794 lines
78 KiB
XML
794 lines
78 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.886.38511" ID-GBIF-Dataset="92414cf7-8f9d-40bf-96aa-1999f0f5e80e" ID-PMC="PMC6851206" ID-Pensoft-Pub="1313-2970-886-1" ID-Pensoft-UUID="F8981F284ACA5F88ADEC048F0DA101FA" ID-PubMed="31736621" ID-ZooBank="811E2B80FEC6419B8F3772000C68BD1E" ModsDocID="1313-2970-886-1" checkinTime="1573015838763" checkinUser="pensoft" docAuthor="Jung, Tae Won, Coleman, Charles Oliver & Yoon, Seong Myeong" docDate="2019" docId="93FECD02FCD5588CADADC77C85642F46" docLanguage="en" docName="ZooKeys 886: 1-59" docOrigin="ZooKeys 886" docSource="http://dx.doi.org/10.3897/zookeys.886.38511" docTitle="Photis bronca Jung, Coleman & Yoon, 2019, sp. nov." docType="treatment" docUuid="CE27564C-91FB-47A7-8500-9740799302A0" docUuidSource="ZooBank" docVersion="5" id="F8981F284ACA5F88ADEC048F0DA101FA" lastPageNumber="1" masterDocId="F8981F284ACA5F88ADEC048F0DA101FA" masterDocTitle="Taxonomic study on the photid amphipods (Senticaudata, Corophiida, Photoidea, Photidae) from Korean waters, with descriptions of a new genus and seven new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="1" updateTime="1668168022767" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic study on the photid amphipods (Senticaudata, Corophiida, Photoidea, Photidae) from Korean waters, with descriptions of a new genus and seven new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jung, Tae Won</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Coleman, Charles Oliver</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoon, Seong Myeong</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>886</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>59</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.886.38511</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.886.38511</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-886-1</mods:identifier>
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<mods:identifier type="ZooBank">811E2B80FEC6419B8F3772000C68BD1E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F8981F284ACA5F88ADEC048F0DA101FA</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="160140693" LSID="urn:lsid:zoobank.org:act:CE27564C-91FB-47A7-8500-9740799302A0" httpUri="http://treatment.plazi.org/id/93FECD02FCD5588CADADC77C85642F46" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="93FECD02-FCD5-588C-ADAD-C77C85642F46" class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">Photis bronca</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Figs 8</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A maxilliped B gnathopod 1 C gnathopod 2 D pereopod 3 E coxa of pereopod 3 F pereopod 4. Scale bars: 0.1 mm (A), 0.2 mm (B-F)." figureDoi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure9">9</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">10</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Photis bronca sp. nov., paratype, NIBRIV 0000848473, female, 5.3 mm, Daryeo-do Island, South Korea. A gnathopod 1 B gnathopod 2 C coxa and oostegite of pereopod 3 D pereopod 4 E oostegite of pereopod 5. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure11">11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis longicaudata" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longicaudata">Photis longicaudata</taxonomicName>
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:
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<bibRefCitation author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" pagination="131 - 170" publicationUrl="http://hdl.handle.net/2433/175398" refId="B36" refString="Nagata, K, 1965. Studies on marine gammaridean Amphipoda of the Seto Inland Sea. I. Publications of the Seto Marine Biological Laboratory 13: 131 - 170" title="Studies on marine gammaridean Amphipoda of the Seto Inland Sea. I." url="http://hdl.handle.net/2433/175398" volume="13" year="1965">Nagata 1965</bibRefCitation>
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: 310, fig. 35;
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<bibRefCitation DOI="https://doi.org/10.5134/176060" author="Hirayama, A" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" pagination="75 - 150" refId="B21" refString="Hirayama, A, 1984. Taxonomic studies on the shallow water gammaridean Amphipoda of the West Kyushu, Japan. II. Corophiidae. Publications of the Seto Marine Biological Laboratory 28: 75 - 150" title="Taxonomic studies on the shallow water gammaridean Amphipoda of the West Kyushu, Japan. II. Corophiidae." url="https://doi.org/10.5134/176060" volume="28" year="1984">Hirayama 1984</bibRefCitation>
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: 42, fig. 71;
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<bibRefCitation author="Kim, Y-H" journalOrPublisher="Korean Journal of Systematic Zoology Special issue" pageId="0" pageNumber="1" pagination="1 - 11" publicationUrl="http://www.e-ased.org/PublishedPaper/before_list.asp" refId="B23" refString="Kim, Y-H, Eun, Y, Lee, K-S, 2005. Amphipods (Gammaridea and Caprellidea) fauna of Jindo Island in Korea. Korean Journal of Systematic Zoology Special issue 5: 1 - 11" title="Amphipods (Gammaridea and Caprellidea) fauna of Jindo Island in Korea." url="http://www.e-ased.org/PublishedPaper/before_list.asp" volume="5" year="2005">Kim et al. 2005</bibRefCitation>
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: 3, figs 1, 2;
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<bibRefCitation author="Ren, XQ" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" refId="B37" refString="Ren, XQ, 1992. Studies on the Gammaridea (Crustacea: Amphipoda) from Jiaozhou Bay (Yellow sea). In: Transactions of the Chinese Crustacean Society Vol. 3. Qingdao Ocean University Press, Qingdao, 214-317." title="Studies on the Gammaridea (Crustacea: Amphipoda) from Jiaozhou Bay (Yellow sea). In: Transactions of the Chinese Crustacean Society Vol. 3. Qingdao Ocean University Press, Qingdao, 214 - 317." year="1992">Ren 1992</bibRefCitation>
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: 259, fig. 28; 2000: 141, fig. 4; 2006: 398, fig. 171 [non
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<taxonomicName family="Photidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. longicaudata" order="Amphipoda" pageId="0" pageNumber="1" rank="species" species="longicaudata">P. longicaudata</taxonomicName>
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(Spence Bate & Westwood, 1862)].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The epithet of the specific name,
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<emphasis italics="true" pageId="0" pageNumber="1">bronca</emphasis>
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, has its origin from the Latin word
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<emphasis italics="true" pageId="0" pageNumber="1">broncus</emphasis>
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. This name refers to the bearing of a quadrate tooth medially on the gnathopod 2 palmar margin in both sexes.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype: ♂ (5.8 mm), NIBRIV0000848472; Daryeo-do Island, Bukchon, Jeju-do, South Korea (
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<geoCoordinate degrees="34" direction="north" minutes="33" orientation="latitude" precision="15" seconds="27" value="34.5575">34°33'27"N</geoCoordinate>
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,
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<geoCoordinate degrees="126" direction="east" minutes="41" orientation="longitude" precision="15" seconds="49" value="126.696945">126°41'49"E</geoCoordinate>
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); 30 Nov, 2012; grab sampler (about 20 m depth), by Prof. HY Soh. Paratypes: 2 ♂♂ (2.3 and 2.5 mm), 3 ♀♀ (ovigerous; 3.2-5.3 mm), NIBRIV0000848473; same data as holotype.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Additional materials: 3 ♂♂ 2 ♀♀, NIBRIV0000848930, Saekkiseom, Ae-do Island, Mijo-myeon, Gyeongsangnam-do, South Korea (
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<geoCoordinate degrees="34" direction="north" minutes="41" orientation="latitude" precision="15" seconds="31" value="34.691944">34°41'31"N</geoCoordinate>
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,
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<geoCoordinate degrees="128" direction="east" minutes="02" orientation="longitude" precision="15" seconds="00" value="128.03334">128°02'00"E</geoCoordinate>
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), 1 May 2009, Scuba diving (about 20 m depth), by TW Jung; 2 ♂♂, NIBRIV0000848931, Jeolmyeongyeo, Chuja-do Island, Jeju-do, South Korea (
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<geoCoordinate degrees="33" direction="north" minutes="52" orientation="latitude" precision="15" seconds="04" value="33.867775">33°52'04"N</geoCoordinate>
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,
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<geoCoordinate degrees="126" direction="east" minutes="18" orientation="longitude" precision="15" seconds="43" value="126.31195">126°18'43"E</geoCoordinate>
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), 12 Jul, 2016, Scuba diving (about 16 m depth), by CH Yi; 3 ♂♂ 2 ♀♀, NIBRIV0000848932, Yenae-ri, Goheung-gun, Jeollanam-do, South Korea (
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<geoCoordinate degrees="34" direction="north" minutes="27" orientation="latitude" precision="15" seconds="24" value="34.45667">34°27'24"N</geoCoordinate>
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,
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<geoCoordinate degrees="127" direction="east" minutes="31" orientation="longitude" precision="15" seconds="16" value="127.52112">127°31'16"E</geoCoordinate>
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), 28 Mar 2017, washing macro-algae, by SH Kim.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">In both sexes, gnathopod 2 propodus palmar margin with subrectangular cavity bearing one rectangular tooth medially. Male gnathopod 2 basis lateral margin forming a well-developed sac-like lobe. Stridulated ridges only present on male gnathopod 2 basis and coxae 3 and 4 (absent in females).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Holotype male.</emphasis>
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Head (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8A</figureCitation>
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) as long as pereonites 1 and 2 combined; lateral cephalic lobe rounded; eye circular, moderate in size, located in the middle of lateral lobe; antennal sinus deep.
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</paragraph>
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<caption doi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
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<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
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<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
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</taxonomicName>
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sp. nov., holotype, NIBRIV0000848472, male, 5.8 mm, Daryeo-do Island, South Korea.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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habitus
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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antenna 1
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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antenna 2
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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upper lip
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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lower lip
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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left mandible
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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right mandible
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
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maxilla 1
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<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
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maxilla 2. Scale bars: 0.1 mm (
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<emphasis bold="true" pageId="0" pageNumber="1">
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<normalizedToken originalValue="D–I">D-I</normalizedToken>
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</emphasis>
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), 0.2 mm (
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<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
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), 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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Antenna 1 (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8B</figureCitation>
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) 0.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as body; peduncle 1st article stout, 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as head; 2nd article slender, 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as 1st article; 3rd article 0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as 2nd article; accessory flagellum absent; flagellum 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as peduncle 1
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<normalizedToken originalValue="st–">st-</normalizedToken>
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3rd articles combined, composed of ten articles (terminal article rudimentary).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Antenna 2 (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8C</figureCitation>
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) 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as antenna 1; peduncle 3th article reaching end of lateral cephalic lobe; 4th article as long as 2nd article of antenna 1; 5th article as long as 4th article; flagellum 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as peduncle 3
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<normalizedToken originalValue="rd–">rd-</normalizedToken>
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5th articles combined, composed of ten articles (terminal article rudimentary).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Upper lip (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8D</figureCitation>
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) convex anteriorly, with notch in the middle, covered with minute setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Lower lip (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8E</figureCitation>
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) inner lobe subovoid, outer lobe apex rounded, covered with minute setae, with one robust seta at mediodistal corner; mandibular process well developed.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Mandibles (
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8F, G</figureCitation>
|
||
) with 4-dentate incisor, 4-dentate lacinia mobilis, and four raker setae on left mandible; with 5-dentate incisor, minutely dentate lacinia mobilis, and three raker setae on right mandible; molar well developed, triturative, with seven setae along the distal margin of right mandible; palp asymmetrical, composed of three articles, 3rd article distally rounded, 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as 2nd article, with setae extending along most of posteriodistal margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Maxilla 1 (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8H</figureCitation>
|
||
) inner lobe small, without minute setae; outer lobe with eight dentate robust setae on apical margin; palp biarticulated, distal article curved, with four dentate setae on apical margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Maxilla 2 (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A habitus B antenna 1 C antenna 2 D upper lip E lower lip F left mandible G right mandible H maxilla 1 I maxilla 2. Scale bars: 0.1 mm (D-I), 0.2 mm (B, C), 0.5 mm (A)." figureDoi="10.3897/zookeys.886.38511.figure8" httpUri="https://binary.pensoft.net/fig/352720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure8">Fig. 8I</figureCitation>
|
||
) inner lobe with an oblique row of plumose setae on surface; outer lobe longer and slightly dilated distally than inner lobe.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Maxilliped (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A maxilliped B gnathopod 1 C gnathopod 2 D pereopod 3 E coxa of pereopod 3 F pereopod 4. Scale bars: 0.1 mm (A), 0.2 mm (B-F)." figureDoi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure9">Fig. 9A</figureCitation>
|
||
) inner lobe subrectangular, a little expanded distally, with three nodular setae apically and one medial nodular seta subdistally; outer lobe exceeding half of palp 2nd article, lined with seven robust setae along apex to medial margin; palp composed of four articles, 3rd article slightly expanded distally, half as long as 2nd article, 4th article 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as 3rd article, with elongate seta apically (as long as 4th article).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, NIBRIV0000848472, male, 5.8 mm, Daryeo-do Island, South Korea.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
maxilliped
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
gnathopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
gnathopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
pereopod 3
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
coxa of pereopod 3
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
pereopod 4. Scale bars: 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), 0.2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<normalizedToken originalValue="B–F">B-F</normalizedToken>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Gnathopod 1 (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A maxilliped B gnathopod 1 C gnathopod 2 D pereopod 3 E coxa of pereopod 3 F pereopod 4. Scale bars: 0.1 mm (A), 0.2 mm (B-F)." figureDoi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure9">Fig. 9B</figureCitation>
|
||
) coxa subtrapezoidal, slightly extended distally, 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long; basis as long as coxa, anterior margin lateral border forming weak lobe distally, with three elongate setae subproximally, posterior margin with three elongate setae at the middle; carpus subtrapezoidal, 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis, half as wide as long, posterior lobe blunt; propodus as long and wide as carpus, rounded anteriorly, with minute serrations irregularly along palm and posterior margin, palm 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as posterior margin, defined by one robust seta medially; dactylus 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus, exceeding palm, inner margin serrated, with two teeth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Gnathopod 2 (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A maxilliped B gnathopod 1 C gnathopod 2 D pereopod 3 E coxa of pereopod 3 F pereopod 4. Scale bars: 0.1 mm (A), 0.2 mm (B-F)." figureDoi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure9">Fig. 9C</figureCitation>
|
||
) stout, coxa subrectangular, 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, slightly extended and rounded anterioventrally; basis anterior margin lateral borders forming well-developed lobe distally (sac-like lobe reaching middle of merus) bearing oblique stridulated ridges on surface; ischium with small anterior lobe only; merus subrectangular, 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis; carpus stout, anterior margin convex, carpal lobe well developed; propodus stout, as long as basis, 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, slightly widened distally, palmar margin defined by one large blunt spine, with small obtuse spine near dactylus base and robust seta near defining spine, concave subrectangularly, bearing quadrate tooth medially; dactylus half as long as propodus, with two teeth on inner margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 3 (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A maxilliped B gnathopod 1 C gnathopod 2 D pereopod 3 E coxa of pereopod 3 F pereopod 4. Scale bars: 0.1 mm (A), 0.2 mm (B-F)." figureDoi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure9">Fig. 9D, E</figureCitation>
|
||
) coxa widened distally, produced anterioventrally, 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, with stridulated ridges on medial surface and short stridulated ridges near the posterioventral corner on lateral surface submarginally; basis 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, with three plumose setae along distal half of anterior margin, posterior margin expanded, evenly rounded, with two elongate setae at the middle, with a pair of plumose and minute setae at distal corner; merus 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis, anterior margin weakly expanded, with two plumose setae submarginally, distal corner somewhat produced, with one plumose seta subapically; carpus subrectangular, half as long as and 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as merus; propodus slender, diminished distally, 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis; dactylus 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 4 (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A maxilliped B gnathopod 1 C gnathopod 2 D pereopod 3 E coxa of pereopod 3 F pereopod 4. Scale bars: 0.1 mm (A), 0.2 mm (B-F)." figureDoi="10.3897/zookeys.886.38511.figure9" httpUri="https://binary.pensoft.net/fig/352721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure9">Fig. 9F</figureCitation>
|
||
) coxa slightly widened distally, as long as that of pereopod 3, with stridulated ridges along anterioventral corner more oblique than that of female; basis 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, with one plumose seta at the middle of anterior margin, posterior margin expanded, evenly rounded, with three elongate setae at the middle, with three plumose setae distally; merus 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis, anterior margin weakly expanded, distal corner somewhat produced; carpus half as long as and 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as merus, propodus slender, diminished distally, 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis; dactylus 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 5 (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10A</figureCitation>
|
||
) coxa bilobed, large, anterior lobe subovoid, 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide, expanded ventrally, posterior lobe small, expanded backwards; basis subovoid, broad, more expanded proximally, 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long; merus subrectangular, half as long as basis, half as wide as long; carpus subrectangular, as long and wide as merus; propodus 1.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as carpus, with a pair of distal locking setae unequal in length (longer seta 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as dactylus), with a group of four setae (longest seta 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus) at anteriodistal corner; dactylus 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus, armed with one accessory cusp on outer margin.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, NIBRIV0000848472, male, 5.8 mm, Daryeo-do Island, South Korea.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
pereopod 5
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
, pereopod 6
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
dactylus of pereopod 6
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
coxa of pereopod 7
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
pereopod 7
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
uropod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
uropod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||
uropod 3 and telson. Scale bars: 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
C, E,
|
||
<normalizedToken originalValue="G–I">G-I</normalizedToken>
|
||
</emphasis>
|
||
), 0.2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, B, D, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 6 (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10B, C</figureCitation>
|
||
) 1.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as pereopod 5; coxa bilobed, anterior lobe small, posterior lobe weakly dilated posterioventrally; basis subovoid, 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, anterior margin convex, posterior margin slightly dilated proximally; merus half as long as basis, 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long; carpus as long and 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as merus; propodus 1.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as carpus, with one pair of distal locking setae unequal in length (longer seta 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as dactylus), with one group of five setae (longest seta half as long as propodus) at anteriodistal corner; dactylus 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus, armed with one accessory cusp on outer margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 7 (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10D, E</figureCitation>
|
||
) 1.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as pereopod 6; coxa unilobed, dilated posterioventrally; basis subovoid, 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as that of pereopod 6, 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, anterior margin rather convex, posterior margin with one blunt extension proximally; merus subrectangular, a little extended posteriodistally, 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis; carpus rectangular, 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as merus, 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide; propodus 1.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as carpus, with one pair of distal locking setae subequal (smaller than those of pereopods 5 and 6); dactylus 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as propodus, armed with one accessory cusp on outer margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Epimeron 1 slightly extended anterioventrally. Epimera 2 and 3 with each posterioventral corner produced backwards, but not acute (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Uropod 1 (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10G</figureCitation>
|
||
) peduncle without distoventral spine, but bearing small acute extension, with six dorsolateral and three dorsomedial robust setae; inner ramus 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as peduncle, with one subapical seta only; outer ramus 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as inner ramus, with three dorsolateral setae and one subapical seta.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Uropod 2 (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10H</figureCitation>
|
||
) 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as uropod 1; peduncle 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as that of uropod 1, with small acute extension distoventrally; inner ramus 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as peduncle, with two dorsomedial setae and one subapical seta; outer ramus 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as inner ramus, with two dorsolateral setae and one subapical seta.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Uropod 3 (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10I</figureCitation>
|
||
) 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as uropod 2; peduncle 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as that of uropod 2; outer ramus biarticular, 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as peduncle, last article vestigial, with two elongate setae subapically; inner ramus scale-like, 0.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as outer ramus.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Telson (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photis bronca sp. nov., holotype, NIBRIV 0000848472, male, 5.8 mm, Daryeo-do Island, South Korea. A pereopod 5 B, pereopod 6 C dactylus of pereopod 6 D coxa of pereopod 7 E pereopod 7 F uropod 1 G uropod 2 H uropod 3 and telson. Scale bars: 0.1 mm (C, E, G-I), 0.2 mm (A, B, D, F)." figureDoi="10.3897/zookeys.886.38511.figure10" httpUri="https://binary.pensoft.net/fig/352722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure10">Fig. 10I</figureCitation>
|
||
) subtriangular in dorsal view, with a pair of simple setae, a pair of sensory setae, and one nodular robust seta on each side.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype female.</emphasis>
|
||
Gnathopod 1 (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Photis bronca sp. nov., paratype, NIBRIV 0000848473, female, 5.3 mm, Daryeo-do Island, South Korea. A gnathopod 1 B gnathopod 2 C coxa and oostegite of pereopod 3 D pereopod 4 E oostegite of pereopod 5. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure11">Fig. 11A</figureCitation>
|
||
) not sexually dimorphic.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, NIBRIV0000848473, female, 5.3 mm, Daryeo-do Island, South Korea.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
gnathopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
gnathopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
coxa and oostegite of pereopod 3
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
pereopod 4
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
oostegite of pereopod 5. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Gnathopod 2 (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Photis bronca sp. nov., paratype, NIBRIV 0000848473, female, 5.3 mm, Daryeo-do Island, South Korea. A gnathopod 1 B gnathopod 2 C coxa and oostegite of pereopod 3 D pereopod 4 E oostegite of pereopod 5. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure11">Fig. 11B</figureCitation>
|
||
) stouter than female gnathopod 1, slightly smaller than that of male; coxa subrectangular, 1.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide, evenly rounded anterioventrally, oostegite 1.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis, 2.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide; basis anterior margin lateral borders forming lobe distally, but smaller than that of male (not exceeding ischium), without stridulated ridges on surface; ischium with small anterior lobe only; merus rectangular, 0.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis; carpus stout, anterior margin convex, carpal lobe well developed; propodus stout, 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as basis, 0.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long, not widened distally, palmar margin cavity subquadrate, bearing one quadrate tooth medially, proximal and defining spines weaker than those of male, with one robust defining seta medially; dactylus half as long as propodus, inner margin minutely serrated, with three teeth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 3 (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Photis bronca sp. nov., paratype, NIBRIV 0000848473, female, 5.3 mm, Daryeo-do Island, South Korea. A gnathopod 1 B gnathopod 2 C coxa and oostegite of pereopod 3 D pereopod 4 E oostegite of pereopod 5. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure11">Fig. 11C</figureCitation>
|
||
) not sexually dimorphic except for the absence of both medial and lateral stridulated ridges on coxa; oostegite 1.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as that of gnathopod 2.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 4 (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Photis bronca sp. nov., paratype, NIBRIV 0000848473, female, 5.3 mm, Daryeo-do Island, South Korea. A gnathopod 1 B gnathopod 2 C coxa and oostegite of pereopod 3 D pereopod 4 E oostegite of pereopod 5. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure11">Fig. 11D</figureCitation>
|
||
) coxa without stridulated ridges at anterioventral corner, basis posterior margin more expanded distally, with twelve plumose setae along distal half.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Pereopod 5 oostegite (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Photis bronca sp. nov., paratype, NIBRIV 0000848473, female, 5.3 mm, Daryeo-do Island, South Korea. A gnathopod 1 B gnathopod 2 C coxa and oostegite of pereopod 3 D pereopod 4 E oostegite of pereopod 5. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.886.38511.figure11" httpUri="https://binary.pensoft.net/fig/352723" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure11">Fig. 11E</figureCitation>
|
||
) half as long as that of pereopod 4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has been misidentified as
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis longicaudata" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis longicaudata</emphasis>
|
||
</taxonomicName>
|
||
(Spence Bate & Westwood, 1862) in the Far East including Korea (
|
||
<bibRefCitation author="Kim, Y-H" journalOrPublisher="Korean Journal of Systematic Zoology Special issue" pageId="0" pageNumber="1" pagination="1 - 11" publicationUrl="http://www.e-ased.org/PublishedPaper/before_list.asp" refId="B23" refString="Kim, Y-H, Eun, Y, Lee, K-S, 2005. Amphipods (Gammaridea and Caprellidea) fauna of Jindo Island in Korea. Korean Journal of Systematic Zoology Special issue 5: 1 - 11" title="Amphipods (Gammaridea and Caprellidea) fauna of Jindo Island in Korea." url="http://www.e-ased.org/PublishedPaper/before_list.asp" volume="5" year="2005">Kim et al. 2005</bibRefCitation>
|
||
), Japan (
|
||
<bibRefCitation author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" pagination="131 - 170" publicationUrl="http://hdl.handle.net/2433/175398" refId="B36" refString="Nagata, K, 1965. Studies on marine gammaridean Amphipoda of the Seto Inland Sea. I. Publications of the Seto Marine Biological Laboratory 13: 131 - 170" title="Studies on marine gammaridean Amphipoda of the Seto Inland Sea. I." url="http://hdl.handle.net/2433/175398" volume="13" year="1965">Nagata 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.5134/176060" author="Hirayama, A" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" pagination="75 - 150" refId="B21" refString="Hirayama, A, 1984. Taxonomic studies on the shallow water gammaridean Amphipoda of the West Kyushu, Japan. II. Corophiidae. Publications of the Seto Marine Biological Laboratory 28: 75 - 150" title="Taxonomic studies on the shallow water gammaridean Amphipoda of the West Kyushu, Japan. II. Corophiidae." url="https://doi.org/10.5134/176060" volume="28" year="1984">Hirayama 1984</bibRefCitation>
|
||
), and China (
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" refId="B37" refString="Ren, XQ, 1992. Studies on the Gammaridea (Crustacea: Amphipoda) from Jiaozhou Bay (Yellow sea). In: Transactions of the Chinese Crustacean Society Vol. 3. Qingdao Ocean University Press, Qingdao, 214-317." title="Studies on the Gammaridea (Crustacea: Amphipoda) from Jiaozhou Bay (Yellow sea). In: Transactions of the Chinese Crustacean Society Vol. 3. Qingdao Ocean University Press, Qingdao, 214 - 317." year="1992">Ren 1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Studia Marina Sinica" pageId="0" pageNumber="1" pagination="136 - 145" refId="B38" refString="Ren, XQ, 2000. Studies on the genus Photis (Crustacea: Amphipoda) from the South China Sea. Studia Marina Sinica 42: 136 - 145" title="Studies on the genus Photis (Crustacea: Amphipoda) from the South China Sea." volume="42" year="2000">2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B39" refString="Ren, XQ, 2006. Fauna Sinica, Invertebrata Vol 41, Crustacea, Amphipoda, Gammaridea (I). Science Press, Beijing" title="Fauna Sinica, Invertebrata Vol 41, Crustacea, Amphipoda, Gammaridea (I)." year="2006">2006</bibRefCitation>
|
||
). The authors re-examined the syntype series of
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
deposited at the Natural History Museum, London (NHMUK 1911.20899-20906;
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Photis longicaudata (Spence Bate & Westwood, 1862), lectotype, NHMUK 1911.20899, male. 3.2 mm, Shetland Islands, UK. A gnathopod 1 B gnathopod 2 C coxa of pereopod 3 D coxa of pereopod 4. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.38511.figure12" httpUri="https://binary.pensoft.net/fig/352724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure12">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Photis longicaudata (Spence Bate & Westwood, 1862), paralectotype, NHMUK 1911.20890 - 20896, females. 5.3 mm (A, B) and 3.4 mm (C, D), Shetland Islands, UK A gnathopod 1 B gnathopod 2 C gnathopod 1 D gnathopod 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.38511.figure13" httpUri="https://binary.pensoft.net/fig/352725" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure13">13</figureCitation>
|
||
) and several specimens, which were collected from the North Sea, at the Museum
|
||
<normalizedToken originalValue="für">fuer</normalizedToken>
|
||
Naturkunde Berlin (ZMB 11656) to confirm the validity of
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
in Korean waters. As a result, the material formerly identified as
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
from Japan and China, as well as the Korean material, is herein described as a new species based on the following differences when compared with the newly designated
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype (NHMUK 1911.20899;
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Photis longicaudata (Spence Bate & Westwood, 1862), lectotype, NHMUK 1911.20899, male. 3.2 mm, Shetland Islands, UK. A gnathopod 1 B gnathopod 2 C coxa of pereopod 3 D coxa of pereopod 4. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.38511.figure12" httpUri="https://binary.pensoft.net/fig/352724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure12">Fig. 12</figureCitation>
|
||
): gnathopod 1 palmar margin has a robust defining seta medially in both sexes (absent in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype); gnathopod 2 palmar margin has a quadrate tooth medially in both sexes (obviously absent in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype); the sac-like lobe of male gnathopod 2 basis is much larger than that of
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
(slightly exceeding the ischium in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype); the anterior margin of male gnathopod 2 propodus is a little rounded (slightly shorter and more swollen in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype); the palmar margin of male gnathopod 2 propodus has more produced spines (slightly produced in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype); male pereopod 3 coxa has a different pattern of stridulated ridges; the male pereopod 4 coxa has stridulated ridges on the anterior half of the ventral margin (absent in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype); and female gnathopod 2 palmar margin is concave subrectangularly (more oblique and recessed in the
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
lectotype) (
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.9917" author="Bate, CS" journalOrPublisher="John van Voorst, London" pageId="0" pageNumber="1" refId="B7" refString="Bate, CS, Westwood, JL, 1863. A history of the British sessile-eyed Crustacea, Vol 1. John van Voorst, London" title="A history of the British sessile-eyed Crustacea, Vol 1." url="https://doi.org/10.5962/bhl.title.9917" year="1863">Bate and Westwood 1863</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.1164" author="Sars, GO" journalOrPublisher="ALB Cammermeyers Forlag, Copenhagen" pageId="0" pageNumber="1" refId="B42" refString="Sars, GO, 1890-1895. An account of the Crustacea of Norway with short descriptions and figures of all the species, Vol I. Amphipoda. ALB Cammermeyers Forlag, Copenhagen" title="An account of the Crustacea of Norway with short descriptions and figures of all the species, Vol I. Amphipoda." url="https://doi.org/10.5962/bhl.title.1164" year="1890 - 1895">Sars 1895</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" pagination="131 - 170" publicationUrl="http://hdl.handle.net/2433/175398" refId="B36" refString="Nagata, K, 1965. Studies on marine gammaridean Amphipoda of the Seto Inland Sea. I. Publications of the Seto Marine Biological Laboratory 13: 131 - 170" title="Studies on marine gammaridean Amphipoda of the Seto Inland Sea. I." url="http://hdl.handle.net/2433/175398" volume="13" year="1965">Nagata 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.5134/176060" author="Hirayama, A" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" pagination="75 - 150" refId="B21" refString="Hirayama, A, 1984. Taxonomic studies on the shallow water gammaridean Amphipoda of the West Kyushu, Japan. II. Corophiidae. Publications of the Seto Marine Biological Laboratory 28: 75 - 150" title="Taxonomic studies on the shallow water gammaridean Amphipoda of the West Kyushu, Japan. II. Corophiidae." url="https://doi.org/10.5134/176060" volume="28" year="1984">Hirayama 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="0" pageNumber="1" refId="B37" refString="Ren, XQ, 1992. Studies on the Gammaridea (Crustacea: Amphipoda) from Jiaozhou Bay (Yellow sea). In: Transactions of the Chinese Crustacean Society Vol. 3. Qingdao Ocean University Press, Qingdao, 214-317." title="Studies on the Gammaridea (Crustacea: Amphipoda) from Jiaozhou Bay (Yellow sea). In: Transactions of the Chinese Crustacean Society Vol. 3. Qingdao Ocean University Press, Qingdao, 214 - 317." year="1992">Ren 1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Studia Marina Sinica" pageId="0" pageNumber="1" pagination="136 - 145" refId="B38" refString="Ren, XQ, 2000. Studies on the genus Photis (Crustacea: Amphipoda) from the South China Sea. Studia Marina Sinica 42: 136 - 145" title="Studies on the genus Photis (Crustacea: Amphipoda) from the South China Sea." volume="42" year="2000">2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B39" refString="Ren, XQ, 2006. Fauna Sinica, Invertebrata Vol 41, Crustacea, Amphipoda, Gammaridea (I). Science Press, Beijing" title="Fauna Sinica, Invertebrata Vol 41, Crustacea, Amphipoda, Gammaridea (I)." year="2006">2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Kim, Y-H" journalOrPublisher="Korean Journal of Systematic Zoology Special issue" pageId="0" pageNumber="1" pagination="1 - 11" publicationUrl="http://www.e-ased.org/PublishedPaper/before_list.asp" refId="B23" refString="Kim, Y-H, Eun, Y, Lee, K-S, 2005. Amphipods (Gammaridea and Caprellidea) fauna of Jindo Island in Korea. Korean Journal of Systematic Zoology Special issue 5: 1 - 11" title="Amphipods (Gammaridea and Caprellidea) fauna of Jindo Island in Korea." url="http://www.e-ased.org/PublishedPaper/before_list.asp" volume="5" year="2005">Kim et al. 2005</bibRefCitation>
|
||
; see
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Photis longicaudata (Spence Bate & Westwood, 1862), lectotype, NHMUK 1911.20899, male. 3.2 mm, Shetland Islands, UK. A gnathopod 1 B gnathopod 2 C coxa of pereopod 3 D coxa of pereopod 4. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.38511.figure12" httpUri="https://binary.pensoft.net/fig/352724" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure12">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Photis longicaudata (Spence Bate & Westwood, 1862), paralectotype, NHMUK 1911.20890 - 20896, females. 5.3 mm (A, B) and 3.4 mm (C, D), Shetland Islands, UK A gnathopod 1 B gnathopod 2 C gnathopod 1 D gnathopod 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.38511.figure13" httpUri="https://binary.pensoft.net/fig/352725" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.886.38511.figure13">13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.886.38511.figure12" httpUri="https://binary.pensoft.net/fig/352724" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis longicaudata" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis longicaudata</emphasis>
|
||
</taxonomicName>
|
||
(Spence Bate & Westwood, 1862), lectotype, NHMUK 1911.20899, male. 3.2 mm, Shetland Islands, UK.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
gnathopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
gnathopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
coxa of pereopod 3
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
coxa of pereopod 4. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.886.38511.figure13" httpUri="https://binary.pensoft.net/fig/352725" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis longicaudata" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis longicaudata</emphasis>
|
||
</taxonomicName>
|
||
(Spence Bate & Westwood, 1862), paralectotype, NHMUK 1911.20890-20896, females. 5.3 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
) and 3.4 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||
), Shetland Islands, UK
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
gnathopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
gnathopod 2
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
gnathopod 1
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
gnathopod 2. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
It is well known that both
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis aina" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis aina</emphasis>
|
||
</taxonomicName>
|
||
JL Barnard, 1970 and
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis kapapa" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis kapapa</emphasis>
|
||
</taxonomicName>
|
||
JL Barnard, 1970 from the Hawaiian Islands have affinities with
|
||
<taxonomicName lsidName="P. longicaudata" pageId="0" pageNumber="1" rank="species" species="longicaudata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. longicaudata</emphasis>
|
||
</taxonomicName>
|
||
in that gnathopod 2 has very well-developed (sac-like) anterior lobe of the basis (JL
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.34" author="Barnard, JL" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="1" pagination="1 - 286" refId="B4" refString="Barnard, JL, 1970. Sublittoral Gammaridea (Amphipoda) of the Hawaiian Islands. Smithsonian Contributions to Zoology 34: 1 - 286" title="Sublittoral Gammaridea (Amphipoda) of the Hawaiian Islands." url="https://doi.org/10.5479/si.00810282.34" volume="34" year="1970">Barnard 1970</bibRefCitation>
|
||
). Among them,
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName lsidName="P. aina" pageId="0" pageNumber="1" rank="species" species="aina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aina</emphasis>
|
||
</taxonomicName>
|
||
, regarding the eyes being in the middle of the cephalic lobe (occupying the end of the cephalic lobe in
|
||
<taxonomicName lsidName="P. aina" pageId="0" pageNumber="1" rank="species" species="aina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aina</emphasis>
|
||
</taxonomicName>
|
||
), lacking the defining seta on the palmar margin of male gnathopod 1 (with one robust seta medially in
|
||
<taxonomicName lsidName="P. aina" pageId="0" pageNumber="1" rank="species" species="aina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aina</emphasis>
|
||
</taxonomicName>
|
||
), lacking posterioventral notches of epimera 2 and 3, and a shorter inner ramus of uropod 2 (about half length in
|
||
<taxonomicName lsidName="P. aina" pageId="0" pageNumber="1" rank="species" species="aina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. aina</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.34" author="Barnard, JL" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="1" pagination="1 - 286" refId="B4" refString="Barnard, JL, 1970. Sublittoral Gammaridea (Amphipoda) of the Hawaiian Islands. Smithsonian Contributions to Zoology 34: 1 - 286" title="Sublittoral Gammaridea (Amphipoda) of the Hawaiian Islands." url="https://doi.org/10.5479/si.00810282.34" volume="34" year="1970">Barnard 1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.5.1985.99" author="Myers, AA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="1" pagination="1 - 143" refId="B29" refString="Myers, AA, 1985. Shallow-water, coral reef and mangrove Amphipoda (Gammaridea) of Fiji. Records of the Australian Museum Supplement 5: 1 - 143" title="Shallow-water, coral reef and mangrove Amphipoda (Gammaridea) of Fiji." url="https://doi.org/10.3853/j.0812-7387.5.1985.99" volume="5" year="1985">Myers 1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2260.1.43" author="Myers, AA" editor="Lowry, JK" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="771 - 799" refId="B31" refString="Myers, AA, 2009. Photidae. In: Lowry, JK, Myers, AA, Eds., Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef, Australia. . Zootaxa 2260: 771 - 799" title="Photidae." url="https://doi.org/10.11646/zootaxa.2260.1.43" volume="2260" volumeTitle="Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef, Australia." year="2009">2009</bibRefCitation>
|
||
).
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
by the following differences: the eyes occupying at the middle of the cephalic lobe (at the end of the cephalic lobe in
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
), lacking stridulated ridges in both posterior margins of the coxae of gnathopods 1 and 2, 0.7 times as wide as long male gnathopod 2 (0.8 times in
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
), the female palmar margin of gnathopod 2 with a small obtuse spine near the dactylus base (without spine in
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
), the presence of one accessory cusp on pereopod 5 dactylus (two accessory cusps in
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
), subacutely produced posterior margin of epimeron 3 (rounded in
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
), and uropod 3 outer ramus with the length of 0.6 times as long as the peduncle (0.9 times in
|
||
<taxonomicName lsidName="P. kapapa" pageId="0" pageNumber="1" rank="species" species="kapapa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. kapapa</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.34" author="Barnard, JL" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="1" pagination="1 - 286" refId="B4" refString="Barnard, JL, 1970. Sublittoral Gammaridea (Amphipoda) of the Hawaiian Islands. Smithsonian Contributions to Zoology 34: 1 - 286" title="Sublittoral Gammaridea (Amphipoda) of the Hawaiian Islands." url="https://doi.org/10.5479/si.00810282.34" volume="34" year="1970">Barnard 1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ren, XQ" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B39" refString="Ren, XQ, 2006. Fauna Sinica, Invertebrata Vol 41, Crustacea, Amphipoda, Gammaridea (I). Science Press, Beijing" title="Fauna Sinica, Invertebrata Vol 41, Crustacea, Amphipoda, Gammaridea (I)." year="2006">Ren 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis bronca" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. resembles
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis davei" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="davei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis davei</emphasis>
|
||
</taxonomicName>
|
||
Myers, 2009 in the shapes of elongate propodus of pereopods 3 and 4 as well as the outlines of gnathopods in both sexes, but the former differs from the latter in the presence of a well-developed sac-like lateral lobe on the basis of male gnathopod 2 (present but very weak in
|
||
<taxonomicName lsidName="P. davei" pageId="0" pageNumber="1" rank="species" species="davei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. davei</emphasis>
|
||
</taxonomicName>
|
||
) and not-elongate rami of uropods 1 and 2 (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2260.1.43" author="Myers, AA" editor="Lowry, JK" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="771 - 799" refId="B31" refString="Myers, AA, 2009. Photidae. In: Lowry, JK, Myers, AA, Eds., Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef, Australia. . Zootaxa 2260: 771 - 799" title="Photidae." url="https://doi.org/10.11646/zootaxa.2260.1.43" volume="2260" volumeTitle="Benthic Amphipoda (Crustacea: Peracarida) of the Great Barrier Reef, Australia." year="2009">Myers 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis fischmanni" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fischmanni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis fischmanni</emphasis>
|
||
</taxonomicName>
|
||
Gurjanova, 1951 also has a similar gnathopod 2 shape in both sexes, but
|
||
<taxonomicName lsidName="P. bronca" pageId="0" pageNumber="1" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is clearly different from this species by elongate antenna 1 and 2 (0.4 times as long as the body in
|
||
<taxonomicName lsidName="P. bronca" pageId="0" pageNumber="1" rank="species" species="bronca">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. bronca</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., but less than 0.3 times in
|
||
<taxonomicName lsidName="P. fischmanni" pageId="0" pageNumber="1" rank="species" species="fischmanni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. fischmanni</emphasis>
|
||
</taxonomicName>
|
||
), less expanded carpus and propodus of gnathopod 1, and less setose appendages than
|
||
<taxonomicName lsidName="P. fischmanni" pageId="0" pageNumber="1" rank="species" species="fischmanni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. fischmanni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Gurjanova, E" journalOrPublisher="Opredeliteli po faune SSSR, Akademiya Nauk SSSR, Opredeliteli po Faune SSSR" pageId="0" pageNumber="1" pagination="1 - 1029" refId="B19" refString="Gurjanova, E, 1951. Bokoplavy morei SSSR i sopredelnykh vod (Amphipoda,Gammaridea). Opredeliteli po faune SSSR, Akademiya Nauk SSSR, Opredeliteli po Faune SSSR 41: 1 - 1029" title="Bokoplavy morei SSSR i sopredelnykh vod (Amphipoda, Gammaridea)." volume="41" year="1951">Gurjanova 1951</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Gurjanova, E" journalOrPublisher="Zoologicheskogo Instituta Akademii Nauk SSSR, Trudy" pageId="0" pageNumber="1" pagination="166 - 218" refId="B20" refString="Gurjanova, E, 1955. Novye vidy bokoplavov (Amphipoda, Gammaridea) iz servernoi chasti Tixogo Okeana. Zoologicheskogo Instituta Akademii Nauk SSSR, Trudy 18: 166 - 218" title="Novye vidy bokoplavov (Amphipoda, Gammaridea) iz servernoi chasti Tixogo Okeana." volume="18" year="1955">1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Conlan, KE" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="1" refId="B16" refString="Conlan, KE, 1983. The amphipod superfamily Corophioidea in the Northeastern Pacific region. 3. Family Isaeidae: Systematics and Distributional Ecology. Publications in Natural Sciences 4: 1-75. (National Museum of Natural Science: Ottawa, Canada.)" title="The amphipod superfamily Corophioidea in the Northeastern Pacific region. 3. Family Isaeidae: Systematics and Distributional Ecology. Publications in Natural Sciences 4: 1 - 75. (National Museum of Natural Science: Ottawa, Canada.)" year="1983">Conlan 1983</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This new species differs from
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis paeowai" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paeowai">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis paeowai</emphasis>
|
||
</taxonomicName>
|
||
Myers, 1995 and
|
||
<taxonomicName class="Malacostraca" family="Photidae" genus="Photis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Photis pirloti" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pirloti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Photis pirloti</emphasis>
|
||
</taxonomicName>
|
||
Myers, 1985 by bearing a well-developed sac-like lateral lobe on the basis of male gnathopod 2 (present but rather weak in
|
||
<taxonomicName lsidName="P. paeowai" pageId="0" pageNumber="1" rank="species" species="paeowai">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. paeowai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="P. pirloti" pageId="0" pageNumber="1" rank="species" species="pirloti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. pirloti</emphasis>
|
||
</taxonomicName>
|
||
) and a quadrate medial tooth on the palmar margin of gnathopod 2 in both sexes (absent in
|
||
<taxonomicName lsidName="P. paeowai" pageId="0" pageNumber="1" rank="species" species="paeowai">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. paeowai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="P. pirloti" pageId="0" pageNumber="1" rank="species" species="pirloti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. pirloti</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.5.1985.99" author="Myers, AA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="1" pagination="1 - 143" refId="B29" refString="Myers, AA, 1985. Shallow-water, coral reef and mangrove Amphipoda (Gammaridea) of Fiji. Records of the Australian Museum Supplement 5: 1 - 143" title="Shallow-water, coral reef and mangrove Amphipoda (Gammaridea) of Fiji." url="https://doi.org/10.3853/j.0812-7387.5.1985.99" volume="5" year="1985">Myers 1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0812-7387.22.1995.121" author="Myers, AA" journalOrPublisher="Records of the Australian Museum Supplement" pageId="0" pageNumber="1" pagination="25 - 95" refId="B30" refString="Myers, AA, 1995. The Amphipoda (Crustacea) of Madang Lagoon: Aoridae, Isaeidae, Neomegamphopidae. Records of the Australian Museum Supplement 22: 25 - 95" title="The Amphipoda (Crustacea) of Madang Lagoon: Aoridae, Isaeidae, Neomegamphopidae." url="https://doi.org/10.3853/j.0812-7387.22.1995.121" volume="22" year="1995">1995</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |