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<document ID-DOI="http://doi.org/10.5281/zenodo.4386548" ID-GBIF-Dataset="92d2405f-68eb-4b38-ae95-34f307af641e" ID-GBIF-Taxon="160681416" ID-Zenodo-Dep="4386548" approvedBy="valdenar" checkinTime="1575905809981" checkinUser="valdenar" docAuthor="M., Osvaldo Villarreal, Sánchez, Nestor &amp; Ascenção, Antonio De" docDate="2019" docId="03E287C97876187F9D2141D7A9ABF81B" docLanguage="en" docName="IheringiaSérieZoologia.109.e2019008.pdf.pdf" docOrigin="Iheringia, Sér. Zool. 109" docStyle="DocumentStyle{}" docStyleName="iheringia.2019.journal_article" docTitle="Eutimesius punctatus M. &amp; Sánchez &amp; Ascenção 2019, comb. nov." docType="treatment" docVersion="11" lastPageId="3" lastPageNumber="3" masterDocId="FFDBFFB17877187C9E734355AD3EFFB9" masterDocTitle="New generic assignment to the harvestman Metaphareus punctatus (Opiliones: Stygnidae) and observations about it reproductive behavior" masterLastPageNumber="6" masterPageNumber="1" pageId="1" pageNumber="2" updateTime="1643456739563" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New generic assignment to the harvestman Metaphareus punctatus (Opiliones: Stygnidae) and observations about it reproductive behavior</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>M., Osvaldo Villarreal</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sánchez, Nestor</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ascenção, Antonio De</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Iheringia, Sér. Zool.</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-02-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>109</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>6</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/1678-4766e2019008</mods:identifier>
<mods:identifier type="GBIF-Dataset">92d2405f-68eb-4b38-ae95-34f307af641e</mods:identifier>
<mods:identifier type="Zenodo-Dep">3569290</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4386548" ID-GBIF-Taxon="160681416" ID-Zenodo-Dep="4386548" LSID="urn:lsid:plazi:treatment:03E287C97876187F9D2141D7A9ABF81B" httpUri="http://treatment.plazi.org/id/03E287C97876187F9D2141D7A9ABF81B" lastPageId="3" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection box="[850,1399,642,667]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph blockId="1.[850,1399,642,702]" box="[850,1399,642,667]" pageId="1" pageNumber="2">
<heading bold="true" box="[850,1399,642,667]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="0">
<emphasis bold="true" box="[850,1399,642,667]" pageId="1" pageNumber="2">
<taxonomicName authority="(Roewer, 1913)" authorityName="M. &amp; Sánchez &amp; Ascenção" authorityYear="2019" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[850,1265,642,666]" class="Arachnida" family="Stygnidae" genus="Eutimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="punctatus" status="comb. nov.">
<emphasis bold="true" box="[850,1087,642,666]" italics="true" pageId="1" pageNumber="2">Eutimesius punctatus</emphasis>
(
<bibRefCitation author="ROEWER, C. F." box="[1101,1257,642,666]" journalOrPublisher="Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten" pageId="1" pageNumber="2" pagination="1 - 256" part="79" refId="ref4399" refString="ROEWER, C. F. 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten 79 (4): 1 - 256." title="Die Familie der Gonyleptiden der Opiliones-Laniatores" type="journal article" year="1913">Roewer, 1913</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel box="[1277,1399,643,667]" pageId="1" pageNumber="2" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1012,1237,678,702]" pageId="1" pageNumber="2" type="description">
<paragraph blockId="1.[850,1399,642,702]" box="[1012,1237,678,702]" pageId="1" pageNumber="2">
<heading box="[1012,1237,678,702]" centered="true" fontSize="10" level="3" pageId="1" pageNumber="2" reason="1">
(
<figureCitation box="[1019,1115,678,702]" captionStart-0="Figs 1-6" captionStart-1="Figs 7-9" captionStartId-0="1.[151,188,1927,1946]" captionStartId-1="2.[151,188,946,965]" captionTargetBox-0="[331,1260,847,1905]" captionTargetBox-1="[212,1377,216,908]" captionTargetId-0="figure@1.[331,1260,847,1905]" captionTargetId-1="figure@2.[212,1377,187,908]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 1-6. Eutimesius punctatus (Roewer, 1913), comb. nov. (male MIZA 1749), habitus: 1, dorsal, panoramic view; 2, frontal view; 3, dorsal view, detail of scutum; 4, lateral view; Figs 5-6, leg III, tarsomerus, dorsal and prolateral views 4. Bars of scale = 1 mm." captionText-1="Figs 7-9. Penis of Eutimesius punctatus (Roewer, 1913), comb. nov., apical portion (MIZA 0065969): 7, lateral view; 8, ventral view; 9, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.3569292" figureDoi-1="http://doi.org/10.5281/zenodo.3569294" httpUri-0="https://zenodo.org/record/3569292/files/figure.png" httpUri-1="https://zenodo.org/record/3569294/files/figure.png" pageId="1" pageNumber="2">Figs 1–9</figureCitation>
,
<figureCitation box="[1127,1190,678,702]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="1" pageNumber="2">11-14</figureCitation>
,
<figureCitation box="[1201,1228,678,702]" captionStart="Fig" captionStartId="4.[151,182,953,972]" captionTargetBox="[168,1419,169,906]" captionTargetId="figure@4.[148,1436,151,926]" captionTargetPageId="4" captionText="Fig. 18. Geographical distribution of Eutimesius punctatus (Roewer, 1913), comb. nov. and E. albicinctus (Roewer, 1915)." figureDoi="http://doi.org/10.5281/zenodo.3569298" httpUri="https://zenodo.org/record/3569298/files/figure.png" pageId="1" pageNumber="2">18</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="reference_group">
<paragraph blockId="1.[813,1437,730,808]" pageId="1" pageNumber="2">
<taxonomicName authority="ROEWER, 1913: 432" authorityName="Roewer" authorityYear="1913" box="[813,1189,730,749]" class="Arachnida" family="Stygnidae" genus="Metaphareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="1" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis box="[813,1014,730,749]" italics="true" pageId="1" pageNumber="2">Metaphareus punctatus</emphasis>
<bibRefCitation author="ROEWER, C. F." box="[1020,1189,730,749]" journalOrPublisher="Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten" pageId="1" pageNumber="2" pagination="1 - 256" part="79" refId="ref4399" refString="ROEWER, C. F. 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten 79 (4): 1 - 256." title="Die Familie der Gonyleptiden der Opiliones-Laniatores" type="journal article" year="1913">ROEWER, 1913:432</bibRefCitation>
</taxonomicName>
, fig. 170; 1923:573 fig. 719;
<bibRefCitation author="CAPORIACCO, L. DI" box="[850,1049,759,779]" journalOrPublisher="Acta Biologica Venezuelica" pageId="1" pageNumber="2" pagination="1 - 46" part="1" refId="ref3843" refString="CAPORIACCO, L. DI 1951. Studi sugli Aracnidi del Venezuela racolti dalla sezione di Biologia (Universita Centrale del Venezuela). I parte: Scorpiones, Solifuga, Opiliones e Chernetes. Acta Biologica Venezuelica 1 (1): 1 - 46." title="Studi sugli Aracnidi del Venezuela racolti dalla sezione di Biologia (Universita Centrale del Venezuela). I parte: Scorpiones, Solifuga, Opiliones e Chernetes" type="journal article" year="1951">CAPORIACCO, 1951:42</bibRefCitation>
;
<bibRefCitation author="PINTO-DA-ROCHA, R." box="[1059,1310,759,779]" journalOrPublisher="Arquivos de Zoologia" pageId="1" pageNumber="2" pagination="163 - 342" part="33" refId="ref4366" refString="PINTO-DA-ROCHA, R. 1997. Systematic review of the neotropical family Stygnidae (Opiliones: Laniatores, Gonyleptoidea). Arquivos de Zoologia 33 (4): 163 - 342." title="Systematic review of the neotropical family Stygnidae (Opiliones: Laniatores, Gonyleptoidea)" type="journal article" year="1997">PINTO-DA-ROCHA, 1997:297</bibRefCitation>
, figs 427-431, 604;
<bibRefCitation accessDate="2018-12-27" author="KURY, A. B." box="[894,1033,789,808]" journalOrPublisher="Rio de Janeiro, Museu Nacional / UFRJ website" pageId="1" pageNumber="2" publicationUrl="http://www.museunacional.ufrj.br/mndi/Aracnologia/pdfliteratura/pdfs%20opiliones.htm" refId="ref4097" refString="KURY, A. B. 2003, onwards. OmniPaper Project - The ARACNOLAB internet resource on Opilionological Taxonomic Literature. Rio de Janeiro, Museu Nacional / UFRJ website. Avaliable at http: // www. museunacional. ufrj. br / mndi / Aracnologia / pdfliteratura / pdfs % 20 opiliones. htm. Accessed on 27 December 2018." title="onwards. OmniPaper Project - The ARACNOLAB internet resource on Opilionological Taxonomic Literature" type="url" year="2003">KURY, 2003:229</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3569292" ID-Zenodo-Dep="3569292" httpUri="https://zenodo.org/record/3569292/files/figure.png" pageId="1" pageNumber="2" startId="1.[151,188,1927,1946]" targetBox="[331,1260,847,1905]" targetPageId="1">
<paragraph blockId="1.[151,1436,1927,1972]" pageId="1" pageNumber="2">
Figs 1-6.
<taxonomicName authority="(Roewer, 1913)" authorityName="M. &amp; Sánchez &amp; Ascenção" authorityYear="2019" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[231,550,1927,1946]" class="Arachnida" family="Stygnidae" genus="Eutimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="punctatus" status="comb. nov.">
<emphasis box="[231,412,1927,1945]" italics="true" pageId="1" pageNumber="2">Eutimesius punctatus</emphasis>
(
<bibRefCitation author="ROEWER, C. F." box="[422,544,1927,1946]" journalOrPublisher="Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten" pageId="1" pageNumber="2" pagination="1 - 256" part="79" refId="ref4399" refString="ROEWER, C. F. 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten 79 (4): 1 - 256." title="Die Familie der Gonyleptiden der Opiliones-Laniatores" type="journal article" year="1913">Roewer, 1913</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel box="[558,653,1927,1946]" pageId="1" pageNumber="2" rank="species">comb. nov.</taxonomicNameLabel>
(male MIZA 1749), habitus: 1, dorsal, panoramic view; 2, frontal view; 3, dorsal view, detail of scutum; 4, lateral view; Figs 5-6, leg III, tarsomerus, dorsal and prolateral views 4. Bars of scale = 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3569294" ID-Zenodo-Dep="3569294" box="[151,1422,946,966]" httpUri="https://zenodo.org/record/3569294/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,188,946,965]" targetBox="[212,1377,216,908]" targetPageId="2">
<paragraph blockId="2.[151,1422,946,966]" box="[151,1422,946,966]" pageId="2" pageNumber="3">
Figs 7-9. Penis of
<taxonomicName authority="(Roewer, 1913)" authorityName="M. &amp; Sánchez &amp; Ascenção" authorityYear="2019" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[307,629,946,965]" class="Arachnida" family="Stygnidae" genus="Eutimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus" status="comb. nov.">
<emphasis box="[307,489,947,965]" italics="true" pageId="2" pageNumber="3">Eutimesius punctatus</emphasis>
(
<bibRefCitation author="ROEWER, C. F." box="[500,623,946,965]" journalOrPublisher="Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten" pageId="2" pageNumber="3" pagination="1 - 256" part="79" refId="ref4399" refString="ROEWER, C. F. 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten 79 (4): 1 - 256." title="Die Familie der Gonyleptiden der Opiliones-Laniatores" type="journal article" year="1913">Roewer, 1913</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel box="[638,735,946,966]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
, apical portion (MIZA 0065969): 7, lateral view; 8, ventral view; 9, dorsal view.
</paragraph>
</caption>
<subSubSection pageId="2" pageNumber="3" type="type_taxon">
<paragraph blockId="2.[151,776,1053,1957]" pageId="2" pageNumber="3">
<typeStatus box="[227,279,1054,1078]" pageId="2" pageNumber="3">Type</typeStatus>
data.
<materialsCitation ID-GBIF-Occurrence="2465044320" collectionCode="SMF RI, 832" country="Venezuela" location="Merida" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Merida" typeStatus="holotype">
<collectingCountry box="[338,493,1054,1078]" name="Venezuela" pageId="2" pageNumber="3">VENEZUELA</collectingCountry>
,
<emphasis bold="true" box="[504,586,1053,1077]" pageId="2" pageNumber="3">
<collectingRegion box="[504,586,1053,1077]" country="Venezuela" name="Merida" pageId="2" pageNumber="3">Mérida</collectingRegion>
</emphasis>
,
<typeStatus box="[598,689,1053,1077]" pageId="2" pageNumber="3">holotype</typeStatus>
<specimenCount box="[694,715,1054,1077]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
,
<typeStatus box="[730,775,1054,1078]" pageId="2" pageNumber="3">type</typeStatus>
<collectionCode box="[151,294,1086,1110]" pageId="2" pageNumber="3">SMF RI, 832</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="materials_examined">
<paragraph blockId="2.[151,776,1053,1957]" pageId="2" pageNumber="3">
Studied material.
<materialsCitation ID-GBIF-Occurrence="2465044326" collectingDate="2006-08" collectingDateMax="2017-02-03" collectingDateMin="2006-08" collectionCode="MIZA" collectorName="RodrIguez C. &amp; Villarreal, M. O. &amp; GarcIa, A. &amp; SAnchez" country="Venezuela" elevation="2" latitude="8.583611" location="Sector Piedra Blanca" longLatPrecision="1" longitude="-71.32611" municipality="Campo ElIas" pageId="2" pageNumber="3" specimenCode="MIZA 1749" specimenCount="2118" specimenCount-female="1" specimenCount-male="2117" stateProvince="Merida">
<collectingCountry box="[421,581,1119,1143]" name="Venezuela" pageId="2" pageNumber="3">VENEZUELA</collectingCountry>
,
<emphasis bold="true" box="[596,681,1118,1142]" pageId="2" pageNumber="3">
<collectingRegion box="[596,681,1118,1142]" country="Venezuela" name="Merida" pageId="2" pageNumber="3">Mérida</collectingRegion>
</emphasis>
:
<collectingMunicipality pageId="2" pageNumber="3">Campo ElÍas</collectingMunicipality>
,
<specimenCount box="[216,238,1152,1175]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
,
<specimenCount box="[248,268,1152,1175]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
(
<specimenCode box="[280,411,1151,1175]" pageId="2" pageNumber="3">MIZA 1749</specimenCode>
),
<location LSID="urn:lsid:plazi:treatment:03E287C97876187F9D2141D7A9ABF81B:8E9460047875187E9FDF472AAFB4FB2E" box="[428,650,1151,1175]" country="Venezuela" latitude="8.583611" longLatPrecision="1" longitude="-71.32611" municipality="Campo ElIas" name="Sector Piedra Blanca" pageId="2" pageNumber="3" stateProvince="Merida">Sector Piedra Blanca</location>
, [
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],
<quantity box="[284,368,1184,1208]" metricMagnitude="3" metricUnit="m" metricValue="2.114" pageId="2" pageNumber="3" unit="m" value="2114.0">
<specimenCount box="[284,368,1184,1208]" pageId="2" pageNumber="3" type="male">2114 m</specimenCount>
</quantity>
,
<date box="[381,485,1184,1208]" pageId="2" pageNumber="3" value="2006-08">
<collectingDate box="[381,485,1184,1208]" pageId="2" pageNumber="3" value="2006-08">Viii.2006</collectingDate>
</date>
,
<collectorName box="[497,642,1182,1208]" pageId="2" pageNumber="3">RodrÍguez C.</collectorName>
leg;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2465044327" collectingDate="2006-08" collectingDateMax="2017-02-03" collectingDateMin="2006-08" collectionCode="MIZA" collectorName="RodrIguez C. &amp; Villarreal, M. O. &amp; GarcIa, A. &amp; SAnchez" country="Venezuela" elevation="2" latitude="8.583611" location="Sector Piedra Blanca" longLatPrecision="1" longitude="-71.32611" municipality="Campo ElIas" pageId="2" pageNumber="3" specimenCode="MIZA 1749" specimenCount="2118" specimenCount-female="1" specimenCount-male="2117" stateProvince="Merida">
AndrÉs
<location LSID="urn:lsid:plazi:treatment:03E287C97876187F9D2141D7A9ABF81B:8E9460047875187E9EE44795ADE7FB61" box="[151,217,1216,1240]" country="Venezuela" latitude="8.583611" longLatPrecision="1" longitude="-71.32611" municipality="Campo ElIas" name="Bello" pageId="2" pageNumber="3" stateProvince="Merida">Bello</location>
,
<specimenCount box="[233,269,1216,1240]" pageId="2" pageNumber="3" type="male">2♂</specimenCount>
(
<collectionCode box="[287,361,1216,1240]" collectionName="Venezuela, Maracay, Museuo del Instituto de Zoologia Agricola" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="2" pageNumber="3" type="Museum">MIZA</collectionCode>
s/n)
<location LSID="urn:lsid:plazi:treatment:03E287C97876187F9D2141D7A9ABF81B:8E9460047875187E9FD84795AF00FB60" box="[427,574,1215,1241]" country="Venezuela" latitude="8.583611" longLatPrecision="1" longitude="-71.32611" municipality="Campo ElIas" name="road MErida" pageId="2" pageNumber="3" stateProvince="Merida">road MÉrida</location>
-
<location LSID="urn:lsid:plazi:treatment:03E287C97876187F9D2141D7A9ABF81B:8E9460047875187E9C2E4794AFE0FB61" box="[605,734,1216,1240]" country="Venezuela" latitude="8.583611" longLatPrecision="1" longitude="-71.32611" municipality="Campo ElIas" name="La Azulita" pageId="2" pageNumber="3" stateProvince="Merida">La Azulita</location>
,
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,
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,
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,
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,
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<elevation metricMagnitude="0" metricUnit="m" metricValue="2.255" pageId="2" pageNumber="3" unit="m" value="2.255">2.255 m</elevation>
</quantity>
,
<date box="[187,298,1281,1305]" pageId="2" pageNumber="3" value="2017-02-03">
<collectingDate box="[187,298,1281,1305]" pageId="2" pageNumber="3" value="2017-02-03">03.ii.2017</collectingDate>
</date>
,
<collectorName box="[310,485,1281,1306]" pageId="2" pageNumber="3">Villarreal, M.O.</collectorName>
;
<collectorName box="[497,737,1280,1306]" pageId="2" pageNumber="3">GarcÍa, A. &amp; SÁnchez</collectorName>
, N. leg;
</materialsCitation>
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<collectingMunicipality box="[198,315,1314,1338]" pageId="2" pageNumber="3">Libertador</collectingMunicipality>
:
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,
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, (
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),
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,
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<collectingDate box="[519,639,1347,1371]" pageId="2" pageNumber="3" value="2010-04-11">11.IV.2010</collectingDate>
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,
<collectorName box="[651,774,1345,1371]" pageId="2" pageNumber="3">N. SÁnchez</collectorName>
&amp;
<collectorName box="[180,302,1379,1403]" pageId="2" pageNumber="3">Mendoza I.</collectorName>
leg;
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(
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<collectingDate box="[519,639,1412,1436]" pageId="2" pageNumber="3" value="2010-04-11">11.IV.2010</collectingDate>
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,
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leg;
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,
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(
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,
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,
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<collectingDate box="[390,512,1477,1501]" pageId="2" pageNumber="3" value="2010-04-17">17.IV.2010</collectingDate>
</date>
,
<collectorName box="[525,649,1475,1501]" pageId="2" pageNumber="3">SÁnchez N.</collectorName>
leg;
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<materialsCitation ID-GBIF-Occurrence="2465044322" collectingDate="2010-05-25" collectionCode="CAULA-Op-0051" collectorName="SAnchez N. &amp; De AscenCAo A." elevation="2160" location="Monte Zerpa" pageId="2" pageNumber="3" specimenCount="8" specimenCount-female="1" specimenCount-male="7">
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),
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,
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<collectingDate box="[654,771,1509,1533]" pageId="2" pageNumber="3" value="2010-05-25">25.V.2010</collectingDate>
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,
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leg;
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<materialsCitation ID-GBIF-Occurrence="2465044323" collectingDate="2011-09-24" collectionCode="CAULA- Op-0243" collectorName="SAnchez N. &amp; Mendoza I." elevation="2170" latitude="8.639722" location="Monte Zerpa" longLatPrecision="1" longitude="-71.16744" pageId="2" pageNumber="3" specimenCount="7" specimenCount-female="5" specimenCount-male="2">
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,
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(
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),
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[
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,
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<collectingDate box="[188,331,1608,1632]" pageId="2" pageNumber="3" value="2011-09-24">24. IX. 2011</collectingDate>
</date>
,
<collectorName box="[343,470,1606,1632]" pageId="2" pageNumber="3">SÁnchez N.</collectorName>
&amp;
<collectorName box="[507,633,1607,1631]" pageId="2" pageNumber="3">Mendoza I.</collectorName>
leg;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2465044325" collectingDate="2011-10-08" collectionCode="CAULA-Op-0302" collectorName="SAnchez N. &amp; Mendoza I." latitude="8.641472" location="Monte Zerpa" longLatPrecision="1" longitude="-71.162674" pageId="2" pageNumber="3" specimenCount="2145" specimenCount-female="3" specimenCount-male="2142">
<specimenCount box="[692,728,1608,1632]" pageId="2" pageNumber="3" type="male">2♂</specimenCount>
,
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(
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),
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,
<location LSID="urn:lsid:plazi:treatment:03E287C97876187F9D2141D7A9ABF81B:8E9460047875187E9C53453DAC04F918" latitude="8.641472" longLatPrecision="1" longitude="-71.162674" name="Quebrada La Honda, RIo Albarregas" pageId="2" pageNumber="3">Quebrada La Honda, RÍo Albarregas</location>
, [
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<date box="[676,772,1673,1697]" pageId="2" pageNumber="3" value="2011-10-08">
<collectingDate box="[676,772,1673,1697]" pageId="2" pageNumber="3" value="2011-10-08">8.x.2011</collectingDate>
</date>
.
<collectorName box="[151,273,1703,1729]" pageId="2" pageNumber="3">SÁnchez N.</collectorName>
&amp;
<collectorName box="[307,428,1705,1729]" pageId="2" pageNumber="3">Mendoza I.</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="diagnosis">
<paragraph blockId="2.[151,776,1053,1957]" pageId="2" pageNumber="3">
Diagnosis. This species can be distinguished from its other all congeneric species by the black spots on the base of mesotergal tubercles contrasting with the yellow greenish background (
<figureCitation box="[292,383,1835,1860]" captionStart="Figs 1-6" captionStartId="1.[151,188,1927,1946]" captionTargetBox="[331,1260,847,1905]" captionTargetId="figure@1.[331,1260,847,1905]" captionTargetPageId="1" captionText="Figs 1-6. Eutimesius punctatus (Roewer, 1913), comb. nov. (male MIZA 1749), habitus: 1, dorsal, panoramic view; 2, frontal view; 3, dorsal view, detail of scutum; 4, lateral view; Figs 5-6, leg III, tarsomerus, dorsal and prolateral views 4. Bars of scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3569292" httpUri="https://zenodo.org/record/3569292/files/figure.png" pageId="2" pageNumber="3">Figs 1-4</figureCitation>
,
<figureCitation box="[396,463,1836,1860]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">11, 14</figureCitation>
) and pattern of the dry-mark (
<figureCitation box="[159,228,1868,1892]" captionStart="Figs 1-6" captionStartId="1.[151,188,1927,1946]" captionTargetBox="[331,1260,847,1905]" captionTargetId="figure@1.[331,1260,847,1905]" captionTargetPageId="1" captionText="Figs 1-6. Eutimesius punctatus (Roewer, 1913), comb. nov. (male MIZA 1749), habitus: 1, dorsal, panoramic view; 2, frontal view; 3, dorsal view, detail of scutum; 4, lateral view; Figs 5-6, leg III, tarsomerus, dorsal and prolateral views 4. Bars of scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3569292" httpUri="https://zenodo.org/record/3569292/files/figure.png" pageId="2" pageNumber="3">Figs 3</figureCitation>
,
<figureCitation box="[239,308,1868,1892]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">11, 14</figureCitation>
).
</paragraph>
<paragraph blockId="2.[151,776,1053,1957]" lastBlockId="2.[813,1437,1053,1957]" pageId="2" pageNumber="3">
Genital description (males) (
<figureCitation box="[560,662,1901,1925]" captionStart="Figs 7-9" captionStartId="2.[151,188,946,965]" captionTargetBox="[212,1377,216,908]" captionTargetId="figure@2.[212,1377,187,908]" captionTargetPageId="2" captionText="Figs 7-9. Penis of Eutimesius punctatus (Roewer, 1913), comb. nov., apical portion (MIZA 0065969): 7, lateral view; 8, ventral view; 9, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3569294" httpUri="https://zenodo.org/record/3569294/files/figure.png" pageId="2" pageNumber="3">Figs 7-9</figureCitation>
). General pattern typical for
<taxonomicName authorityName="Roewer" authorityYear="1913" box="[351,530,1933,1957]" class="Arachnida" family="Stygnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Heterostygninae">Heterostygninae</taxonomicName>
: with the hammer well differentiated and carrying the MS A1-A2 and MS B, all aligned forming a ring slightly basal at the beginning of the lamina parva (LP), which is clearly differentiated from the truncus and have lateral margins convergent and curved, with apical circular and deep cleft, and a basal constriction forming a basal neck. MS C1-C2 similar in size, positioned slightly apical at neck of the LP. Only one pair of MS D, located on a monticule, dorsally on the base of LP. Stylus curved dorsally, with small dorsal process.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="biology_ecology">
<paragraph blockId="2.[813,1437,1053,1957]" pageId="2" pageNumber="3">
Natural history. This species inhabits exclusively in cloud forests located in the Andean mountain range system in
<collectingRegion box="[840,975,1410,1436]" country="Venezuela" name="Merida" pageId="2" pageNumber="3">MÉrida state</collectingRegion>
, particularly in protected areas in Sierra de La Culata National Park (
<figureCitation box="[1076,1197,1444,1468]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">Figs 14–16</figureCitation>
).
<taxonomicName authorityName="M. &amp; Sánchez &amp; Ascenção" authorityYear="2019" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[1214,1436,1445,1468]" class="Arachnida" family="Stygnidae" genus="Eutimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis box="[1214,1436,1445,1468]" italics="true" pageId="2" pageNumber="3">Eutimesius punctatus</emphasis>
</taxonomicName>
is characterized for occupying microhabitats above the undergrowth, being frequent to find it on shrubs near small water courses (
<figureCitation box="[965,1039,1542,1566]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">Fig. 17</figureCitation>
). Remarkably, in Monte Zerpa,
<collectingRegion country="Venezuela" name="Merida" pageId="2" pageNumber="3">Mérida state</collectingRegion>
, it was frequently found on a species of scaly tree fern
<taxonomicName authority="(H. Karst)" authorityName="H. Karst" baseAuthorityName="H. Karst" box="[813,1285,1607,1632]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="variety" species="caracasana" variety="meridensis">
<emphasis box="[813,1022,1607,1631]" italics="true" pageId="2" pageNumber="3">Cyathea caracasana</emphasis>
var.
<emphasis box="[1069,1179,1607,1631]" italics="true" pageId="2" pageNumber="3">meridensis</emphasis>
(H. Karst)
</taxonomicName>
(
<taxonomicName authorityName="Kaulfuss" authorityYear="1827" box="[1297,1429,1607,1631]" class="Polypodiopsida" family="Cyatheaceae" kingdom="Plantae" order="Cyatheales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="family">Cyatheaceae</taxonomicName>
) a very common species on the edges of streams, which is used by
<taxonomicName authorityName="M. &amp; Sánchez &amp; Ascenção" authorityYear="2019" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[904,1039,1673,1696]" class="Arachnida" family="Stygnidae" genus="Eutimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis box="[904,1039,1673,1696]" italics="true" pageId="2" pageNumber="3">E. punctatus</emphasis>
</taxonomicName>
to lay his eggs (
<figureCitation box="[1220,1346,1673,1697]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">Figs 13, 17</figureCitation>
). In this regard, one of the authors observed four events involving posture or care of eggs inthese plants, always the eggswere placed on the back of the leaves (at the bottom) (
<figureCitation box="[1343,1423,1770,1794]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">Fig. 13</figureCitation>
). All these cases were observed at the same locality: brook El Macanal, Monte Zerpa,
<collectingRegion box="[1070,1202,1835,1859]" country="Venezuela" name="Merida" pageId="2" pageNumber="3">Mérida state</collectingRegion>
(
<figureCitation box="[1216,1339,1835,1860]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="2" pageNumber="3">Figs 15, 17</figureCitation>
).
</paragraph>
<paragraph blockId="2.[813,1437,1053,1957]" lastBlockId="3.[813,1437,1741,1954]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
The first case only one female was observed next to the clutch (about
<specimenCount box="[997,1081,1901,1925]" pageId="2" pageNumber="3" type="egg">50 eggs</specimenCount>
), which has a semicircular shape; a second case was observed both male and female next to the clutch (eggs arranged in a similar way to the previous case), in this case after being disturbed they both left the clutch, they were dropped in a vacuum to fall into the leaf litter; a third case was observed, which only the female after the disturbance fled the clutch, in this case the eggs had a different appearance, some of which located on a side edge looked more recent; and a fourth case in which only a male and some scattered eggs were observed (about
<specimenCount box="[1300,1380,1779,1803]" pageId="3" pageNumber="4" type="egg">15 eggs</specimenCount>
in an approximate diameter of
<quantity box="[1083,1151,1817,1841]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="cm" value="10.0">10 cm</quantity>
) (
<figureCitation box="[1174,1252,1817,1841]" captionStart="Figs 10-17" captionStartId="3.[151,188,1610,1629]" captionTargetBox="[151,1436,155,1570]" captionTargetId="figure@3.[151,1436,155,1570]" captionTargetPageId="3" captionText="Figs 10-17. Fig. 10, Eutimesius albicinctus (Roewer, 1915), male at live from Chorotal, MÉrida, Venezuela; Figs 11-12, Eutimesius punctatus (Roewer, 1913), comb. nov., male at live from Chorotal, MÉrida, Venezuela; Figs 13, 14, male and egg on fern from Monte Zerpa, MÉrida; Figs 15-17, cloud forest in MÉrida state, Chorotal (15) and Monte Zerpa (16 and 17)." figureDoi="http://doi.org/10.5281/zenodo.3569296" httpUri="https://zenodo.org/record/3569296/files/figure.png" pageId="3" pageNumber="4">Fig. 13</figureCitation>
), this could have several interpretations, maybe it is the remnant of a larger clutch cared by this male, or maybe the male was preying on the eggs of an old alien clutch.
</paragraph>
</subSubSection>
</treatment>
</document>
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