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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.221.3234" ID-GBIF-Dataset="b6521187-a030-4b18-8f9d-f35084c9336a" ID-PMC="PMC3487634" ID-Pensoft-Pub="1313-2970-221-31" ID-PubMed="23129980" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-221-31" ModsDocOrigin="ZooKeys 221" ModsDocTitle="A revision of the Megachile subgenus Litomegachile Mitchell with an illustrated key and description of a new species (Hymenoptera, Megachilidae, Megachilini)" checkinTime="1451248746570" checkinUser="pensoft" docAuthor="Bzdyk, Emily L." docDate="2012" docId="5688FCCE85727B7875378FF3C1B3A4FB" docLanguage="en" docName="ZooKeys 221: 31-61" docOrigin="ZooKeys 221" docSource="http://dx.doi.org/10.3897/zookeys.221.3234" docTitle="Megachile (Litomegachile) pankus Bzdyk, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="48" masterDocId="CB68FFB2082C476FFFF5FFD7FFFB0B6B" masterDocTitle="A revision of the Megachile subgenus Litomegachile Mitchell with an illustrated key and description of a new species (Hymenoptera, Megachilidae, Megachilini)" masterLastPageNumber="61" masterPageNumber="31" pageNumber="46" updateTime="1668154456179" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Megachile subgenus Litomegachile Mitchell with an illustrated key and description of a new species (Hymenoptera, Megachilidae, Megachilini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bzdyk, Emily L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>221</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>61</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.221.3234</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.221.3234</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-221-31</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152037030" LSID="urn:lsid:zoobank.org:act:80ED5270-BA6B-42C8-AB4B-AEFA7A531D7A" httpUri="http://treatment.plazi.org/id/5688FCCE85727B7875378FF3C1B3A4FB" lastPageId="17" lastPageNumber="48" pageId="15" pageNumber="46">
<subSubSection pageId="15" pageNumber="46" type="nomenclature">
<paragraph pageId="15" pageNumber="46">
<taxonomicName LSID="urn:lsid:zoobank.org:act:80ED5270-BA6B-42C8-AB4B-AEFA7A531D7A" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile (Litomegachile) pankus" order="Hymenoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="pankus" subGenus="Litomegachile">Megachile (Litomegachile) pankus</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="46">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="46" type="type material">
<paragraph pageId="15" pageNumber="46">Type material.</paragraph>
<paragraph pageId="15" pageNumber="46">Holotype female: MEXICO: Hidalgo, Pachuca, 11 Jun 1935, R. M. and G. E. Bohart (BBSL). Paratypes: 1 female: MEXICO: Sonora, Alamos, 4 Sep 1991 (AMNH), 1 female: MEXICO: Sinaloa, Mazatlan 28 Oct 1969 (BBSL); 1 female: MEXICO: Sinaloa, 4 mi NW Choix, 31 Aug 1968 (BMEC); 1 female: MEXICO: Sinaloa, 6 mi NW Choix, 6 Aug 1968 (BMEC).</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="47" pageId="15" pageNumber="46" type="diagnosis">
<paragraph pageId="15" pageNumber="46">Diagnosis.</paragraph>
<paragraph lastPageId="16" lastPageNumber="47" pageId="15" pageNumber="46">
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile pankus" order="Hymenoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="pankus">Megachile pankus</taxonomicName>
is unique among
<taxonomicName class="Insecta" family="Megachilidae" genus="Litomegachile" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Litomegachile" order="Hymenoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">Litomegachile</taxonomicName>
species because the female has a mandible with an angulation between teeth 3 and 4, and T6 is basally convex and apically concave. No other species in the subgenus has this combination of characters. The female
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile onobrychidis" order="Hymenoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="onobrychidis">Megachile onobrychidis</taxonomicName>
has similar metasomal features, but has more
<pageBreakToken pageId="16" pageNumber="47" start="start">black</pageBreakToken>
setae on S6, while
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile pankus" order="Hymenoptera" pageId="16" pageNumber="47" phylum="Arthropoda" rank="species" species="pankus">Megachile pankus</taxonomicName>
has only a few black setae on T6. It can also be further distinguished from
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile onobrychidis" order="Hymenoptera" pageId="16" pageNumber="47" phylum="Arthropoda" rank="species" species="onobrychidis">Megachile onobrychidis</taxonomicName>
and
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile brevis" order="Hymenoptera" pageId="16" pageNumber="47" phylum="Arthropoda" rank="species" species="brevis">Megachile brevis</taxonomicName>
by the angulation between teeth 3 and 4 of the mandible. T6 is convex basally and concave apically in profile, and concave laterally in dorsal view, which distinguishes it from
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile mendica" order="Hymenoptera" pageId="16" pageNumber="47" phylum="Arthropoda" rank="species" species="mendica">Megachile mendica</taxonomicName>
or
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile gentilis" order="Hymenoptera" pageId="16" pageNumber="47" phylum="Arthropoda" rank="species" species="gentilis">Megachile gentilis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="48" pageId="16" pageNumber="47" type="female description">
<paragraph pageId="16" pageNumber="47">Female description.</paragraph>
<paragraph lastPageId="17" lastPageNumber="48" pageId="16" pageNumber="47">
Body length 10 mm. Forewing length 7 mm. Head: HL 0.7
<normalizedToken originalValue="×">x</normalizedToken>
HW; compound eyes convergent below, with upper inner margins slightly convergent above; lateral ocelli closer to margin of vertex than edge of compound eye; compound eye width 1.1
<normalizedToken originalValue="×">x</normalizedToken>
width of genal area in lateral view; clypeus twice as wide as high (Figure 3B); clypeus and supraclypeal area slightly convex; punctation fine, with larger punctures on clypeus, becoming smaller on supraclypeal, paraocular area, vertex and rest of head; punctures never separated by more than 0.3
<normalizedToken originalValue="×">x</normalizedToken>
puncture diameter; labrum width 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length; AD 3
<normalizedToken originalValue="×">x</normalizedToken>
width of antennal socket, ID 0.6
<normalizedToken originalValue="×">x</normalizedToken>
ASO; ID 1.36
<normalizedToken originalValue="×">x</normalizedToken>
length of scape; mandible with recessed cutting edges between teeth 3 and 4 and incomplete recessed cutting edge that forms rough right angle between teeth 3 and 2; surface between teeth 3 and 4 angulate (Figure 2C, 4B); scape length 4.3
<normalizedToken originalValue="×">x</normalizedToken>
width, with white setae; pedicel and F1 width 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length; pedicel length 0.8
<normalizedToken originalValue="×">x</normalizedToken>
F1; F2-6 length equal to width; F7-8 length 0.9
<normalizedToken originalValue="×">x</normalizedToken>
width; F9 length 0.8
<normalizedToken originalValue="×">x</normalizedToken>
width; F10 length 0.7
<normalizedToken originalValue="×">x</normalizedToken>
width; Mesosoma: mesepisternum convex, large and pronounced, twice as wide as pronotum; scutum length 0.8
<normalizedToken originalValue="×">x</normalizedToken>
width; scutellum length 0.3
<normalizedToken originalValue="×">x</normalizedToken>
scutum length, scutellum width 0.4
<normalizedToken originalValue="×">x</normalizedToken>
scutellum length; tegula twice as long as wide; scutum 7.2
<normalizedToken originalValue="×">x</normalizedToken>
width of tegula. Wings: forewing length 2.7
<normalizedToken originalValue="×">x</normalizedToken>
width; WCL 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of wing; SL 0.2
<normalizedToken originalValue="×">x</normalizedToken>
MCL; with two submarginal cells, first submar
<pageBreakToken pageId="17" pageNumber="48" start="start">ginal</pageBreakToken>
crossvein angled parallel to medial vein, second submarginal crossvein angulate; distance from distal edge of stigma to wing base 0.7
<normalizedToken originalValue="×">x</normalizedToken>
distance from wing base to distal edge of marginal cell; hindwing with jugal lobe that does not extend past cubital cell; LTJ 0.3
<normalizedToken originalValue="×">x</normalizedToken>
HWL; LTV 0.5
<normalizedToken originalValue="×">x</normalizedToken>
HWL (Figure 3A). Legs: ratio of segment length of foreleg (compared to FL): CL 0.6
<normalizedToken originalValue="×">x</normalizedToken>
, TL 0.3
<normalizedToken originalValue="×">x</normalizedToken>
, FL 1
<normalizedToken originalValue="×">x</normalizedToken>
, TBL 0.9
<normalizedToken originalValue="×">x</normalizedToken>
, TRL 1.2
<normalizedToken originalValue="×">x</normalizedToken>
, BTL 0.5
<normalizedToken originalValue="×">x</normalizedToken>
, DTL 0.3
<normalizedToken originalValue="×">x</normalizedToken>
; foreleg with tibial spur modified as antennal cleaner, TSL 0.2
<normalizedToken originalValue="×">x</normalizedToken>
; midleg segment ratios: CL 0.7
<normalizedToken originalValue="×">x</normalizedToken>
, TL 0.4
<normalizedToken originalValue="×">x</normalizedToken>
, FL 1
<normalizedToken originalValue="×">x</normalizedToken>
, TBL 1
<normalizedToken originalValue="×">x</normalizedToken>
, TRL 1.3
<normalizedToken originalValue="×">x</normalizedToken>
, BTL 0.7
<normalizedToken originalValue="×">x</normalizedToken>
, DTL 0.3
<normalizedToken originalValue="×">x</normalizedToken>
; foreleg with tibial spur, TSL 0.3
<normalizedToken originalValue="×">x</normalizedToken>
TBL; hindleg segment ratios: CL 0.5
<normalizedToken originalValue="×">x</normalizedToken>
, TL 0.3
<normalizedToken originalValue="×">x</normalizedToken>
, FL 1
<normalizedToken originalValue="×">x</normalizedToken>
, TBL 0.9
<normalizedToken originalValue="×">x</normalizedToken>
, TRL 1.4
<normalizedToken originalValue="×">x</normalizedToken>
, BTL 0.7
<normalizedToken originalValue="×">x</normalizedToken>
, DTL 0.3
<normalizedToken originalValue="×">x</normalizedToken>
; tibia with two spurs, TSL 0.4
<normalizedToken originalValue="×">x</normalizedToken>
TBL; hindleg with basitarsus dilated 4.5
<normalizedToken originalValue="×">x</normalizedToken>
width of distitarsus (Figure 3C). Metasoma: T2-4 with shallow transverse basal grooves; T1-5 with apical fringes of white hair covering marginal zone, T1-2 fringe widths 0.2
<normalizedToken originalValue="×">x</normalizedToken>
width of discal surface medially, T3-5 fringe widths 0.3a width of discal surface medially; T1-2 apical fringes of white hair more sparse, marginal zone slightly visible between hairs; T1-5 with white discal pubescence; T6 discal surface with black appressed pubescence and black erect setae; T6 convex basally and concave apically in profile, and concave laterally in dorsal view;S1-5 with yellow setae; S6 with yellow setae and some black setae apically (Figure 5F). Color: Body black, legs brownish distally, wing membrane slightly tinted brown, veins brown (Figure 2A-B). Pubescence: White on head except ocellar region black; paraocular area, supraclypeal area and clypeus with dense pubescence obscuring view of integument; vertex with sparse pubescence with integument visible beneath; genal area with pubescence sparse beginning at dorsal surface, progressively more dense toward malar area. Mesosomal pubescence white; dense around tegula and behind scutellum, sparse on scutum, dense on ventral mesosomal surface.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="48" type="male">
<paragraph pageId="17" pageNumber="48">Male.</paragraph>
<paragraph pageId="17" pageNumber="48">unknown</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="48" type="etymology">
<paragraph pageId="17" pageNumber="48">Etymology.</paragraph>
<paragraph pageId="17" pageNumber="48">
The species name
<normalizedToken originalValue="pankus">'pankus'</normalizedToken>
is a nonsense combination.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="48" type="distribution">
<paragraph pageId="17" pageNumber="48">Distribution.</paragraph>
<paragraph pageId="17" pageNumber="48">
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile pankus" order="Hymenoptera" pageId="17" pageNumber="48" phylum="Arthropoda" rank="species" species="pankus">Megachile pankus</taxonomicName>
has only been collected in Mexico (Figure 14).
</paragraph>
<caption pageId="17" pageNumber="48">
<paragraph pageId="17" pageNumber="48">
Figure 14. Distribution of
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile pankus" order="Hymenoptera" pageId="17" pageNumber="48" phylum="Arthropoda" rank="species" species="pankus">Megachile pankus</taxonomicName>
(red indicates locality of holotype).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="17" pageNumber="48" type="flower records">
<paragraph pageId="17" pageNumber="48">Flower records.</paragraph>
<paragraph pageId="17" pageNumber="48">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Petalostemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petalostemon" order="Fabales" pageId="17" pageNumber="48" phylum="Tracheophyta" rank="genus">Petalostemon</taxonomicName>
sp. (
<taxonomicName genus="Fabaceae" lsidName="Fabaceae" pageId="17" pageNumber="48" rank="genus">Fabaceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>