills Wildlife Sanctuary</collectorName>
,
<collectingMunicipality id="372E4A7CFF8CB910FCDEFA54FBDDFA03" box="[769,1055,1498,1523]" pageId="4" pageNumber="343">Chamarajanagar District</collectingMunicipality>
,
<collectingRegion id="15311EE4FF8CB910FBECFA54FB6BFA03" box="[1075,1193,1498,1523]" country="India" name="Karnataka" pageId="4" pageNumber="343">Karnataka</collectingRegion>
,
<collectingCountry id="AFE29096FF8CB910FB63FA54FB3AFA03" box="[1212,1272,1498,1523]" name="India" pageId="4" pageNumber="343">India</collectingCountry>
, collected by
<collectorName id="7A00B5D0FF8CB910FF48FA70FE7BF9E7" box="[151,441,1534,1559]" pageId="4" pageNumber="343">Chintan Sheth, V. Deepak</collectorName>
and
<collectorName id="7A00B5D0FF8CB910FE30FA70FD54F9E7" box="[495,662,1534,1559]" pageId="4" pageNumber="343">Ishan Agarwal</collectorName>
,
<date id="A34BF6C6FF8CB910FD7EFA71FCE1F9E7" box="[673,803,1535,1559]" pageId="4" pageNumber="343" value="2013-04-13">
<collectingDate id="B30F0F2EFF8CB910FD7EFA71FCE1F9E7" box="[673,803,1535,1559]" pageId="4" pageNumber="343" value="2013-04-13">13.04.2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="D74AD006FF8CB910FF18F9ACFB53F973" blockId="4.[151,1437,1282,2030]" pageId="4" pageNumber="343">
<materialsCitation id="679DDA5BFF8CB910FF18F9ACFB53F973" ID-GBIF-Occurrence="1314772784" collectingDate="2012-05-26" collectorName="V. Deepak &amp; Ishan Agarwal" country="India" county="Salem District" location="An" municipality="Mettur Taluk" pageId="4" pageNumber="343" specimenCount="1" stateProvince="Tamil Nadu">
<location id="D22A86DDFF8CB910FF18F9ACFF2BF9CA" LSID="urn:lsid:plazi:treatment:5F5C6110FF8CB911FF48FB99FAE0F858:D22A86DDFF8CB910FF18F9ACFF2BF9CA" box="[199,233,1570,1594]" country="India" county="Salem District" municipality="Mettur Taluk" name="An" pageId="4" pageNumber="343" stateProvince="Tamil Nadu">An</location>
additional specimen, though differing in coloration from topotypical
<taxonomicName id="10F5AB85FF8CB910FBDDF9ACFB74F9CA" box="[1026,1206,1570,1594]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="343" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8CB910FBDDF9ACFB74F9CA" box="[1026,1206,1570,1594]" italics="true" pageId="4" pageNumber="343">C. collegalensis</emphasis>
</taxonomicName>
and 4.7% divergent (uncorrected p-distance, ND2), is tentatively referred to this species. CES/09/1403, adult female,
<collectingMunicipality id="372E4A7CFF8CB910FB20F9C9FA5AF9AF" box="[1279,1432,1606,1631]" pageId="4" pageNumber="343">Mettur Taluk</collectingMunicipality>
,
<collectingCounty id="3E2BA88AFF8CB910FF48F9E4FEF9F973" box="[151,315,1642,1667]" pageId="4" pageNumber="343">Salem District</collectingCounty>
,
<collectingRegion id="15311EE4FF8CB910FE99F9E4FE0FF973" box="[326,461,1642,1667]" country="India" name="Tamil Nadu" pageId="4" pageNumber="343">Tamil Nadu</collectingRegion>
,
<collectingCountry id="AFE29096FF8CB910FE07F9E4FDD6F973" box="[472,532,1642,1667]" name="India" pageId="4" pageNumber="343">India</collectingCountry>
, collected by
<collectorName id="7A00B5D0FF8CB910FD70F9E5FCE6F973" box="[687,804,1642,1667]" pageId="4" pageNumber="343">V. Deepak</collectorName>
and
<collectorName id="7A00B5D0FF8CB910FC85F9E4FBC2F973" box="[858,1024,1642,1667]" pageId="4" pageNumber="343">Ishan Agarwal</collectorName>
,
<date id="A34BF6C6FF8CB910FBD3F9E4FB4CF973" box="[1036,1166,1642,1667]" pageId="4" pageNumber="343" value="2012-05-26">
<collectingDate id="B30F0F2EFF8CB910FBD3F9E4FB4CF973" box="[1036,1166,1642,1667]" pageId="4" pageNumber="343" value="2012-05-26">26.05.2012</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="D74AD006FF8CB910FF18F900FAE4F81E" blockId="4.[151,1437,1282,2030]" pageId="4" pageNumber="343">
<emphasis id="E5810C14FF8CB910FF18F900FE84F957" bold="true" box="[199,326,1678,1703]" pageId="4" pageNumber="343">Definition.</emphasis>
A small
<taxonomicName id="10F5AB85FF8CB910FE6DF900FD8DF956" box="[434,591,1678,1702]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="343" phylum="Chordata" rank="genus">
<emphasis id="E5810C14FF8CB910FE6DF900FD8DF956" box="[434,591,1678,1702]" italics="true" pageId="4" pageNumber="343">Cyrtodactylus</emphasis>
</taxonomicName>
, snout-vent length to at least
<quantity id="100D7DE3FF8CB910FC70F900FBC1F956" box="[943,1027,1678,1703]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="343" unit="mm" value="54.0">54 mm</quantity>
; head short, body moderately stout, limbs and digits not long, slender; two pairs of enlarged postmentals, outer pair smaller than and separated by inner pair, which are in broad contact; homogeneous dorsal scalation consisting of small, granular, roughly circular scales; 2729 ventral scales across belly; no precloacal groove, no enlarged precloacal or femoral scales, no precloacal or femoral pores. Nine to ten lamellae in basal series and
<quantity id="100D7DE3FF8CB910FC6AF8AFFC3DF8CA" box="[949,1023,1825,1850]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="4" pageNumber="343" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">69 in</quantity>
apical series beneath 4th toe of pes. Subcaudals on original tail roughly subequal. Dorsal pattern of three pairs of dark spots that may be fused into 8- shaped markings between limb insertions — one beginning behind forelimb insertion largest and the last ending just above the hindlimb insertion smallest; lighter interspaces narrower than largest spot and broader than smallest, often with a few small dark markings; broken elongate spots/streaks along flank; ground colouration the colour of milky tea in the faded
<typeStatus id="084E6EA4FF8CB910FE4DF85BFE36F81E" box="[402,500,2005,2030]" pageId="4" pageNumber="343" type="holotype">holotype</typeStatus>
and tan to brown in other specimens; dark streaks covering gular region.
</paragraph>
</subSubSection>
<caption id="838A808EFF8DB911FF48FBE1FC55FB11" httpUri="https://zenodo.org/record/259266/files/figure.png" pageId="5" pageNumber="344" targetBox="[150,1432,227,1099]" targetPageId="5">
<paragraph id="D74AD006FF8DB911FF48FBE1FC55FB11" blockId="5.[151,1436,1135,1249]" pageId="5" pageNumber="344">
<emphasis id="E5810C14FF8DB911FF48FBE1FED0FB74" bold="true" box="[151,274,1135,1157]" pageId="5" pageNumber="344">FIGURE 3.</emphasis>
Dorsal view of
<taxonomicName id="10F5AB85FF8DB911FE6AFBE1FD0FFB75" box="[437,717,1135,1157]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FE6AFBE1FD0FFB75" box="[437,717,1135,1157]" italics="true" pageId="5" pageNumber="344">Cyrtodactylus collegalensis</emphasis>
</taxonomicName>
(top row, AF) and
<taxonomicName id="10F5AB85FF8DB911FC44FBFEFBD8FB76" box="[923,1050,1136,1158]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="speciosus">
<emphasis id="E5810C14FF8DB911FC44FBFEFBD8FB76" box="[923,1050,1136,1158]" italics="true" pageId="5" pageNumber="344">C. speciosus</emphasis>
</taxonomicName>
(bottom row, GL). A, holotype of
<taxonomicName id="10F5AB85FF8DB911FA5BFBFEFED9FB54" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FA5BFBFEFED9FB54" italics="true" pageId="5" pageNumber="344">C. collegalensis</emphasis>
</taxonomicName>
(BMNH 1946.8.25.28); B, CES/09/1444; C, CES/09/1463; D, CES/09/1442; E, CES/09/1443; F,
<taxonomicName id="10F5AB85FF8DB911FA5BFB01FED9FB32" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FA5BFB01FED9FB32" italics="true" pageId="5" pageNumber="344">C. collegalensis</emphasis>
</taxonomicName>
, CES/09/1403. G, holotype of
<taxonomicName id="10F5AB85FF8DB911FD8BFB23FD16FB33" box="[596,724,1197,1219]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="speciosus">
<emphasis id="E5810C14FF8DB911FD8BFB23FD16FB33" box="[596,724,1197,1219]" italics="true" pageId="5" pageNumber="344">C. speciosus</emphasis>
</taxonomicName>
, BMNH 1946.9.4.88; H, presumed sketch of
<taxonomicName id="10F5AB85FF8DB911FB7BFB23FF17FB11" authority="Boulenger 1885" authorityName="Boulenger" authorityYear="1885" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="speciosus">
<emphasis id="E5810C14FF8DB911FB7BFB23FAE1FB33" box="[1188,1315,1197,1219]" italics="true" pageId="5" pageNumber="344">C. speciosus</emphasis>
(Boulenger 1885)
</taxonomicName>
; I, CES/09/1405; J, CES/09/1406; K, CES/09/1407; L, CES/09/1408.
</paragraph>
</caption>
<subSubSection id="9FEF838DFF8DB911FF18FA86FE7EF9ED" pageId="5" pageNumber="344" type="discussion">
<paragraph id="D74AD006FF8DB911FF18FA86FE7EF9ED" blockId="5.[151,1437,1288,1961]" pageId="5" pageNumber="344">
<emphasis id="E5810C14FF8DB911FF18FA86FE8DFAD1" bold="true" box="[199,335,1288,1313]" pageId="5" pageNumber="344">Comments.</emphasis>
Collections from around the
<typeStatus id="084E6EA4FF8DB911FD45FA87FD08FAD1" box="[666,714,1289,1313]" pageId="5" pageNumber="344">type</typeStatus>
locality resulted in the rediscovery of this species as well as the discovery of a moderately genetically divergent specimen with a drastically different colour pattern from an isolated hill. Further sampling is needed to ascertain if that is an aberrant individual,
<taxonomicName id="10F5AB85FF8DB911FBBBFADFFA5EFA99" box="[1124,1436,1361,1385]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FBBBFADFFA5EFA99" box="[1124,1436,1361,1385]" italics="true" pageId="5" pageNumber="344">Cyrtodactylus collegalensis</emphasis>
</taxonomicName>
shows clinal geographic variation, or if that population is truly divergent. This species is variable in pattern, with stippling on the lighter interspaces between spots absent in a few specimens, the collar separated from the postorbital streak in CES/09/1403, dorsal pairs of spots fused occasionally. Interestingly, CES/09/1442 and CES/ 09/1443 that have identical ND2 sequences have different head markings, body colouration and vary in the degree of fusion of spots (
<figureCitation id="4FCECC83FF8DB911FEB6F98BFE6CF9ED" box="[361,430,1541,1566]" captionStart="FIGURE 3" captionStartId="5.[151,250,1135,1157]" captionTargetBox="[150,1432,227,1099]" captionTargetId="figure@5.[147,1432,227,1100]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of Cyrtodactylus collegalensis (top row, A F) and C. speciosus (bottom row, G L). A, holotype of C. collegalensis (BMNH 1946.8.25.28); B, CES / 09 / 1444; C, CES / 09 / 1463; D, CES / 09 / 1442; E, CES / 09 / 1443; F, C. collegalensis, CES / 09 / 1403. G, holotype of C. speciosus, BMNH 1946.9.4.88; H, presumed sketch of C. speciosus (Boulenger 1885); I, CES / 09 / 1405; J, CES / 09 / 1406; K, CES / 09 / 1407; L, CES / 09 / 1408." httpUri="https://zenodo.org/record/259266/files/figure.png" pageId="5" pageNumber="344">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9FEF838DFF8DB911FF18F9A6FAE0F858" pageId="5" pageNumber="344" type="diagnosis">
<paragraph id="D74AD006FF8DB911FF18F9A6FAE0F858" blockId="5.[151,1437,1288,1961]" pageId="5" pageNumber="344">
<emphasis id="E5810C14FF8DB911FF18F9A6FE83F9B1" bold="true" box="[199,321,1576,1601]" pageId="5" pageNumber="344">Diagnosis.</emphasis>
<taxonomicName id="10F5AB85FF8DB911FE97F9A7FDBDF9B1" box="[328,639,1577,1601]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FE97F9A7FDBDF9B1" box="[328,639,1577,1601]" italics="true" pageId="5" pageNumber="344">Cyrtodactylus collegalensis</emphasis>
</taxonomicName>
(and members of the
<taxonomicName id="10F5AB85FF8DB911FCA8F9A7FBE9F9B1" box="[887,1067,1577,1601]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FCA8F9A7FBE9F9B1" box="[887,1067,1577,1601]" italics="true" pageId="5" pageNumber="344">C. collegalensis</emphasis>
</taxonomicName>
complex) can be separated from all other
<taxonomicName id="10F5AB85FF8DB911FF21F9C3FE59F995" box="[254,411,1613,1637]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="genus">
<emphasis id="E5810C14FF8DB911FF21F9C3FE59F995" box="[254,411,1613,1637]" italics="true" pageId="5" pageNumber="344">Cyrtodactylus</emphasis>
</taxonomicName>
species by entirely homogeneous dorsal scalation, the absence of a precloacal groove, no enlarged precloacal or femoral scales, and no precloacal or femoral pores.
<taxonomicName id="10F5AB85FF8DB911FC3FF9FFFB51F979" box="[992,1171,1649,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FC3FF9FFFB51F979" box="[992,1171,1649,1673]" italics="true" pageId="5" pageNumber="344">C. collegalensis</emphasis>
</taxonomicName>
and members of the
<taxonomicName id="10F5AB85FF8DB911FA5DF9FFFEEBF95D" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FA5DF9FFFEEBF95D" italics="true" pageId="5" pageNumber="344">C. collegalensis</emphasis>
</taxonomicName>
complex differ from other
<taxonomicName id="10F5AB85FF8DB911FD82F91BFD0DF95D" box="[605,719,1685,1709]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="genus">
<emphasis id="E5810C14FF8DB911FD82F91BFD0DF95D" box="[605,719,1685,1709]" italics="true" pageId="5" pageNumber="344">Geckoella</emphasis>
</taxonomicName>
species in homogeneous dorsal scalation that consists of small, granular scales (vs. large flattened or granular dorsal scales, interspersed with a few or numerous tubercles in
<emphasis id="E5810C14FF8DB911FA5DF937FE21F905" italics="true" pageId="5" pageNumber="344">
<taxonomicName id="10F5AB85FF8DB911FA5DF937FEEDF905" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="albofasciatus">C. albofasciatus</taxonomicName>
,
<taxonomicName id="10F5AB85FF8DB911FEE4F953FE21F905" box="[315,483,1757,1781]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="deccanensis">C. deccanensis</taxonomicName>
</emphasis>
and
<taxonomicName id="10F5AB85FF8DB911FDC5F953FD56F905" box="[538,660,1757,1781]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="triedrus">
<emphasis id="E5810C14FF8DB911FDC5F953FD56F905" box="[538,660,1757,1781]" italics="true" pageId="5" pageNumber="344">C. triedrus</emphasis>
</taxonomicName>
; enlarged, hexagonal scales in
<taxonomicName id="10F5AB85FF8DB911FC2DF953FB53F905" box="[1010,1169,1757,1781]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="jeyporensis">
<emphasis id="E5810C14FF8DB911FC2DF953FB53F905" box="[1010,1169,1757,1781]" italics="true" pageId="5" pageNumber="344">C. jeyporensis</emphasis>
</taxonomicName>
; tubercles present in
<taxonomicName id="10F5AB85FF8DB911FA5DF953FEC5F8E9" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="E5810C14FF8DB911FA5DF953FEC5F8E9" italics="true" pageId="5" pageNumber="344">C. nebulosus</emphasis>
</taxonomicName>
; presence of few enlarged scales near hind-limbs in
<taxonomicName id="10F5AB85FF8DB911FC53F88FFBD0F8E9" box="[908,1042,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="yakhuna">
<emphasis id="E5810C14FF8DB911FC53F88FFBD0F8E9" box="[908,1042,1793,1817]" italics="true" pageId="5" pageNumber="344">C. yakhuna</emphasis>
</taxonomicName>
).
<taxonomicName id="10F5AB85FF8DB911FBF0F88FFA82F8E9" box="[1071,1344,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FBF0F88FFA82F8E9" box="[1071,1344,1793,1817]" italics="true" pageId="5" pageNumber="344">Geckoella collegalensis</emphasis>
</taxonomicName>
can be additionally be differentiated from the
<collectingRegion id="15311EE4FF8DB911FD94F8ABFDAEF8CE" box="[587,620,1829,1854]" country="United Kingdom" name="Surrey" pageId="5" pageNumber="344">Sri</collectingRegion>
Lankan
<taxonomicName id="10F5AB85FF8DB911FD11F8ABFC8CF8CD" box="[718,846,1829,1853]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="yakhuna">
<emphasis id="E5810C14FF8DB911FD11F8ABFC8CF8CD" box="[718,846,1829,1853]" italics="true" pageId="5" pageNumber="344">C. yakhuna</emphasis>
</taxonomicName>
by dorsal colour pattern (three rows of paired spots/ 8-shaped markings between limb insertions, equal to or wider than interspaces vs. one or two rows of spots/bands, equal to or narrower than interspaces).
<taxonomicName id="10F5AB85FF8DB911FDBCF8E3FC5FF875" box="[611,925,1901,1925]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="collegalensis">
<emphasis id="E5810C14FF8DB911FDBCF8E3FC5FF875" box="[611,925,1901,1925]" italics="true" pageId="5" pageNumber="344">Cyrtodactylus collegalensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="10F5AB85FF8DB911FB5FF8E3FACDF875" box="[1152,1295,1901,1925]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="speciosus">
<emphasis id="E5810C14FF8DB911FB5FF8E3FACDF875" box="[1152,1295,1901,1925]" italics="true" pageId="5" pageNumber="344">C. speciosus</emphasis>
</taxonomicName>
, but can be differentiated by a number of characters which are detailed after the systematic account of
<taxonomicName id="10F5AB85FF8DB911FB50F81FFADEF858" box="[1167,1308,1937,1960]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="344" phylum="Chordata" rank="species" species="speciosus">
<emphasis id="E5810C14FF8DB911FB50F81FFADEF858" box="[1167,1308,1937,1960]" italics="true" pageId="5" pageNumber="344">C. speciosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>