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<document id="777D872F8F1D123CE87219343960FAFD" ID-DOI="10.1515/aemnp-2017-0084" ID-ISSN="0374-1036" ID-Zenodo-Dep="5315312" ID-ZooBank="03305E03-AF44-4C6D-9E2B-9A3EE979C5AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630122446806" checkinUser="marcus" docAuthor="Wolski, Andrzej" docDate="2017" docId="03E287C9F446FFFFFE44FB06CF62F9D8" docLanguage="en" docName="ActaEntMusNatPra.2017.57.2.399-455.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57 (2)" docSource="http://dx.doi.org/10.1515/aemnp-2017-0084" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Cylapus luridus Wolski 2017, sp. nov" docType="treatment" docVersion="4" lastPageNumber="430" masterDocId="FFDBFFB1F458FFE0FFB8FFFACD36FFEB" masterDocTitle="Taxonomic review of the plant bug genera Amapacylapus and Cylapus with descriptions of two new species and a key to the genera of Cylapini (Hemiptera: Heteroptera: Miridae)" masterLastPageNumber="455" masterPageNumber="399" pageNumber="429" updateTime="1699161098555" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="12145A57666285724CD22A27BA47CC82">Taxonomic review of the plant bug genera Amapacylapus and Cylapus with descriptions of two new species and a key to the genera of Cylapini (Hemiptera: Heteroptera: Miridae)</mods:title>
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<mods:title id="B35F0F954FA2BDDA5BA7E3D26E7F68C3">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="64B3958848B8779FC7380ECEE3292178">2017</mods:date>
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<mods:number id="15C5D4445C96F9599F67EBE52CF46E9C">2017-12-31</mods:number>
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<paragraph id="8BF436DFF446FFFEFE44FB06CE37FAF8" blockId="30.[508,769,1276,1299]" box="[508,769,1276,1299]" pageId="30" pageNumber="429">
<heading id="D0BC81B3F446FFFEFE44FB06CE37FAF8" bold="true" box="[508,769,1276,1299]" centered="true" fontSize="8" level="2" pageId="30" pageNumber="429" reason="3">
<emphasis id="B93FEACDF446FFFEFE44FB06CE37FAF8" bold="true" box="[508,769,1276,1299]" pageId="30" pageNumber="429">
<taxonomicName id="4C4B4D5CF446FFFEFE44FB06CF9DFAF8" authority="Wolski, 2017" authorityName="Wolski" authorityYear="2017" box="[508,683,1276,1299]" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="429" phylum="Arthropoda" rank="species" species="luridus" status="sp. nov.">
<emphasis id="B93FEACDF446FFFEFE44FB06CF9DFAF8" bold="true" box="[508,683,1276,1299]" italics="true" pageId="30" pageNumber="429">Cylapus luridus</emphasis>
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<taxonomicNameLabel id="A20C57B6F446FFFEFD09FB06CE37FAF8" box="[689,769,1276,1299]" pageId="30" pageNumber="429" rank="species">sp. nov</taxonomicNameLabel>
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<subSubSection id="C3516554F446FFFEFD90FAE0CFE0FAC6" box="[552,726,1306,1325]" pageId="30" pageNumber="429" type="description">
<paragraph id="8BF436DFF446FFFEFD90FAE0CFE0FAC6" blockId="30.[151,1127,1306,1421]" box="[552,726,1306,1325]" pageId="30" pageNumber="429">
(
<figureCitation id="13702A5AF446FFFEFD96FAE0CF46FAC6" box="[558,624,1306,1325]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="30" pageNumber="429">Figs 43</figureCitation>
,
<figureCitation id="13702A5AF446FFFEFDC1FAE0CFA6FAC6" box="[633,656,1306,1325]" captionStart="Figs 5665" captionStartId="28.[151,188,1220,1239]" captionTargetBox="[151,1126,219,1183]" captionTargetId="figure-10@28.[151,1126,219,1183]" captionTargetPageId="28" captionText="Figs 5665. Lateral view of Cylapus species: 56 C. amazonicus Carvalho, 1989 (J); 57 C. antennatus Carvalho &amp; Fontes, 1968 (J); 58 C. citus Bergroth, 1922 (♀); 59 C. luridus sp. nov. (J); 60 C. marginicollis (Distant, 1883) (J); 61 C. ruficeps Bergroth, 1922 (♀); 62 C. stellatus (Distant, 1883) (♀); 63 C. striatus Reuter, 1907 (♀); 64 C. tenuicornis (Say, 1832) (J); 65 C. tucuruiensis Carvalho, 1989 (J)." figureDoi="http://doi.org/10.5281/zenodo.5315326" httpUri="https://zenodo.org/record/5315326/files/figure.png" pageId="30" pageNumber="429">59</figureCitation>
,
<figureCitation id="13702A5AF446FFFEFD21FAE0CFF9FAC6" box="[665,719,1306,1325]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="30" pageNumber="429">8589</figureCitation>
)
</paragraph>
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<subSubSection id="C3516554F446FFFEFF2FFABECF14FA67" pageId="30" pageNumber="429" type="materials_examined">
<paragraph id="8BF436DFF446FFFEFF2FFABECF14FA67" blockId="30.[151,1127,1306,1421]" pageId="30" pageNumber="429">
<emphasis id="B93FEACDF446FFFEFF2FFABECC2DFABC" bold="true" box="[151,283,1348,1367]" pageId="30" pageNumber="429">Type material.</emphasis>
<materialsCitation id="3B233C82F446FFFEFEA7FABEC932FA99" collectingDate="1997-11-04" collectingDateMax="1997-11-16" collectingDateMin="1997-11-04" collectionCode="USNM" country="Brazil" location="Ariquemes" pageId="30" pageNumber="429" specimenCount="1" stateProvince="Rondonia" typeStatus="holotype">
<emphasis id="B93FEACDF446FFFEFEA7FABECCEDFABC" bold="true" box="[287,475,1348,1367]" pageId="30" pageNumber="429">
<collectingCountry id="F35C764FF446FFFEFEA7FABECC42FABC" box="[287,372,1348,1367]" name="Brazil" pageId="30" pageNumber="429">BRAZIL</collectingCountry>
:
<collectingRegion id="498FF83DF446FFFEFEC4FABECCE2FABE" box="[380,468,1348,1367]" country="Brazil" name="Rondonia" pageId="30" pageNumber="429">RONDÔNIA</collectingRegion>
:
</emphasis>
<typeStatus id="54F0887DF446FFFEFE67FABECF05FABE" box="[479,563,1348,1367]" pageId="30" pageNumber="429" type="holotype">HOLOTYPE</typeStatus>
: J, “Brazil: Rondônia,
<quantity id="4CB39B3AF446FFFEFD43FABECE06FABC" box="[763,816,1348,1367]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="30" pageNumber="429" unit="km" value="62.0">62 km</quantity>
SW
<location id="8E946004F446FFFEFCEFFABECE81FABC" LSID="urn:lsid:plazi:treatment:03E287C9F446FFFFFE44FB06CF62F9D8:8E946004F446FFFEFCEFFABECE81FABC" box="[855,951,1348,1367]" country="Brazil" name="Ariquemes" pageId="30" pageNumber="429" stateProvince="Rondonia">Ariquemes</location>
, nr Fzda [= Fazenda, farm] Rancho Grande,
<date id="FFF5101FF446FFFEFEEDFAA5CCE3FA99" box="[341,469,1375,1394]" pageId="30" pageNumber="429" value="1997-11-04" valueMax="1997-11-16" valueMin="1997-11-04">
<collectingDate id="EFB1E9F7F446FFFEFEEDFAA5CCE3FA99" box="[341,469,1375,1394]" pageId="30" pageNumber="429" value="1997-11-04" valueMax="1997-11-16" valueMin="1997-11-04">416XI1997</collectingDate>
</date>
JE Eger; Drake Collection Eger Accesion 1998” (
<figureCitation id="13702A5AF446FFFEFCC8FAA5CE9AFA99" box="[880,940,1375,1394]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="30" pageNumber="429">Fig.43</figureCitation>
) (
<collectionCode id="ED5AAE1AF446FFFEFC04FAA5CEC8FA99" box="[956,1022,1375,1394]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="429" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B233C82F446FFFEFBB2FAA5CF28FA67" collectingDate="1997-11-04" collectingDateMax="1997-11-16" collectingDateMin="1997-11-04" collectionCode="USNM" country="Brazil" location="Ariquemes" pageId="30" pageNumber="429" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<typeStatus id="54F0887DF446FFFEFBB2FAA5C956FA9B" box="[1034,1120,1375,1394]" pageId="30" pageNumber="429" type="paratype">PARATYPES</typeStatus>
: 2 JJ, the same data as for holotype (
<collectionCode id="ED5AAE1AF446FFFEFE6EFA83CF2EFA67" box="[470,536,1401,1420]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="429" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3516554F446FFFFFF2FFA5FCC19FDF8" lastPageId="31" lastPageNumber="430" pageId="30" pageNumber="429" type="diagnosis">
<paragraph id="8BF436DFF446FFFFFF2FFA5FCE2DFEB8" blockId="30.[151,1127,1445,1564]" lastBlockId="31.[113,1089,220,1587]" lastPageId="31" lastPageNumber="430" pageId="30" pageNumber="429">
<emphasis id="B93FEACDF446FFFEFF2FFA5FCC3AFA57" bold="true" box="[151,268,1445,1468]" pageId="30" pageNumber="429">Diagnosis.</emphasis>
Recognized by the following combinations of characters: antennal segment II in males thicker than segment I (
<figureCitation id="13702A5AF446FFFEFE64FA3FCF1BFA37" box="[476,557,1477,1500]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="30" pageNumber="429">Fig. 43</figureCitation>
); pronotum black, tinged with red laterally (
<figureCitation id="13702A5AF446FFFEFBB1FA3FC96CFA37" box="[1033,1114,1477,1500]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="30" pageNumber="429">Fig. 43</figureCitation>
); outer margin of clavus with narrow, yellowish stripe along entire length (
<figureCitation id="13702A5AF446FFFEFC2AFA1FCED7FA17" box="[914,993,1509,1532]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="30" pageNumber="429">Fig. 43</figureCitation>
); endosomal sclerites: es1 straight, cylindrical, serrate at apical one fourth; es2 weakly arcuate, cylindrical, weakly broadened and serrate at apical one sixth; es3 and es4 longest, moderately broadened, sharply pointed; es3 with dextrolateral margin straight and sinistrolateral margin arcuate; es4 with dextrolateral margin arcuate, sinistrolateral margin sinuate (
<figureCitation id="13702A5AF447FFFFFCAFFEE6CE53FED8" box="[791,869,284,307]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Fig. 85</figureCitation>
); left paramere when viewed dorsally with sensory lobe strongly developed (
<figureCitation id="13702A5AF447FFFFFD07FEC6CE39FEB8" box="[703,783,316,339]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Fig. 86</figureCitation>
).
</paragraph>
<paragraph id="8BF436DFF447FFFFFF28FEA6CC19FDF8" blockId="31.[113,1089,220,1587]" pageId="31" pageNumber="430">
Most similar to
<taxonomicName id="4C4B4D5CF447FFFFFE87FEA6CCE8FE98" authorityName="Carvalho" authorityYear="1989" box="[319,478,348,371]" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="430" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B93FEACDF447FFFFFE87FEA6CCE8FE98" box="[319,478,348,371]" italics="true" pageId="31" pageNumber="430">C. amazonicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B4D5CF447FFFFFE55FEA6CFB2FE98" authorityName="Carvalho &amp; Fontes" authorityYear="1968" box="[493,644,348,371]" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="430" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="B93FEACDF447FFFFFE55FEA6CFB2FE98" box="[493,644,348,371]" italics="true" pageId="31" pageNumber="430">C. antennatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B4D5CF447FFFFFD2BFEA6CE77FE98" baseAuthorityName="Distant" baseAuthorityYear="1883" box="[659,833,348,371]" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="430" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B93FEACDF447FFFFFD2BFEA6CE77FE98" box="[659,833,348,371]" italics="true" pageId="31" pageNumber="430">C. marginicollis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4B4D5CF447FFFFFC38FEA6C915FE98" authorityName="Carvalho" authorityYear="1989" box="[896,1059,348,371]" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="430" phylum="Arthropoda" rank="species" species="tucuruiensis">
<emphasis id="B93FEACDF447FFFFFC38FEA6C915FE98" box="[896,1059,348,371]" italics="true" pageId="31" pageNumber="430">C. tucuruiensis</emphasis>
</taxonomicName>
in sharing the thick antennal segment in males (
<figureCitation id="13702A5AF447FFFFFDECFE86CFE6FE78" box="[596,720,380,403]" captionStart="Figs 3640" captionStartId="22.[151,188,1541,1560]" captionTargetBox="[153,1127,234,1526]" captionTargetId="figure-10@22.[151,1127,219,1527]" captionTargetPageId="22" captionText="Figs 3640. Dorsal habitus photographs of Cylapus species: 36 C. amazonicus Carvalho, 1989 (J); 3739 C. antennatus Carvalho &amp; Fontes, 1968 (37 J holotype, 38 J, 39 ♀); 40 C. citus Bergroth, 1922 (J)." figureDoi="http://doi.org/10.5281/zenodo.5315324" httpUri="https://zenodo.org/record/5315324/files/figure.png" pageId="31" pageNumber="430">Figs 36, 38</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFD63FE86CFCEFE78" box="[731,760,380,403]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="31" pageNumber="430">43</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFCBBFE86CE16FE78" box="[771,800,380,403]" captionStart="Figs 4855" captionStartId="28.[151,187,1514,1533]" captionText="Figs 4855. Dorsal habitus photographs of Cylapus species:48, 49 C.stellatus (Distant, 1883) (48 J paralectotype, 49 ♀ lectotype); 50, 51 C. striatus Reuter, 1907 (50 ♀, 51 J); 52, 53 C. tenuicornis (Say, 1832) (52 ♀, 53 J); 54, 55 C. tucuruiensis Carvalho, 1989 (54 ♀, 55 J)." pageId="31" pageNumber="430">55</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFC93FE86CEF7FE78" box="[811,961,380,403]" captionStart="Figs 5665" captionStartId="28.[151,188,1220,1239]" captionTargetBox="[151,1126,219,1183]" captionTargetId="figure-10@28.[151,1126,219,1183]" captionTargetPageId="28" captionText="Figs 5665. Lateral view of Cylapus species: 56 C. amazonicus Carvalho, 1989 (J); 57 C. antennatus Carvalho &amp; Fontes, 1968 (J); 58 C. citus Bergroth, 1922 (♀); 59 C. luridus sp. nov. (J); 60 C. marginicollis (Distant, 1883) (J); 61 C. ruficeps Bergroth, 1922 (♀); 62 C. stellatus (Distant, 1883) (♀); 63 C. striatus Reuter, 1907 (♀); 64 C. tenuicornis (Say, 1832) (J); 65 C. tucuruiensis Carvalho, 1989 (J)." figureDoi="http://doi.org/10.5281/zenodo.5315326" httpUri="https://zenodo.org/record/5315326/files/figure.png" pageId="31" pageNumber="430">56, 57, 59, 60</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFC75FE86CEDDFE78" box="[973,1003,380,403]" captionStart="Figs 9196" captionStartId="36.[151,188,1484,1503]" captionText="Figs 9196. Scanning electron micrographs of Cylapus marginicollis (Distant, 1883) (9194) and C. ruficeps Bergroth, 1922 (9596): 91 dorsal habitus (♀); 92 lateral view (J); 93 thoracic pleura; 94 mesotarsus; 95 lateral views; 96 thoracic pleura. Abbreviations: ea = evaporative areas; msp = metathoracic spiracle; pc = posterior carina; per = peritreme." pageId="31" pageNumber="430">92</figureCitation>
). It can, however, be distinguished by the shape of the male genitalia (
<figureCitation id="13702A5AF447FFFFFCBAFE66CE49FE58" box="[770,895,412,435]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Figs 8589</figureCitation>
). With
<taxonomicName id="4C4B4D5CF447FFFFFC74FE66CD90FE38" authorityName="Carvalho" authorityYear="1989" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="430" phylum="Arthropoda" rank="species" species="tucuruiensis">
<emphasis id="B93FEACDF447FFFFFC74FE66CD90FE38" italics="true" pageId="31" pageNumber="430">C. tucuruiensis</emphasis>
</taxonomicName>
in shares outer margin of clavus with yellow stripe along entire length. It can be easily distinguished by having yellow patch on corium (
<figureCitation id="13702A5AF447FFFFFD30FE26CFEAFE18" box="[648,732,476,499]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="31" pageNumber="430">Figs 43</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFD50FE26CE1BFE18" box="[744,813,476,499]" captionStart="Figs 4855" captionStartId="28.[151,187,1514,1533]" captionText="Figs 4855. Dorsal habitus photographs of Cylapus species:48, 49 C.stellatus (Distant, 1883) (48 J paralectotype, 49 ♀ lectotype); 50, 51 C. striatus Reuter, 1907 (50 ♀, 51 J); 52, 53 C. tenuicornis (Say, 1832) (52 ♀, 53 J); 54, 55 C. tucuruiensis Carvalho, 1989 (54 ♀, 55 J)." pageId="31" pageNumber="430">54, 55</figureCitation>
) and male genitalia (
<figureCitation id="13702A5AF447FFFFFBA9FE26CD80FDF8" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Figs 8589</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFF79FE06CC15FDF8" box="[193,291,508,531]" captionStart="Figs 135144" captionStartId="48.[151,188,1320,1339]" captionTargetBox="[229,1073,210,1302]" captionTargetId="figure-154@48.[208,1077,205,1302]" captionTargetPageId="48" captionText="Figs 135144. Male genitalia of Cylapus tenuicornis (Say, 1832) (135139) and C. tucuruiensis Carvalho, 1989 (140144): 135, 140 endosoma; 136, 141 left paramere (dorsal view); 137, 142 left paramere (right lateral view); 138, 143 apical process of left paramere; 139, 144 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315354" httpUri="https://zenodo.org/record/5315354/files/figure.png" pageId="31" pageNumber="430">140144</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3516554F447FFFFFFC9FDE6CFCDFA58" pageId="31" pageNumber="430" type="description">
<paragraph id="8BF436DFF447FFFFFFC9FDE6C977FC78" blockId="31.[113,1089,220,1587]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9FDE6CC8BFDD8" bold="true" box="[113,445,540,563]" pageId="31" pageNumber="430">
Description.
<emphasis id="B93FEACDF447FFFFFEB8FDE6CC0EFDD8" bold="true" box="[256,312,540,563]" italics="true" pageId="31" pageNumber="430">Male</emphasis>
. Coloration
</emphasis>
(
<figureCitation id="13702A5AF447FFFFFE71FDE6CF2CFDD8" box="[457,538,540,563]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="31" pageNumber="430">Figs 43</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFD9BFDE6CF77FDD8" box="[547,577,540,563]" captionStart="Figs 5665" captionStartId="28.[151,188,1220,1239]" captionTargetBox="[151,1126,219,1183]" captionTargetId="figure-10@28.[151,1126,219,1183]" captionTargetPageId="28" captionText="Figs 5665. Lateral view of Cylapus species: 56 C. amazonicus Carvalho, 1989 (J); 57 C. antennatus Carvalho &amp; Fontes, 1968 (J); 58 C. citus Bergroth, 1922 (♀); 59 C. luridus sp. nov. (J); 60 C. marginicollis (Distant, 1883) (J); 61 C. ruficeps Bergroth, 1922 (♀); 62 C. stellatus (Distant, 1883) (♀); 63 C. striatus Reuter, 1907 (♀); 64 C. tenuicornis (Say, 1832) (J); 65 C. tucuruiensis Carvalho, 1989 (J)." figureDoi="http://doi.org/10.5281/zenodo.5315326" httpUri="https://zenodo.org/record/5315326/files/figure.png" pageId="31" pageNumber="430">59</figureCitation>
). Dorsum black with red and yellow areas.
<emphasis id="B93FEACDF447FFFFFC47FDE6C977FDD8" bold="true" box="[1023,1089,540,563]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFC47FDE6C90CFDD8" bold="true" box="[1023,1082,540,563]" italics="true" pageId="31" pageNumber="430">Head</emphasis>
.
</emphasis>
Vertex dark brown, tinged with dirty yellow; frons, mandibular and maxillary plates, clypeus, and buccula black; frons indistinctly tinged with yellow basally; gula yellow; antennal segments I and II black; segments III and IV dark brown; segment III with relatively broad, yellow annulation basally; labium black.
<emphasis id="B93FEACDF447FFFFFD9FFD66CF49FD58" bold="true" box="[551,639,668,691]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFD9FFD66CF4EFD58" bold="true" box="[551,632,668,691]" italics="true" pageId="31" pageNumber="430">Thorax</emphasis>
.
</emphasis>
<emphasis id="B93FEACDF447FFFFFD3DFD66CFD9FD58" box="[645,751,668,691]" italics="true" pageId="31" pageNumber="430">Pronotum</emphasis>
black, tinged with red laterally; collar dirty yellow; humeral angle yellow.
<emphasis id="B93FEACDF447FFFFFD94FD46CE7FFD38" box="[556,841,700,723]" italics="true" pageId="31" pageNumber="430">Mesoscutum and scutellum</emphasis>
black, narrowly yellow apically.
<emphasis id="B93FEACDF447FFFFFF6BFD26CCB4FD18" box="[211,386,732,755]" italics="true" pageId="31" pageNumber="430">Thoracic pleura</emphasis>
black; posterior margin of mesepimeron yellow red along entire length; metathoracic scent gland evaporative area yellow tinged with fuscous.
<emphasis id="B93FEACDF447FFFFFC7CFD06C977FCF8" box="[964,1089,764,787]" italics="true" pageId="31" pageNumber="430">Hemelytron</emphasis>
black; outer margin of clavus and inner margin of corium with yellow, narrow stripe along entire length; apical margin of corium and basal margin of cuneus yellow along entire length.
<emphasis id="B93FEACDF447FFFFFFC9FCA6CD95FC98" box="[113,163,860,883]" italics="true" pageId="31" pageNumber="430">Legs</emphasis>
. Coxae black; remaining segments dark brown.
<emphasis id="B93FEACDF447FFFFFD2FFCA6CE36FC98" bold="true" box="[663,768,860,883]" italics="true" pageId="31" pageNumber="430">Abdomen</emphasis>
black.
<emphasis id="B93FEACDF447FFFFFCF1FCA6CDE2FC78" bold="true" pageId="31" pageNumber="430">Structure, texture and vestiture</emphasis>
(
<figureCitation id="13702A5AF447FFFFFF59FC86CC04FC78" box="[225,306,892,915]" captionStart="Figs 4147" captionStartId="24.[151,188,1517,1536]" captionText="Figs 4147. Dorsal habitus photographs of Cylapus species: 41, 42 C. citus Bergroth, 1922 (41 ♀, 42 holotype); 43 C. luridus sp. nov. (J holotype); 44, 45 C. marginicollis (Distant, 1883) (44 J, 45 ♀); 46, 47 C. ruficeps Bergroth, 1922 (46 J holotype, 47 ♀)." pageId="31" pageNumber="430">Figs 43</figureCitation>
,
<figureCitation id="13702A5AF447FFFFFE83FC86CC6FFC78" box="[315,345,892,915]" captionStart="Figs 5665" captionStartId="28.[151,188,1220,1239]" captionTargetBox="[151,1126,219,1183]" captionTargetId="figure-10@28.[151,1126,219,1183]" captionTargetPageId="28" captionText="Figs 5665. Lateral view of Cylapus species: 56 C. amazonicus Carvalho, 1989 (J); 57 C. antennatus Carvalho &amp; Fontes, 1968 (J); 58 C. citus Bergroth, 1922 (♀); 59 C. luridus sp. nov. (J); 60 C. marginicollis (Distant, 1883) (J); 61 C. ruficeps Bergroth, 1922 (♀); 62 C. stellatus (Distant, 1883) (♀); 63 C. striatus Reuter, 1907 (♀); 64 C. tenuicornis (Say, 1832) (J); 65 C. tucuruiensis Carvalho, 1989 (J)." figureDoi="http://doi.org/10.5281/zenodo.5315326" httpUri="https://zenodo.org/record/5315326/files/figure.png" pageId="31" pageNumber="430">59</figureCitation>
). Dorsum covered with relatively long, semirecumbent and erect setae.
</paragraph>
<paragraph id="8BF436DFF447FFFFFFC9FC66C977FC58" blockId="31.[113,1089,220,1587]" box="[113,1089,924,947]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9FC66CD85FC58" bold="true" box="[113,179,924,947]" italics="true" pageId="31" pageNumber="430">Head.</emphasis>
antennal segment II thicker than segment I, gradually becoming narrower toward apex.
</paragraph>
<paragraph id="8BF436DFF447FFFFFFC9FC46CCB1FB38" blockId="31.[113,1089,220,1587]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9FC46CDFFFC38" bold="true" box="[113,201,956,979]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9FC46CDF4FC38" bold="true" box="[113,194,956,979]" italics="true" pageId="31" pageNumber="430">Thorax</emphasis>
.
</emphasis>
<emphasis id="B93FEACDF447FFFFFF76FC46CC01FC38" box="[206,311,956,979]" italics="true" pageId="31" pageNumber="430">Pronotum</emphasis>
calli moderately developed.
<emphasis id="B93FEACDF447FFFFFDDBFC46CFFAFC38" box="[611,716,956,979]" italics="true" pageId="31" pageNumber="430">Scutellum</emphasis>
moderately convex.
<emphasis id="B93FEACDF447FFFFFC1CFC46C977FC38" bold="true" box="[932,1089,956,979]" italics="true" pageId="31" pageNumber="430">Male genitalia</emphasis>
(
<figureCitation id="13702A5AF447FFFFFFC1FC26CDC0FC18" box="[121,246,988,1011]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Figs 8589</figureCitation>
).
<emphasis id="B93FEACDF447FFFFFEB0FC26CC46FC18" box="[264,368,988,1011]" italics="true" pageId="31" pageNumber="430">Aedeagus</emphasis>
(
<figureCitation id="13702A5AF447FFFFFEC7FC26CCF9FC18" box="[383,463,988,1011]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Fig. 85</figureCitation>
). Endosoma with four sclerites (es1es4): es1 straight, cylindrical, serrate at apical one fourth; es2 weakly arcuate, cylindrical, weakly broadened and serrate at apical one sixth; es3 and es4 longest, moderately broadened, sharply pointed; es3 with dextrolateral margin straight and sinistrolateral margin arcuate; es4 with dextrolateral margin arcuate, sinistrolateral margin sinuate.
<emphasis id="B93FEACDF447FFFFFDD0FBA6CE37FB98" box="[616,769,1116,1139]" italics="true" pageId="31" pageNumber="430">Left paramere</emphasis>
(
<figureCitation id="13702A5AF447FFFFFCA8FBA6CEB9FB98" box="[784,911,1116,1139]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Figs 8688</figureCitation>
). Apical process weakly arcuate and tapering toward apex; paramere body arcuate in lateral view, with strongly developed sensory lobe.
<emphasis id="B93FEACDF447FFFFFECCFB66CF2FFB58" box="[372,537,1180,1203]" italics="true" pageId="31" pageNumber="430">Right paramere</emphasis>
(
<figureCitation id="13702A5AF447FFFFFD9EFB66CF42FB58" box="[550,628,1180,1203]" captionStart="Figs 8089" captionStartId="34.[151,188,1367,1386]" captionTargetBox="[139,1129,234,1319]" captionTargetId="figure-97@34.[130,1136,223,1322]" captionTargetPageId="34" captionText="Figs 8089. Male genitalia of Cylapus citus Bergroth, 1922 (8084) and C. luridus sp. nov. (8589): 80, 85 endosoma; 81, 86 left paramere (dorsal view); 82, 87 left paramere (right lateral view); 83, 88 apical process of left paramere; 84, 89 right paramere (left lateral view). Abbreviations: bpr = basal process; dss = sclerotized portion of ductus seminis inside endosoma; es14 = endosomal sclerites 14; pb = paramere body; sg = secondary gonopore; sl = sensory lobe. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5315338" httpUri="https://zenodo.org/record/5315338/files/figure.png" pageId="31" pageNumber="430">Fig. 89</figureCitation>
). Apical process short, obtuse, both margins of paramere body sinuate.
</paragraph>
<paragraph id="8BF436DFF447FFFFFF28FB26CC57FB18" blockId="31.[113,1089,220,1587]" box="[144,353,1244,1267]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFF28FB26CDDFFB18" bold="true" box="[144,233,1244,1267]" italics="true" pageId="31" pageNumber="430">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF436DFF447FFFFFF28FB06CFCDFA58" blockId="31.[113,1089,220,1587]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFF28FB06CC07FAF8" bold="true" box="[144,305,1276,1299]" italics="true" pageId="31" pageNumber="430">Measurements</emphasis>
(in mm). J (*:
<typeStatus id="54F0887DF447FFFFFE6BFB06CF06FAF8" box="[467,560,1276,1299]" pageId="31" pageNumber="430" type="holotype">holotype</typeStatus>
measurements):
<emphasis id="B93FEACDF447FFFFFD5CFB06CE2DFAF8" box="[740,795,1276,1299]" italics="true" pageId="31" pageNumber="430">Body</emphasis>
. Length: 6.60*6.70, width 1.802.00*.
<emphasis id="B93FEACDF447FFFFFF4FFAE6CC07FAD8" box="[247,305,1308,1331]" italics="true" pageId="31" pageNumber="430">Head</emphasis>
. Length: 0.570.62*, width: 1.30*, interocular distance: 0.58*.
<emphasis id="B93FEACDF447FFFFFC59FAE6C90CFAD8" box="[993,1082,1308,1331]" italics="true" pageId="31" pageNumber="430">Antenna</emphasis>
. Length of segment I: 1.10*1.30, II: 2.382.75*, III: 3.00 (in
<typeStatus id="54F0887DF447FFFFFCAFFAC6CE42FAB8" box="[791,884,1340,1363]" pageId="31" pageNumber="430" type="holotype">holotype</typeStatus>
missing), IV: 1.50 (in
<typeStatus id="54F0887DF447FFFFFF2FFAA6CDC2FA98" box="[151,244,1372,1395]" pageId="31" pageNumber="430" type="holotype">holotype</typeStatus>
missing).
<emphasis id="B93FEACDF447FFFFFED0FAA6CCF7FA98" box="[360,449,1372,1395]" italics="true" pageId="31" pageNumber="430">Labium.</emphasis>
Length of segment I: 0.88*, II: 0.880.90*, III: 1.10*, IV: 0.32*0.35.
<emphasis id="B93FEACDF447FFFFFF4EFA86CC56FA78" box="[246,352,1404,1427]" italics="true" pageId="31" pageNumber="430">Pronotum</emphasis>
. Length: 1.001.10*, width of anterior margin: 1.101.20*, length of lateral margin: 1.001.10*, width of posterior margin: 1.90*.
</paragraph>
</subSubSection>
<subSubSection id="C3516554F447FFFFFFC9FA46CEACFA18" pageId="31" pageNumber="430" type="etymology">
<paragraph id="8BF436DFF447FFFFFFC9FA46CEACFA18" blockId="31.[113,1089,220,1587]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9FA46CDC4FA38" bold="true" box="[113,242,1468,1491]" pageId="31" pageNumber="430">Etymology.</emphasis>
The species name is the Latin adjective
<taxonomicName id="4C4B4D5CF447FFFFFD1FFA46CFC5FA38" authorityName="Wolski" authorityYear="2017" box="[679,755,1468,1491]" class="Insecta" family="Miridae" genus="Cylapus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="430" phylum="Arthropoda" rank="species" species="luridus">
<emphasis id="B93FEACDF447FFFFFD1FFA46CFC5FA38" box="[679,755,1468,1491]" italics="true" pageId="31" pageNumber="430">luridus</emphasis>
</taxonomicName>
(-
<emphasis id="B93FEACDF447FFFFFCB4FA46CE2FFA38" box="[780,793,1468,1491]" italics="true" pageId="31" pageNumber="430">a</emphasis>
, -
<emphasis id="B93FEACDF447FFFFFC88FA46CE67FA38" box="[816,849,1468,1491]" italics="true" pageId="31" pageNumber="430">um</emphasis>
) meaning pale yellow and is used to denote the clavus with thin, yellow stripe along outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C3516554F447FFFFFFC9FA06CC72F9F8" box="[113,324,1532,1555]" pageId="31" pageNumber="430" type="biology_ecology">
<paragraph id="8BF436DFF447FFFFFFC9FA06CC72F9F8" blockId="31.[113,1089,220,1587]" box="[113,324,1532,1555]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9FA06CDFBF9F8" bold="true" box="[113,205,1532,1555]" pageId="31" pageNumber="430">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3516554F447FFFFFFC9F9E6CF62F9D8" box="[113,596,1564,1587]" pageId="31" pageNumber="430" type="distribution">
<paragraph id="8BF436DFF447FFFFFFC9F9E6CF62F9D8" blockId="31.[113,1089,220,1587]" box="[113,596,1564,1587]" pageId="31" pageNumber="430">
<emphasis id="B93FEACDF447FFFFFFC9F9E6CC32F9D8" bold="true" box="[113,260,1564,1587]" pageId="31" pageNumber="430">Distribution.</emphasis>
<collectingCountry id="F35C764FF447FFFFFEB2F9E6CC7DF9D8" box="[266,331,1564,1587]" name="Brazil" pageId="31" pageNumber="430">Brazil</collectingCountry>
(
<collectingRegion id="498FF83DF447FFFFFEE1F9E6CCF2F9D8" box="[345,452,1564,1587]" country="Brazil" name="Rondonia" pageId="31" pageNumber="430">Rondônia</collectingRegion>
) (this paper).
</paragraph>
</subSubSection>
</treatment>
</document>