323 lines
38 KiB
XML
323 lines
38 KiB
XML
<document id="777BAD745B297BF8D52C652865D5D08E" ID-CLB-Dataset="22065" ID-DOI="10.5852/ejt.2017.340" ID-GBIF-Dataset="0be43b36-2678-45a5-acfc-bec202ecb14a" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832552" ID-ZooBank="2C6DC937-E543-4B8D-BF24-CED437F97676" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589831085025" checkinUser="carolina" docAuthor="Ricchiardi, Enrico" docDate="2017" docId="03E9E50844567C61FDF7F9D3FD7A4AFC" docLanguage="en" docName="ejt-2017-340.pdf.imf" docOrigin="European Journal of Taxonomy 340 (340)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hybovalgus calvus Ricchiardi 2017, sp. nov." docType="treatment" docUuid="2AE1E317-E510-4210-B843-973C1985BF3E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="16" masterDocId="FFD09D70445A7C6EFF80FFC2FF874B39" masterDocTitle="Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="13" updateTime="1698840126634" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="441971F693F9A54E0CDF6747ED2F4B3E">Revision of Chinese mainland Hybovalgus Kolbe, 1904, with description of a new species, and Excisivalgus Endrödi, 1952 reduced to synonymy with Hybovalgus (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="85B1BDAE447C530EB7559BC1634DC722">Ricchiardi, Enrico</mods:namePart>
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<mods:affiliation id="9508A0A9470974E37509840B4C2D9665">Corso A. Tassoni 79 / 4, 10143 Torino, Italy. Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Box 92, Beichen West road, Chaoyang District, Beijing, 100101, China; University of the Chinese Academy of Sciences, Yuquan road, Shijingshan, Beijing, 100039, China.</mods:affiliation>
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<mods:nameIdentifier id="B5E69B24CB9F6576ADAC038A492C565E" type="email">alericor@fastwebnet.it&545407454@qq.com</mods:nameIdentifier>
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<mods:title id="50961344763095C8F433DD807039D623">European Journal of Taxonomy</mods:title>
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<mods:part id="F9AE2280BA0A9C5A745155B41D23E44D">
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<mods:date id="058D5A7FA4F1658C0EA55BC199908CB0">2017</mods:date>
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<treatment id="03E9E50844567C61FDF7F9D3FD7A4AFC" ID-DOI="http://doi.org/10.5281/zenodo.3850415" ID-GBIF-Taxon="164167212" ID-Zenodo-Dep="3850415" LSID="urn:lsid:zoobank.org:act:2AE1E317-E510-4210-B843-973C1985BF3E" httpUri="http://treatment.plazi.org/id/03E9E50844567C61FDF7F9D3FD7A4AFC" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
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<subSubSection id="C35A079544567C62FDF7F9D3FC3A4D15" box="[631,957,1553,1580]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BFF541E44567C62FDF7F9D3FC3A4D15" blockId="12.[375,1213,1553,1653]" box="[631,957,1553,1580]" pageId="12" pageNumber="13">
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<heading id="D0B7E37244567C62FDF7F9D3FC3A4D15" box="[631,957,1553,1580]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
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<taxonomicName id="4C402F9D44567C62FDF7F9D3FCDC4D15" ID-CoL="6MHRG" authority="Ricchiardi, 2017" authorityName="Ricchiardi" authorityYear="2017" box="[631,859,1553,1580]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
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<emphasis id="B934880C44567C62FDF7F9D3FCDC4D15" bold="true" box="[631,859,1553,1580]" italics="true" pageId="12" pageNumber="13">Hybovalgus calvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A207357744567C62FCE2F9D0FC3A4D15" box="[866,957,1554,1580]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A079544567C62FEF7F9F5FCBB4D4C" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BFF541E44567C62FEF7F9F5FB3A4D6B" blockId="12.[375,1213,1553,1653]" box="[375,1213,1591,1618]" pageId="12" pageNumber="13">
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<uri id="FFD1581C44567C62FEF7F9F5FB3A4D6B" box="[375,1213,1591,1618]" pageId="12" pageNumber="13">
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urn:lsid:zoobank.org:act:
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<uuid id="FFE66ECB44567C62FD19F9F5FB3A4D6B" box="[665,1213,1591,1618]" pageId="12" pageNumber="13">2AE1E317-E510-4210-B843-973C1985BF3E</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BFF541E44567C62FD77F999FCBB4D4C" blockId="12.[375,1213,1553,1653]" box="[759,828,1627,1653]" pageId="12" pageNumber="13">
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<figureCitation id="137B489B44567C62FD77F999FCBB4D4C" box="[759,828,1627,1653]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[216,1363,985,1846]" captionTargetId="figure@13.[202,1385,952,1854]" captionTargetPageId="13" captionText="Fig. 6. Hybovalgus calvus sp. nov. A–C, H–I. HT ♂ (MNHUB). A. Habitus, dorsal view. B. Protibia. C. Pygidium. H–I. Aedeagus (grey: parameres; white: phallobase). H. Dorsal view. I. Lateral view. D–G. PT ♀ (ERC). D. Habitus, dorsal view. E. Protibia. F. Pygidium, frontal view. G. Pygidium, lateral view. J–K. PT ♂ (ERC). J. Habitus, dorsal view. K. Propygidium and pygidium." figureDoi="http://doi.org/10.5281/zenodo.3832564" httpUri="https://zenodo.org/record/3832564/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A079544567C62FF3DF95CFD8B4CD2" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="8BFF541E44567C62FF3DF95CFEB14D81" blockId="12.[189,310,1694,1720]" box="[189,310,1694,1720]" pageId="12" pageNumber="13">
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<heading id="D0B7E37244567C62FF3DF95CFEB14D81" bold="true" box="[189,310,1694,1720]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
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<emphasis id="B934880C44567C62FF3DF95CFEB14D81" bold="true" box="[189,310,1694,1720]" pageId="12" pageNumber="13">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFF541E44567C62FF3DF90DFD8B4CD2" blockId="12.[189,1399,1743,2027]" pageId="12" pageNumber="13">
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<taxonomicName id="4C402F9D44567C62FF3DF90DFE194DD0" authority="Ricchiardi, 2017" authorityName="Ricchiardi" authorityYear="2017" box="[189,414,1743,1769]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
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<emphasis id="B934880C44567C62FF3DF90DFE194DD0" box="[189,414,1743,1769]" italics="true" pageId="12" pageNumber="13">Hybovalgus calvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A207357744567C62FE28F90DFD824DD0" box="[424,517,1743,1769]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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can easily be separated from all other Chinese
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<taxonomicName id="4C402F9D44567C62FBC7F90DFB544DD0" authorityName="Kolbe" authorityYear="1904" box="[1095,1235,1743,1769]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B934880C44567C62FBC7F90DFB544DD0" box="[1095,1235,1743,1769]" italics="true" pageId="12" pageNumber="13">Hybovalgus</emphasis>
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</taxonomicName>
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(except from
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<taxonomicName id="4C402F9D44567C62FF3DF937FEC34C37" authorityName="Ricchiardi" authorityYear="2017" baseAuthorityName="Endrodi" baseAuthorityYear="1952" box="[189,324,1780,1806]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jordansi">
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<emphasis id="B934880C44567C62FF3DF937FEC34C37" box="[189,324,1780,1806]" italics="true" pageId="12" pageNumber="13">H. jordansi</emphasis>
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</taxonomicName>
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) by the V-shaped protrusion at the posterior margin of the pronotum (
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<figureCitation id="137B489B44567C62FB10F936FAB64C37" box="[1168,1329,1780,1806]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[216,1363,985,1846]" captionTargetId="figure@13.[202,1385,952,1854]" captionTargetPageId="13" captionText="Fig. 6. Hybovalgus calvus sp. nov. A–C, H–I. HT ♂ (MNHUB). A. Habitus, dorsal view. B. Protibia. C. Pygidium. H–I. Aedeagus (grey: parameres; white: phallobase). H. Dorsal view. I. Lateral view. D–G. PT ♀ (ERC). D. Habitus, dorsal view. E. Protibia. F. Pygidium, frontal view. G. Pygidium, lateral view. J–K. PT ♂ (ERC). J. Habitus, dorsal view. K. Propygidium and pygidium." figureDoi="http://doi.org/10.5281/zenodo.3832564" httpUri="https://zenodo.org/record/3832564/files/figure.png" pageId="12" pageNumber="13">Fig. 6A, D, J</figureCitation>
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). The female pygidium apex lacks a protrusion. The parameres are different from those of
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<taxonomicName id="4C402F9D44567C62FB36F8DBFAD14C0B" authorityName="Kolbe" authorityYear="1904" box="[1206,1366,1816,1842]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bioculatus">
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<emphasis id="B934880C44567C62FB36F8DBFAD14C0B" box="[1206,1366,1816,1842]" italics="true" pageId="12" pageNumber="13">H. bioculatus</emphasis>
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</taxonomicName>
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in the C-shaped apex (
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<figureCitation id="137B489B44567C62FE24F8FFFD854C61" box="[420,514,1853,1880]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[216,1363,985,1846]" captionTargetId="figure@13.[202,1385,952,1854]" captionTargetPageId="13" captionText="Fig. 6. Hybovalgus calvus sp. nov. A–C, H–I. HT ♂ (MNHUB). A. Habitus, dorsal view. B. Protibia. C. Pygidium. H–I. Aedeagus (grey: parameres; white: phallobase). H. Dorsal view. I. Lateral view. D–G. PT ♀ (ERC). D. Habitus, dorsal view. E. Protibia. F. Pygidium, frontal view. G. Pygidium, lateral view. J–K. PT ♂ (ERC). J. Habitus, dorsal view. K. Propygidium and pygidium." figureDoi="http://doi.org/10.5281/zenodo.3832564" httpUri="https://zenodo.org/record/3832564/files/figure.png" pageId="12" pageNumber="13">Fig. 6H</figureCitation>
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). Females of
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<taxonomicName id="4C402F9D44567C62FD1EF8FFFCFB4C6E" authority="Ricchiardi, 2017" authorityName="Ricchiardi" authorityYear="2017" box="[670,892,1853,1879]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
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<emphasis id="B934880C44567C62FD1EF8FFFCFB4C6E" box="[670,892,1853,1879]" italics="true" pageId="12" pageNumber="13">Hybovalgus calvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A207357744567C62FC03F8FCFC5A4C61" box="[899,989,1854,1880]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from those of
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<taxonomicName id="4C402F9D44567C62FF3DF8A1FEC64C45" authorityName="Ricchiardi" authorityYear="2017" baseAuthorityName="Endrodi" baseAuthorityYear="1952" box="[189,321,1890,1916]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jordansi">
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<emphasis id="B934880C44567C62FF3DF8A1FEC64C45" box="[189,321,1890,1916]" italics="true" pageId="12" pageNumber="13">H. jordansi</emphasis>
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</taxonomicName>
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by the prominent testaceous scale tufts at the posterior margin of the pronotum; by the fringe of small, testaceous scales at the pygidial apex, instead of prominent, double-pointed testaceous scale tufts. The apex of the parameres of both species (
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<figureCitation id="137B489B44567C62FD7AF86EFCDC4CFF" box="[762,859,1964,1990]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[216,1363,985,1846]" captionTargetId="figure@13.[202,1385,952,1854]" captionTargetPageId="13" captionText="Fig. 6. Hybovalgus calvus sp. nov. A–C, H–I. HT ♂ (MNHUB). A. Habitus, dorsal view. B. Protibia. C. Pygidium. H–I. Aedeagus (grey: parameres; white: phallobase). H. Dorsal view. I. Lateral view. D–G. PT ♀ (ERC). D. Habitus, dorsal view. E. Protibia. F. Pygidium, frontal view. G. Pygidium, lateral view. J–K. PT ♂ (ERC). J. Habitus, dorsal view. K. Propygidium and pygidium." figureDoi="http://doi.org/10.5281/zenodo.3832564" httpUri="https://zenodo.org/record/3832564/files/figure.png" pageId="12" pageNumber="13">Figs 6H</figureCitation>
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,
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<figureCitation id="137B489B44567C62FCE7F86EFC064CFF" box="[871,897,1964,1990]" captionStart="Fig" captionStartId="20.[189,232,1752,1778]" captionTargetBox="[236,1372,370,1677]" captionTargetId="figure@20.[212,1376,353,1697]" captionTargetPageId="20" captionText="Fig. 8. A–F. Hybovalgus sexdentatus Arrow, 1944. A–B. ♂ (ERC). A. Habitus, dorsal view. B. Propygidium and pygidium. C–D. ♀ (ERC). C. Habitus, dorsal view. D. Propygidium and pygidium. E–F. PLT, ♂ (NHM). E. Aedeagus, dorsal view. F. Aedeagus, lateral view. G–J. Hybovalgus jordansi (Endrödi, 1952). G–H. ♂ (ERC). G. Habitus, dorsal view. H. Propygidiun and pygidium. I–J. PT, ♂ (HNHM). I. Aedeagus, dorsal view. J. Aedeagus, lateral view. K–N. Hybovalgus fulvosquamosus Moser, 1904. K–L. ♂ (ERC). K. Habitus, dorsal view. L. Propygidiun and pygidium. M–N. LT, ♂ (MNHUB). M. Aedeagus, dorsal view. N. Aedeagus, lateral view. Drawings: grey parts: parameres; white parts: phallobase." figureDoi="http://doi.org/10.5281/zenodo.3832568" httpUri="https://zenodo.org/record/3832568/files/figure.png" pageId="12" pageNumber="13">8I</figureCitation>
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) are inwardly C-shaped, but in
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<taxonomicName id="4C402F9D44567C62FB72F86FFAF04CFF" authorityName="Ricchiardi" authorityYear="2017" baseAuthorityName="Endrodi" baseAuthorityYear="1952" box="[1266,1399,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jordansi">
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<emphasis id="B934880C44567C62FB72F86FFAF04CFF" box="[1266,1399,1964,1990]" italics="true" pageId="12" pageNumber="13">H. jordansi</emphasis>
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</taxonomicName>
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they are thinner and tapered.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A079544577C63FF3DFEC9FE434A4F" pageId="13" pageNumber="14" type="etymology">
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<paragraph id="8BFF541E44577C63FF3DFEC9FEC34A1C" blockId="13.[189,324,267,293]" box="[189,324,267,293]" pageId="13" pageNumber="14">
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<heading id="D0B7E37244577C63FF3DFEC9FEC34A1C" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
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<emphasis id="B934880C44577C63FF3DFEC9FEC34A1C" bold="true" box="[189,324,267,293]" pageId="13" pageNumber="14">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFF541E44577C63FF3DFEFBFE434A4F" blockId="13.[189,1399,313,375]" pageId="13" pageNumber="14">
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The name comes from the Latin word “
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<taxonomicName id="4C402F9D44577C63FD1FFEFBFD6D4A6A" authorityName="Ricchiardi" authorityYear="2017" box="[671,746,313,339]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="calvus">
|
||
<emphasis id="B934880C44577C63FD1FFEFBFD6D4A6A" box="[671,746,313,339]" italics="true" pageId="13" pageNumber="14">calvus</emphasis>
|
||
</taxonomicName>
|
||
”, meaning glabrous, as the integument of this species appears at first glance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A079544577C63FF3DFE5CFCBB4989" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BFF541E44577C63FF3DFE5CFEEB4A81" blockId="13.[189,364,414,440]" box="[189,364,414,440]" pageId="13" pageNumber="14">
|
||
<heading id="D0B7E37244577C63FF3DFE5CFEEB4A81" bold="true" box="[189,364,414,440]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B934880C44577C63FF3DFE5CFEEB4A81" bold="true" box="[189,364,414,440]" pageId="13" pageNumber="14">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B285E4344577C63FF6AFE0FFAA84933" ID-GBIF-Occurrence="2620519302" collectionCode="MNHUB" collectorName="H. Fruhstorfer" country="Vietnam" location="Tonkin" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<paragraph id="8BFF541E44577C63FF6AFE0FFEDB4ADE" blockId="13.[189,1327,461,522]" box="[234,348,461,487]" pageId="13" pageNumber="14">
|
||
<emphasis id="B934880C44577C63FF6AFE0FFEDB4ADE" bold="true" box="[234,348,461,487]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54FBEABC44577C63FF6AFE0FFEDB4ADE" box="[234,348,461,487]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44577C63FF3DFE32FAA84933" blockId="13.[189,1327,461,522]" box="[189,1327,496,522]" pageId="13" pageNumber="14">
|
||
<collectingCountry id="F357148E44577C63FF3DFE32FECE4933" box="[189,329,496,522]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
|
||
:
|
||
<specimenCount id="9D469F9744577C63FED4FE32FEEB4933" box="[340,364,496,522]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
|
||
,
|
||
<location id="8E9F02C544577C63FEF8FE32FE494933" LSID="urn:lsid:plazi:treatment:03E9E50844567C61FDF7F9D3FD7A4AFC:8E9F02C544577C63FEF8FE32FE494933" box="[376,462,496,522]" country="Vietnam" name="Tonkin" pageId="13" pageNumber="14">Tonkin</location>
|
||
,
|
||
<location id="8E9F02C544577C63FE59FE32FD1C4933" LSID="urn:lsid:plazi:treatment:03E9E50844567C61FDF7F9D3FD7A4AFC:8E9F02C544577C63FE59FE32FD1C4933" box="[473,667,496,522]" country="Vietnam" name="Montes Mauson" pageId="13" pageNumber="14">Montes Mauson</location>
|
||
, Apr.–May, 2000–3000’,
|
||
<collectorName id="26B531C844577C63FC45FE32FBEA4933" box="[965,1133,496,522]" pageId="13" pageNumber="14">H. Fruhstorfer</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED51CCDB44577C63FB2FFE32FAA44933" box="[1199,1315,496,522]" collectionName="MNHUB" pageId="13" pageNumber="14">MNHUB</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B285E4344577C63FF6AFDF3FC104956" ID-GBIF-Occurrence="3763235352" collectionCode="ERC, MNHUB" country="Vietnam" location="VIETNAM" pageId="13" pageNumber="14" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<paragraph id="8BFF541E44577C63FF6AFDF3FEE04972" blockId="13.[189,919,561,623]" box="[234,359,561,587]" pageId="13" pageNumber="14">
|
||
<emphasis id="B934880C44577C63FF6AFDF3FEE04972" bold="true" box="[234,359,561,587]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54FBEABC44577C63FF6AFDF3FEE04972" box="[234,359,561,587]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44577C63FF3DFD97FC104956" blockId="13.[189,919,561,623]" box="[189,919,596,623]" pageId="13" pageNumber="14">
|
||
<collectingCountry id="F357148E44577C63FF3DFD97FECE4956" box="[189,329,597,623]" name="Vietnam" pageId="13" pageNumber="14">VIETNAM</collectingCountry>
|
||
:
|
||
<specimenCount id="9D469F9744577C63FED4FD97FE074956" box="[340,384,597,623]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED51CCDB44577C63FE0FFD97FE494956" box="[399,462,597,623]" pageId="13" pageNumber="14">ERC</collectionCode>
|
||
),
|
||
<specimenCount id="9D469F9744577C63FE62FD97FD894956" box="[482,526,597,623]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED51CCDB44577C63FD9EFD97FD164956" box="[542,657,597,623]" collectionName="MNHUB" pageId="13" pageNumber="14">MNHUB</collectionCode>
|
||
), same data as for HT.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BFF541E44577C63FF3DFD54FCBB4989" blockId="13.[189,828,662,688]" box="[189,828,662,688]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="3B285E4344577C63FF3DFD54FCBB4989" ID-GBIF-Occurrence="3763235344" box="[189,828,662,688]" collectingDate="2009-06-01" collectionCode="ERC" country="China" location="Wuzhishan" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
|
||
<collectingCountry id="F357148E44577C63FF3DFD54FE984989" box="[189,287,662,688]" name="China" pageId="13" pageNumber="14">CHINA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D469F9744577C63FEA9FD54FED04989" box="[297,343,662,688]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="49849AFC44577C63FEE4FD54FE3B4989" box="[356,444,662,688]" country="China" name="Hainan" pageId="13" pageNumber="14">Hainan</collectingRegion>
|
||
,
|
||
<location id="8E9F02C544577C63FE47FD54FDCA4989" LSID="urn:lsid:plazi:treatment:03E9E50844567C61FDF7F9D3FD7A4AFC:8E9F02C544577C63FE47FD54FDCA4989" box="[455,589,662,688]" country="China" name="Wuzhishan" pageId="13" pageNumber="14" stateProvince="Hainan">Wuzhishan</location>
|
||
,
|
||
<date id="FFFE72DE44577C63FDD8FD54FD664989" box="[600,737,662,688]" pageId="13" pageNumber="14" value="2009-06-01">
|
||
<collectingDate id="EFBA8B3644577C63FDD8FD54FD664989" box="[600,737,662,688]" pageId="13" pageNumber="14" value="2009-06-01">1 Jun. 2009</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED51CCDB44577C63FD70FD54FCA84989" box="[752,815,662,688]" pageId="13" pageNumber="14">ERC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A079544577C60FF3DFD15FE834CD2" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BFF541E44577C63FF3DFD15FEC949C8" blockId="13.[189,334,727,753]" box="[189,334,727,753]" pageId="13" pageNumber="14">
|
||
<heading id="D0B7E37244577C63FF3DFD15FEC949C8" bold="true" box="[189,334,727,753]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B934880C44577C63FF3DFD15FEC949C8" bold="true" box="[189,334,727,753]" pageId="13" pageNumber="14">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44577C63FF6AFCC4FEDB4819" blockId="13.[189,746,774,836]" box="[234,348,774,800]" pageId="13" pageNumber="14">
|
||
<emphasis id="B934880C44577C63FF6AFCC4FEDB4819" bold="true" box="[234,348,774,800]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54FBEABC44577C63FF6AFCC4FEDB4819" box="[234,348,774,800]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44577C63FF3DFCE8FD6D487D" blockId="13.[189,746,774,836]" box="[189,746,809,836]" pageId="13" pageNumber="14">
|
||
MEASUREMENTS. Length
|
||
<quantity id="4CB8F9FB44577C63FE56FCEBFDB4487D" box="[470,563,809,836]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="13" pageNumber="14" unit="mm" value="8.2">8.2 mm</quantity>
|
||
; width
|
||
<quantity id="4CB8F9FB44577C63FD09FCEBFD61487D" box="[649,742,809,836]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="13" pageNumber="14" unit="mm" value="5.1">5.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44577C60FF3DFCA9FEC64A49" blockId="13.[189,1104,874,901]" lastBlockId="14.[189,1399,267,368]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">BODY. Colour mostly black, with propygidium and pygidium dark castaneous. HEAD. Black, slightly shiny, covered with large round punctures; frons and vertex covered with small, recumbent, testaceous scales that do not form scale tufts. Clypeus black, anteriorly sinuate, suctorial brush long.</paragraph>
|
||
<caption id="DF3F049644577C63FF3DF8A5FBF44CD2" ID-DOI="http://doi.org/10.5281/zenodo.3832564" ID-Zenodo-Dep="3832564" httpUri="https://zenodo.org/record/3832564/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1895,1921]" targetBox="[216,1363,985,1846]" targetPageId="13">
|
||
<paragraph id="8BFF541E44577C63FF3DF8A5FBF44CD2" blockId="13.[189,1399,1895,2027]" pageId="13" pageNumber="14">
|
||
<emphasis id="B934880C44577C63FF3DF8A5FE8C4CB8" bold="true" box="[189,267,1895,1921]" pageId="13" pageNumber="14">Fig. 6.</emphasis>
|
||
<taxonomicName id="4C402F9D44577C63FE94F8A5FE734CB8" authority="Ricchiardi, 2017" authorityName="Ricchiardi" authorityYear="2017" box="[276,500,1895,1921]" class="Insecta" family="Scarabaeidae" genus="Hybovalgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
|
||
<emphasis id="B934880C44577C63FE94F8A5FE734CB8" box="[276,500,1895,1921]" italics="true" pageId="13" pageNumber="14">Hybovalgus calvus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207357744577C63FE7CF8AAFDDF4CBB" box="[508,600,1896,1922]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B934880C44577C63FDE0F8A5FD5D4CB8" bold="true" box="[608,730,1895,1921]" pageId="13" pageNumber="14">A–C, H–I</emphasis>
|
||
. HT ♂ (MNHUB).
|
||
<emphasis id="B934880C44577C63FC48F8A5FC5A4CB8" bold="true" box="[968,989,1895,1921]" pageId="13" pageNumber="14">A</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B934880C44577C63FB6FF8A5FA844CB8" bold="true" box="[1263,1283,1895,1921]" pageId="13" pageNumber="14">B</emphasis>
|
||
. Protibia.
|
||
<emphasis id="B934880C44577C63FF3DF848FF554C9D" bold="true" box="[189,210,1930,1956]" pageId="13" pageNumber="14">C</emphasis>
|
||
. Pygidium.
|
||
<emphasis id="B934880C44577C63FEE8F848FE1E4C9D" bold="true" box="[360,409,1930,1956]" pageId="13" pageNumber="14">H–I</emphasis>
|
||
. Aedeagus (grey: parameres; white: phallobase).
|
||
<emphasis id="B934880C44577C63FC72F848FB8E4C9D" bold="true" box="[1010,1033,1930,1956]" pageId="13" pageNumber="14">H</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="B934880C44577C63FB3DF848FB4F4C9D" bold="true" box="[1213,1224,1930,1956]" pageId="13" pageNumber="14">I</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="B934880C44577C63FF3DF86FFF7F4CFE" bold="true" box="[189,248,1965,1991]" pageId="13" pageNumber="14">D–G</emphasis>
|
||
. PT ♀ (ERC).
|
||
<emphasis id="B934880C44577C63FE24F86CFE3E4CF1" bold="true" box="[420,441,1966,1992]" pageId="13" pageNumber="14">D</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B934880C44577C63FD43F86CFD504CF1" bold="true" box="[707,727,1966,1992]" pageId="13" pageNumber="14">E</emphasis>
|
||
. Protibia.
|
||
<emphasis id="B934880C44577C63FCD0F86CFCE54CF1" bold="true" box="[848,866,1966,1992]" pageId="13" pageNumber="14">F</emphasis>
|
||
. Pygidium, frontal view.
|
||
<emphasis id="B934880C44577C63FB09F86FFB274CFE" bold="true" box="[1161,1184,1965,1991]" pageId="13" pageNumber="14">G</emphasis>
|
||
. Pygidium, lateral view.
|
||
<emphasis id="B934880C44577C63FE83F813FEB04CD2" bold="true" box="[259,311,2001,2027]" pageId="13" pageNumber="14">J–K</emphasis>
|
||
. PT ♂ (ERC).
|
||
<emphasis id="B934880C44577C63FE67F813FE714CD2" bold="true" box="[487,502,2001,2027]" pageId="13" pageNumber="14">J</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B934880C44577C63FC83F813FC9D4CD2" bold="true" box="[771,794,2001,2027]" pageId="13" pageNumber="14">K</emphasis>
|
||
. Propygidium and pygidium.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF541E44547C60FF3DFE58FD584956" blockId="14.[189,1399,410,624]" pageId="14" pageNumber="15">PRONOTUM. Black, nearly glabrous, trapezoidal, slightly crenate at sides, laterally depressed at center of lateral margins; anterior angle acute, posterior angle right-angled, with apex rounded; posterior margin with central part showily produced backwards, V-shaped, with apex rounded. Carinae rounded, parallel, ending around middle of disk; central small carina not present; lateral carinae not present; with 2 testaceous barely noticeable central carina scale tufts; without any other scale tufts. Covered with small, decumbent, scattered, testaceous scales.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFD5BFACE498A" blockId="14.[189,1353,665,691]" box="[189,1353,665,691]" pageId="14" pageNumber="15">SCUTELLUM. Black, long, apically rounded, covered with small, decumbent, thick, testaceous scales.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFD1FFD764878" blockId="14.[189,1399,732,833]" pageId="14" pageNumber="15">ELYTRA. Black, slightly shiny, mostly glabrous, with barely noticeable juxtascutellar depression. Humbones slightly prominent, without scale tufts; anteo-central part of disk covered with hardly noticeable small, thick, decumbent black scales.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFCAEFEB448E9" blockId="14.[189,1399,875,976]" pageId="14" pageNumber="15">PROPYGIDIUM. Dark castaneous, covered with thick, decumbent, small, coffee bean shaped, testaceous scales that form two small propygidium scale tufts; centre of posterior margin depressed; cones short and blunt.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFC38FD8B4FBD" blockId="14.[189,1398,1017,1156]" pageId="14" pageNumber="15">PYGIDIUM. Dark castaneous, wider than long, slightly depressed at sides in ventral half. Covered with thick, decumbent, small, coffee grain shaped, testaceous scales which form a fringe of long scales at apex. Apex rounded, without projection. Under apex, in ventral view, there is a hollowed, semicircular, glabrous, horizontal surface.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFB6CFC4C4FD7" blockId="14.[189,1399,1197,1262]" pageId="14" pageNumber="15">
|
||
PROTIBIA. Black, with 5 external teeth: 1–3, long, 4–5 smaller, all rounded at apex. Teeth 3–4 closer to one another than 1–3; tooth 5 rather spaced apart from 4 (
|
||
<figureCitation id="137B489B44547C60FCE1FB11FC3A4FD7" box="[865,957,1235,1262]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[216,1363,985,1846]" captionTargetId="figure@13.[202,1385,952,1854]" captionTargetPageId="13" captionText="Fig. 6. Hybovalgus calvus sp. nov. A–C, H–I. HT ♂ (MNHUB). A. Habitus, dorsal view. B. Protibia. C. Pygidium. H–I. Aedeagus (grey: parameres; white: phallobase). H. Dorsal view. I. Lateral view. D–G. PT ♀ (ERC). D. Habitus, dorsal view. E. Protibia. F. Pygidium, frontal view. G. Pygidium, lateral view. J–K. PT ♂ (ERC). J. Habitus, dorsal view. K. Propygidium and pygidium." figureDoi="http://doi.org/10.5281/zenodo.3832564" httpUri="https://zenodo.org/record/3832564/files/figure.png" pageId="14" pageNumber="15">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFAD5FDEB4E6F" blockId="14.[189,1399,1303,1367]" pageId="14" pageNumber="15">MESO- AND METAFEMUR. Black, with a fringe of small, testaceous scales at centre of posterior margin which do not form a small scale tuft.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFA42FDE74E86" blockId="14.[189,1399,1408,1471]" pageId="14" pageNumber="15">METATIBIA. Black, centrally enlarged at posterior margin, without a tooth at centre. First metatarsomere long 1.8 times the length of second.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DFA2BFB814D11" blockId="14.[189,1399,1513,1577]" pageId="14" pageNumber="15">ABDOMEN. Dark castaneous, covered with thick, recumbent, testaceous scales, scattered at centre of 1–3 visible sternites. Anal sternites, with same scales but centrally glabrous.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DF990FDDF4D54" blockId="14.[189,600,1616,1645]" box="[189,600,1616,1645]" pageId="14" pageNumber="15">
|
||
<emphasis id="B934880C44547C60FF3DF990FECB4D55" bold="true" box="[189,332,1616,1644]" pageId="14" pageNumber="15">
|
||
<typeStatus id="54FBEABC44547C60FF3DF990FEA84D55" box="[189,303,1618,1644]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
|
||
<specimenCount id="9D469F9744547C60FEB6F992FECB4D55" box="[310,332,1616,1644]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
</emphasis>
|
||
(differences from HT)
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DF941FC7D4DA4" blockId="14.[189,1018,1667,1694]" box="[189,1018,1667,1694]" pageId="14" pageNumber="15">
|
||
MEASUREMENTS. Length
|
||
<quantity id="4CB8F9FB44547C60FE56F941FDB44DA4" box="[470,563,1667,1694]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="15" unit="mm" value="9.5">9.5 mm</quantity>
|
||
; width
|
||
<quantity id="4CB8F9FB44547C60FD09F941FD614DA4" box="[649,742,1667,1694]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="15" unit="mm" value="5.7">5.7 mm</quantity>
|
||
. Body dark castaneous.
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DF904FEF14C68" blockId="14.[189,1399,1734,1873]" pageId="14" pageNumber="15">The female is quite similar to the male, not showing great differences in the shape of the posterior margins of the pronotum or pygidium apex. The first is pointed at the apex, the latter is regularly rounded. The propygidium is much wider than long, shaped as in the male but with a small depression close to each superior corner.</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DF8B9FE564CAD" blockId="14.[189,465,1914,1941]" box="[189,465,1914,1941]" pageId="14" pageNumber="15">
|
||
<heading id="D0B7E37244547C60FF3DF8B9FE564CAD" bold="true" box="[189,465,1914,1941]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B934880C44547C60FF3DF8B9FE564CAD" bold="true" box="[189,465,1914,1941]" pageId="14" pageNumber="15">Type series variability</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44547C60FF3DF869FE834CD2" blockId="14.[189,1399,1963,2027]" pageId="14" pageNumber="15">
|
||
Males: length 8.0–
|
||
<quantity id="4CB8F9FB44547C60FE19F869FE714CFF" box="[409,502,1963,1990]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="14" pageNumber="15" unit="mm" value="8.4">8.4 mm</quantity>
|
||
; width 4.7–5.3 mmm. A PT
|
||
<specimenCount id="9D469F9744547C60FCC3F86EFCDE4CFF" box="[835,857,1964,1990]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
|
||
has the propygidium, pygidium and abdomen black.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A079544557C61FF3DFEC9FA874A6D" pageId="15" pageNumber="16" type="biology_ecology">
|
||
<paragraph id="8BFF541E44557C61FF3DFEC9FEE74A1C" blockId="15.[189,352,267,293]" box="[189,352,267,293]" pageId="15" pageNumber="16">
|
||
<heading id="D0B7E37244557C61FF3DFEC9FEE74A1C" bold="true" box="[189,352,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B934880C44557C61FF3DFEC9FEE74A1C" bold="true" box="[189,352,267,293]" pageId="15" pageNumber="16">Flight period</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44557C61FF3DFEF8FA874A6D" blockId="15.[189,1280,314,341]" box="[189,1280,314,341]" pageId="15" pageNumber="16">
|
||
The Vietnamese
|
||
<typeStatus id="54FBEABC44557C61FE01FEF8FE384A6D" box="[385,447,314,340]" pageId="15" pageNumber="16">types</typeStatus>
|
||
were collected from May to early June at an elevation of
|
||
<quantity id="4CB8F9FB44557C61FBE5FEF8FB7B4A6D" box="[1125,1276,314,341]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="15" pageNumber="16" unit="m" value="1100.0" valueMax="1500.0" valueMin="700.0">700–1500 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A079544557C61FF3DFEB9FD7A4AFC" box="[189,765,379,453]" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="8BFF541E44557C61FF3DFEB9FED14AAC" blockId="15.[189,342,379,405]" box="[189,342,379,405]" pageId="15" pageNumber="16">
|
||
<heading id="D0B7E37244557C61FF3DFEB9FED14AAC" bold="true" box="[189,342,379,405]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B934880C44557C61FF3DFEB9FED14AAC" bold="true" box="[189,342,379,405]" pageId="15" pageNumber="16">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF541E44557C61FF3DFE69FD7A4AFC" blockId="15.[189,765,426,453]" box="[189,765,426,453]" pageId="15" pageNumber="16">
|
||
Present in northern
|
||
<collectingCountry id="F357148E44557C61FE23FE68FD814AFD" box="[419,518,426,452]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
|
||
and
|
||
<collectingCountry id="F357148E44557C61FDBFFE68FD024AFD" box="[575,645,426,452]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49849AFC44557C61FD15FE68FD774AFC" box="[661,752,426,453]" country="China" name="Hainan" pageId="15" pageNumber="16">Hainan</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |