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<mods:title id="B573936454FB50F1DF7613358FF50076">Resolving taxonomic issues of cicadas (Hemiptera: Cicadidae) including new combinations, new synonymies, and revised status, with updates on the diversity of the Brazilian cicada fauna and new records for four South American countries</mods:title>
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<treatment id="03E2878BFFE95928FF50FC94FC19FB9F" ID-DOI="http://doi.org/10.5281/zenodo.8166918" ID-GBIF-Taxon="212427671" ID-Zenodo-Dep="8166918" LSID="urn:lsid:plazi:treatment:03E2878BFFE95928FF50FC94FC19FB9F" httpUri="http://treatment.plazi.org/id/03E2878BFFE95928FF50FC94FC19FB9F" lastPageId="5" lastPageNumber="344" pageId="3" pageNumber="342">
<subSubSection id="C3516516FFE9592EFF50FC94FCD8F913" box="[151,812,1751,1778]" pageId="3" pageNumber="342" type="nomenclature">
<paragraph id="8BF4369DFFE9592EFF50FC94FCD8F913" blockId="3.[151,812,1751,1778]" box="[151,812,1751,1778]" pageId="3" pageNumber="342">
<heading id="D0BC81F1FFE9592EFF50FC94FCD8F913" bold="true" box="[151,812,1751,1778]" fontSize="11" level="1" pageId="3" pageNumber="342" reason="1">
<emphasis id="B93FEA8FFFE9592EFF50FC94FCD8F913" bold="true" box="[151,812,1751,1778]" pageId="3" pageNumber="342">
<taxonomicName id="4C4B4D1EFFE9592EFF50FC94FDBBF913" ID-CoL="1e24e3f4-7743-4c6f-9d27-523129d0f3d1" authority="(Distant, 1892)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1892" box="[151,591,1751,1778]" class="Insecta" family="Cicadidae" genus="Brevialavenosa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="egae" status="comb. nov.">
<emphasis id="B93FEA8FFFE9592EFF50FC94FE7BF910" bold="true" box="[151,399,1751,1778]" italics="true" pageId="3" pageNumber="342">Brevialavenosa egae</emphasis>
(
<bibRefCitation id="EFDA4B6CFFE9592EFE58FC94FDB3F913" author="Distant" box="[415,583,1751,1777]" firstAuthor="Distant" pageId="3" pageNumber="342" pagination="54 - 67" refId="ref12244" refString="Distant, W. L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant, 1892</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20C57F4FFE9592EFD91FC9BFCD8F913" box="[598,812,1751,1778]" pageId="3" pageNumber="342" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3516516FFE9592EFF50FD5FFD4DF8D6" box="[151,697,1820,1844]" pageId="3" pageNumber="342" type="reference_group">
<paragraph id="8BF4369DFFE9592EFF50FD5FFD4DF8D6" blockId="3.[151,697,1820,1844]" box="[151,697,1820,1844]" pageId="3" pageNumber="342">
<treatmentCitationGroup id="AB5B11B3FFE9592EFF50FD5FFE39F8D6" box="[151,461,1820,1844]" pageId="3" pageNumber="342">
<treatmentCitation id="0AEA108CFFE9592EFF50FD5FFE39F8D6" author="Distant, W. L." box="[151,461,1820,1844]" page="64" pageId="3" pageNumber="342" year="1892">
<taxonomicName id="4C4B4D1EFFE9592EFF50FD5FFE39F8D6" ID-CoL="433ecfea-3d6a-4ba8-a8ac-66f954f8844c" authority="Distant, 1892" authorityName="Distant" authorityPageNumber="64" authorityYear="1892" box="[151,461,1820,1844]" class="Insecta" family="Cicadidae" genus="Tibicen" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B93FEA8FFFE9592EFF50FD5FFEECF8D6" box="[151,280,1820,1844]" italics="true" pageId="3" pageNumber="342">Tibicen egae</emphasis>
<bibRefCitation id="EFDA4B6CFFE9592EFED9FD5FFE39F8D6" author="Distant" box="[286,461,1820,1844]" firstAuthor="Distant" page="64" pageId="3" pageNumber="342" pagination="54 - 67" refId="ref12244" refString="Distant, W. L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant, 1892: 64</bibRefCitation>
</taxonomicName>
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(Ega, on the Amazon).
</paragraph>
</subSubSection>
<subSubSection id="C3516516FFE95928FF50FD21FEB1FBBB" lastPageId="5" lastPageNumber="344" pageId="3" pageNumber="342" type="discussion">
<paragraph id="8BF4369DFFE95928FF50FD21FF1AFEA3" blockId="3.[151,1436,1890,2025]" lastBlockId="5.[151,1437,151,1149]" lastPageId="5" lastPageNumber="344" pageId="3" pageNumber="342">
<emphasis id="B93FEA8FFFE9592EFF50FD21FEFAF89E" bold="true" box="[151,270,1890,1916]" pageId="3" pageNumber="342">REMARKS.</emphasis>
<bibRefCitation id="EFDA4B6CFFE9592EFED2FD20FE19F89E" author="Nunes" box="[277,493,1890,1917]" etAl="et al." firstAuthor="Nunes" pageId="3" pageNumber="342" refId="ref14077" refString="Nunes, V. L., Ruschel, T. P., Maccagnan, D. H. B., Simies, P. C. &amp; Acosta, R. C. (2023) Cigarras do Brasil, Guia Fotographica. 1 a edicao. ebook. 46 pp. [https: // zenodo. org / record / 7712483]" type="book" year="2023">
Nunes
<emphasis id="B93FEA8FFFE9592EFEA3FD20FE61F89E" box="[356,405,1890,1916]" italics="true" pageId="3" pageNumber="342">et al</emphasis>
. (2023)
</bibRefCitation>
reported two species,
<taxonomicName id="4C4B4D1EFFE9592EFD2FFD21FBDAF89E" authority="(Distant, 1892)" baseAuthorityName="Distant" baseAuthorityYear="1892" box="[744,1070,1890,1916]" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B93FEA8FFFE9592EFD2FFD21FC89F89E" box="[744,893,1890,1916]" italics="true" pageId="3" pageNumber="342">Abroma egae</emphasis>
(
<bibRefCitation id="EFDA4B6CFFE9592EFC4BFD21FBD1F89E" author="Distant" box="[908,1061,1890,1916]" firstAuthor="Distant" pageId="3" pageNumber="342" pagination="54 - 67" refId="ref12244" refString="Distant, W. L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant, 1892</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4B4D1EFFE9592EFBA3FD21FF2FF843" authority="(Walker, 1858)" baseAuthorityName="Walker" baseAuthorityYear="1858" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="temperata">
<emphasis id="B93FEA8FFFE9592EFBA3FD21FAC2F89E" box="[1124,1334,1890,1916]" italics="true" pageId="3" pageNumber="342">Abroma temperata</emphasis>
(
<bibRefCitation id="EFDA4B6CFFE9592EFA83FD21FF27F843" author="Walker, F." bookContentInfo="117 pp." journalOrPublisher="John van Voorts, London" pageId="3" pageNumber="342" refId="ref16976" refString="Walker, F. (1858 a) Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders, Esq. Homoptera. John van Voorts, London, 117 pp. https: // doi. org / 10.5962 / bhl. title. 5112" title="Insecta Saundersiana: or characters of undescribed insects in the collection of William Wilson Saunders, Esq. Homoptera" type="book" year="1858">Walker, 1858</bibRefCitation>
)
</taxonomicName>
, from the genus
<taxonomicName id="4C4B4D1EFFE9592EFE58FDC5FD81F842" authority="Stal, 1866" authorityName="Stal" authorityYear="1866" box="[415,629,1926,1953]" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEA8FFFE9592EFE58FDC5FE0CF842" box="[415,504,1926,1952]" italics="true" pageId="3" pageNumber="342">Abroma</emphasis>
Stål, 1866
</taxonomicName>
and the tribe
<taxonomicName id="4C4B4D1EFFE9592EFCDFFDC5FBA1F842" authority="Boulard, 1976" authorityName="Boulard" authorityYear="1976" box="[792,1109,1926,1952]" pageId="3" pageNumber="342" rank="tribe" tribe="Lamotialnini">
Lamotialnini
<bibRefCitation id="EFDA4B6CFFE9592EFC75FDC5FBA1F842" author="Boulard" box="[946,1109,1926,1952]" firstAuthor="Boulard" pageId="3" pageNumber="342" pagination="629 - 636" refId="ref11792" refString="Boulard, M. (1976) Sur une deuxieme cigale africaine depourvue d'appareil sonore (Homoptera). Bulletin de l'Institut Fondamental d'Afrique Noire, Series A, 37, 629 - 636." type="journal article" year="1976">
Boulard,
<quantity id="4CB39B78FFE9592EFBDAFDC5FBA1F842" box="[1053,1109,1926,1952]" metricMagnitude="1" metricUnit="m" metricValue="5.0190399999999995" pageId="3" pageNumber="342" unit="in" value="1976.0">1976</quantity>
</bibRefCitation>
</taxonomicName>
in
<collectingCountry id="F35C760DFFE9592EFBBAFDC5FB31F842" box="[1149,1221,1926,1952]" name="Brazil" pageId="3" pageNumber="342">Brazil</collectingCountry>
. Neither the genus nor the tribe is found in South America (
<bibRefCitation id="EFDA4B6CFFE9592EFDA4FDE9FCBBF826" author="Marshall" box="[611,847,1962,1989]" etAl="et al." firstAuthor="Marshall" pageId="3" pageNumber="342" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B93FEA8FFFE9592EFD16FDE8FCF7F826" box="[721,771,1962,1988]" italics="true" pageId="3" pageNumber="342">et al</emphasis>
. 2018
</bibRefCitation>
). Both species were transferred without comment to
<taxonomicName id="4C4B4D1EFFE9592EFF50FD8DFF04F80A" authorityName="Stal" authorityYear="1866" box="[151,240,1998,2024]" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEA8FFFE9592EFF50FD8DFF04F80A" box="[151,240,1998,2024]" italics="true" pageId="3" pageNumber="342">Abroma</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFDA4B6CFFE9592EFED3FD8DFE4EF80A" author="Distant" box="[276,442,1998,2024]" firstAuthor="Distant" pageId="3" pageNumber="342" refId="ref12567" refString="Distant, W. L. (1906) A synonymic catalogue of Homoptera. Part I. Cicadidae. Trustees of the British Museum, London, 207 pp. https: // doi. org / 10.5962 / bhl. title. 8554" type="book" year="1906">Distant (1906)</bibRefCitation>
.
<taxonomicName id="4C4B4D1EFFE9592EFE02FD8DFDEAF80A" authorityName="Stal" authorityYear="1866" box="[453,542,1998,2024]" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEA8FFFE9592EFE02FD8DFDEAF80A" box="[453,542,1998,2024]" italics="true" pageId="3" pageNumber="342">Abroma</emphasis>
</taxonomicName>
was classified in the Taphurini
<bibRefCitation id="EFDA4B6CFFE9592EFC46FD8DFBD0F80B" author="Distant" box="[897,1060,1998,2025]" firstAuthor="Distant" pageId="3" pageNumber="342" pagination="22 - 35" refId="ref12327" refString="Distant, W. L. (1905 a) Rhynchotal notes XXXIII. Annals and Magazine of Natural History, Series 7, 16, 22 - 35. https: // doi. org / 10.1080 / 03745480509443650" type="journal article" year="1905" yearSuffix="a">Distant, 1905a</bibRefCitation>
in
<bibRefCitation id="EFDA4B6CFFE9592EFB8FFD8DFB09F80A" author="Distant's" box="[1096,1277,1998,2024]" firstAuthor="Distant's" pageId="3" pageNumber="342" refId="ref12567" refString="Distant, W. L. (1906) A synonymic catalogue of Homoptera. Part I. Cicadidae. Trustees of the British Museum, London, 207 pp. https: // doi. org / 10.5962 / bhl. title. 8554" type="book" year="1906">Distants (1906)</bibRefCitation>
catalogue but the remaining members of the genus are found in Africa,
<collectingCountry id="F35C760DFFEF5928FCCBFAD4FC61FF53" box="[780,917,151,177]" name="Madagascar" pageId="5" pageNumber="344">Madagascar</collectingCountry>
, and south, southeast and insular Asia (
<bibRefCitation id="EFDA4B6CFFEF5928FA84FAD4FF2BFF37" author="Metcalf" firstAuthor="Metcalf" pageId="5" pageNumber="344" pagination="1 - 492" refId="ref13979" refString="Metcalf, Z. P. (1963 c) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 2. Tibicinidae. North Carolina State College Contribution, 1564, i - vi + 1 - 492." type="journal article" year="1963" yearSuffix="c">Metcalf 1963c</bibRefCitation>
;
<bibRefCitation id="EFDA4B6CFFEF5928FF2DFAF8FDC5FF37" author="Duffels &amp; van der Laan" box="[234,561,186,213]" firstAuthor="Duffels" pageId="5" pageNumber="344" refId="ref12737" refString="Duffels, J. P. &amp; van der Laan, P. A. (1985) Catalogue of the Cicadoidea (Homoptera, Auchenorhyncha) 1956 - 1980. Series Entomologica 34. Dr. W. Junk Publishers, Dordrect, 414 pp." type="book" year="1985">Duffels &amp; van der Laan 1985</bibRefCitation>
;
<bibRefCitation id="EFDA4B6CFFEF5928FDFBFAF8FD29FF37" author="Sanborn, A. F." bookContentInfo="1001 pp." box="[572,733,187,213]" journalOrPublisher="Elsevier / Academic Press, San Diego" pageId="5" pageNumber="344" refId="ref14777" refString="Sanborn, A. F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier / Academic Press, San Diego, 1001 pp. https: // doi. org / 10.1016 / b 978 - 0 - 12 - 416647 - 9.00001 - 2" title="Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet" type="book" year="2013">Sanborn 2013</bibRefCitation>
). Historically, the Taphurini was a tribe where taxa were placed when it was unclear where the taxon actually belonged and had representatives from most of the world (
<bibRefCitation id="EFDA4B6CFFEF5928FAF0FA9CFEE6FEFF" author="Marshall" etAl="et al." firstAuthor="Marshall" pageId="5" pageNumber="344" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B93FEA8FFFEF5928FF50FB40FF3CFEFF" box="[151,200,259,285]" italics="true" pageId="5" pageNumber="344">et al</emphasis>
. 2018
</bibRefCitation>
). However, the tribe has been redefined and currently includes only Neotropical genera (
<bibRefCitation id="EFDA4B6CFFEF5928FB2DFB40FA6CFEFF" author="Sanborn" box="[1258,1432,259,285]" firstAuthor="Sanborn" pageId="5" pageNumber="344" pagination="1 - 79" refId="ref15711" refString="Sanborn, A. F. (2021 a) The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species. Zootaxa, 4937 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 4937.1.1" type="journal article" year="2021" yearSuffix="a">Sanborn 2021a</bibRefCitation>
;
<bibRefCitation id="EFDA4B6CFFEF5928FF50FB64FF16FEA3" author="Sanborn" box="[151,226,295,321]" firstAuthor="Sanborn" pageId="5" pageNumber="344" pagination="190 - 198" refId="ref15802" refString="Sanborn, A. F. (2021 b) A new genus and species of Neotropical Taphurini Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae) from Brazil with a note on the taxonomic position of Prosotettix Jacobi, 1907. Proceedings of the Entomological Society of Washington, 123, 190 - 198. https: // doi. org / 10.4289 / 0013 - 8797.123.1.190" type="journal article" year="2021" yearSuffix="b">2021b</bibRefCitation>
).
</paragraph>
<caption id="DF346615FFEE5929FF5FFDD6FBE7F817" ID-DOI="http://doi.org/10.5281/zenodo.8166902" ID-Zenodo-Dep="8166902" httpUri="https://zenodo.org/record/8166902/files/figure.png" pageId="4" pageNumber="343" startId="4.[152,255,1941,1965]" targetBox="[203,1384,181,1917]" targetPageId="4" targetType="figure">
<paragraph id="8BF4369DFFEE5929FF5FFDD6FBE7F817" blockId="4.[151,1436,1941,2038]" pageId="4" pageNumber="343">
<emphasis id="B93FEA8FFFEE5929FF5FFDD6FEEEF84F" bold="true" box="[152,282,1941,1965]" pageId="4" pageNumber="343">FIGURE 2.</emphasis>
<taxonomicName id="4C4B4D1EFFEE5929FEE5FDD6FDC3F84F" authority="Distant, 1892" authorityName="Distant" authorityYear="1892" box="[290,567,1941,1965]" class="Insecta" family="Cicadidae" genus="Tibicen" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="343" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B93FEA8FFFEE5929FEE5FDD6FE51F84F" box="[290,421,1941,1965]" italics="true" pageId="4" pageNumber="343">Tibicen egae</emphasis>
<bibRefCitation id="EFDA4B6CFFEE5929FE6AFDD6FDC3F84F" author="Distant" box="[429,567,1941,1965]" firstAuthor="Distant" pageId="4" pageNumber="343" pagination="54 - 67" refId="ref12244" refString="Distant, W. L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant, 1892</bibRefCitation>
</taxonomicName>
type specimen (NHMUK010392112) dorsal (top) and ventral (middle) habitus. The inflated aedeagus with membrane and recurved terminal spines are characteristic of
<taxonomicName id="4C4B4D1EFFEE5929FBC8FDFAFB58F833" authorityName="Sanborn" authorityYear="2021" box="[1039,1196,1977,2001]" class="Insecta" family="Cicadidae" genus="Brevialavenosa" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEA8FFFEE5929FBC8FDFAFB58F833" box="[1039,1196,1977,2001]" italics="true" pageId="4" pageNumber="343">Brevialavenosa</emphasis>
</taxonomicName>
. ©The Trustees of the Natural History Museum, London. Photos courtesy of Fernley Symons and Mick Webb.
</paragraph>
</caption>
<paragraph id="8BF4369DFFEF5928FF00FB08FEEDFD67" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="344">
<taxonomicName id="4C4B4D1EFFEF5928FF00FB08FE92FE86" baseAuthorityName="Distant" baseAuthorityYear="1892" box="[199,358,331,357]" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B93FEA8FFFEF5928FF00FB08FE92FE86" box="[199,358,331,357]" italics="true" pageId="5" pageNumber="344">Abroma egae</emphasis>
</taxonomicName>
(
<figureCitation id="13702A18FFEF5928FEB9FB08FE38FE87" box="[382,460,331,357]" captionStart="FIGURE 2" captionStartId="4.[152,255,1941,1965]" captionTargetBox="[203,1384,181,1917]" captionTargetId="figure-24@4.[203,1384,181,1917]" captionTargetPageId="4" captionText="FIGURE 2. Tibicen egae Distant, 1892 type specimen (NHMUK010392112) dorsal (top) and ventral (middle) habitus. The inflated aedeagus with membrane and recurved terminal spines are characteristic of Brevialavenosa. ©The Trustees of the Natural History Museum, London. Photos courtesy of Fernley Symons and Mick Webb." figureDoi="http://doi.org/10.5281/zenodo.8166902" httpUri="https://zenodo.org/record/8166902/files/figure.png" pageId="5" pageNumber="344">Fig. 2</figureCitation>
,
<typeStatus id="54F0883FFFEF5928FE26FB08FDE5FE87" box="[481,529,331,357]" pageId="5" pageNumber="344">type</typeStatus>
specimen BMNH, specimen NHMUK010392112) possesses the following combination of structures that place the species in the Cicadettinae
<bibRefCitation id="EFDA4B6CFFEF5928FC46FB2CFBDDFE6B" author="Buckton" box="[897,1065,367,393]" firstAuthor="Buckton" pageId="5" pageNumber="344" pagination="65 - 96" refId="ref12047" refString="Buckton, G. B. (1890) s. n. In: Mongraph of the British Cicadae, or Tettigiidae, illustrated by more than four hundred coloured drawings. Vol. I. Part III. Macmillan and Co., London, pp. xxxiv - xlviii + 65 - 96, pls. xxi - xxx. https: // doi. org / 10.5962 / bhl. title. 9473" type="book chapter" year="1890">Buckton, 1890</bibRefCitation>
: partially fused fore wing cubitus posterior and anal vein 1, division of fore wing vein cubitus anterior 1 with the proximal portion shorter than the distal portion, hindwing cubital cell 1 width two or more times the width of cubital cell 2, hindwing radius posterior and median veins fused at their bases, partially visible dorsal metanotum, male opercula not strongly S-shaped and lacking a deeply concave lateral margin, lack of timbal covers, undeveloped pygofer distal shoulder, pygofer upper lobe is present, and claspers that restrain the aedeagus (
<bibRefCitation id="EFDA4B6CFFEF5928FC94F860FBC8FDDF" author="Marshall" box="[851,1084,547,573]" etAl="et al." firstAuthor="Marshall" pageId="5" pageNumber="344" pagination="1 - 64" refId="ref13722" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B93FEA8FFFEF5928FC07F860FC06FDDF" box="[960,1010,547,573]" italics="true" pageId="5" pageNumber="344">et al</emphasis>
. 2018
</bibRefCitation>
;
<bibRefCitation id="EFDA4B6CFFEF5928FB8EF860FAE2FDDF" author="Simon" box="[1097,1302,547,573]" etAl="et al." firstAuthor="Simon" pageId="5" pageNumber="344" pagination="865 - 886" refId="ref16508" refString="Simon, C., Gordon, E. R., Moulds, M. S., Cole, J., Haji, D., Lemmon, A. R., Lemmon, E. M., Kortyna, M., Nazario, K., Wade, E. J., Meister, R., Goemans, G., Chiswell, S. M., Pessacq, P., Veloso, C., McCutchen, J. P. &amp; Lukasik, P. (2019) Off-target capture data, endosymbiont genes and morphology reveal a relict lineage sister to all other singing cicadas. Biological Journal of the Linnean Society, 128, 865 - 886. https: // doi. org / 10.1093 / biolinnean / blz 120" type="journal article" year="2019">
Simon
<emphasis id="B93FEA8FFFEF5928FB5DF860FB38FDDF" box="[1178,1228,547,573]" italics="true" pageId="5" pageNumber="344">et al</emphasis>
. 2019
</bibRefCitation>
;
<bibRefCitation id="EFDA4B6CFFEF5928FAE4F860FEF2FD83" author="Sanborn" etAl="et al." firstAuthor="Sanborn" pageId="5" pageNumber="344" pagination="1 - 129" refId="ref15413" refString="Sanborn, A. F. (2020 a) The cicadas (Hemiptera: Cicadidae) of Peru including the descriptions of twenty-four new species, three new synonymies, and thirty-seven new records. Zootaxa, 4785 (1), 1 - 129. https: // doi. org / 10.11646 / zootaxa. 4785.1.1" type="journal article" year="2020" yearSuffix="a">
Sanborn
<emphasis id="B93FEA8FFFEF5928FA4FF860FF59FD83" italics="true" pageId="5" pageNumber="344">et al</emphasis>
. 2020a
</bibRefCitation>
). This combination of characters eliminates all remaining subfamilies as an option for the placement of the species.
</paragraph>
<paragraph id="8BF4369DFFEF5928FF00F8CCFEB1FBBB" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="344">
In addition,
<taxonomicName id="4C4B4D1EFFEF5928FE91F8CCFE1BFD4A" baseAuthorityName="Distant" baseAuthorityYear="1892" box="[342,495,655,681]" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B93FEA8FFFEF5928FE91F8CCFE1BFD4A" box="[342,495,655,681]" italics="true" pageId="5" pageNumber="344">Abroma egae</emphasis>
</taxonomicName>
can be classified in the Taphurini based on the lack of timbal covers, the small, narrow opercula that curve towards the midline but remain well separated medially and do not cover the tympanal cavity completely, the undeveloped pygofer distal shoulder, the small upper pygofer lobes, the large, flat basal pygofer lobes, the lack of an uncus, and the presence of claspers. Within the Taphurini, the species is very similar to
<emphasis id="B93FEA8FFFEF5928FF50F95CFE3DFCDB" box="[151,457,799,825]" italics="true" pageId="5" pageNumber="344">Brevisalavenosa auripilosa</emphasis>
<bibRefCitation id="EFDA4B6CFFEF5928FE17F95CFD75FCDB" author="Sanborn" box="[464,641,799,825]" firstAuthor="Sanborn" pageId="5" pageNumber="344" pagination="190 - 198" refId="ref15802" refString="Sanborn, A. F. (2021 b) A new genus and species of Neotropical Taphurini Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae) from Brazil with a note on the taxonomic position of Prosotettix Jacobi, 1907. Proceedings of the Entomological Society of Washington, 123, 190 - 198. https: // doi. org / 10.4289 / 0013 - 8797.123.1.190" type="journal article" year="2021" yearSuffix="b">Sanborn, 2021b</bibRefCitation>
differing in that Distants species lacks dense, golden pile covering the body, the anterior abdomen is darker than the rest of the body instead of being lighter, the piceous mesonotal markings are absent, the spot of infuscation in distal fore wing apical cell 1 is smaller, fore wing radiomedial crossvein is less obliquely angled to the radius posterior vein, the costal margin is thicker to the node, the anterolateral margin of the lateral pronotal collar curves ventrally, the angled lateral opercular margin, the semicircular medial opercular margin, the operculum reaching the posterior tympanal cavity, and especially the large aedeagus with the two lateral, terminal spines but the membrane is ventral instead of terminal in Distants species. As a result,
<taxonomicName id="4C4B4D1EFFEF5928FAC3F9B4FEB4FBD7" authority="(Distant, 1892)" baseAuthorityName="Distant" baseAuthorityYear="1892" class="Insecta" family="Cicadidae" genus="Abroma" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="B93FEA8FFFEF5928FAC3F9B4FA68FBF2" box="[1284,1436,1015,1041]" italics="true" pageId="5" pageNumber="344">Abroma egae</emphasis>
(
<bibRefCitation id="EFDA4B6CFFEF5928FF58FE58FECCFBD7" author="Distant" box="[159,312,1051,1077]" firstAuthor="Distant" pageId="5" pageNumber="344" pagination="54 - 67" refId="ref12244" refString="Distant, W. L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant, 1892</bibRefCitation>
)
</taxonomicName>
is reassigned to the genus
<taxonomicName id="4C4B4D1EFFEF5928FDAFFE58FC39FBD7" authority="Sanborn, 2021 b" authorityName="Sanborn" authorityYear="2021" box="[616,973,1051,1077]" class="Insecta" family="Cicadidae" genus="Brevialavenosa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEA8FFFEF5928FDAFFE58FCE1FBD7" box="[616,789,1051,1077]" italics="true" pageId="5" pageNumber="344">Brevialavenosa</emphasis>
<bibRefCitation id="EFDA4B6CFFEF5928FCDBFE58FC39FBD7" author="Sanborn" box="[796,973,1051,1077]" firstAuthor="Sanborn" pageId="5" pageNumber="344" pagination="190 - 198" refId="ref15802" refString="Sanborn, A. F. (2021 b) A new genus and species of Neotropical Taphurini Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae) from Brazil with a note on the taxonomic position of Prosotettix Jacobi, 1907. Proceedings of the Entomological Society of Washington, 123, 190 - 198. https: // doi. org / 10.4289 / 0013 - 8797.123.1.190" type="journal article" year="2021" yearSuffix="b">Sanborn, 2021b</bibRefCitation>
</taxonomicName>
to become
<taxonomicName id="4C4B4D1EFFEF5928FB8BFE58FF2DFBBB" authority="(Distant, 1892)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Distant" baseAuthorityYear="1892" class="Insecta" family="Cicadidae" genus="Brevialavenosa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="344" phylum="Arthropoda" rank="species" species="egae" status="comb. nov.">
<emphasis id="B93FEA8FFFEF5928FB8BFE58FAC1FBD6" box="[1100,1333,1051,1077]" italics="true" pageId="5" pageNumber="344">Brevialavenosa egae</emphasis>
(
<bibRefCitation id="EFDA4B6CFFEF5928FA84FE58FF24FBBB" author="Distant" firstAuthor="Distant" pageId="5" pageNumber="344" pagination="54 - 67" refId="ref12244" refString="Distant, W. L. (1892) On some undescribed Cicadidae, with synonymical notes. Annals and Magazine of Natural History, Series 6, 10, 54 - 67. https: // doi. org / 10.1080 / 00222939208677373" type="journal article" year="1892">Distant, 1892</bibRefCitation>
)
</taxonomicName>
<emphasis id="B93FEA8FFFEF5928FF27FE7CFEB1FBBB" bold="true" box="[224,325,1087,1113]" pageId="5" pageNumber="344">
<taxonomicNameLabel id="A20C57F4FFEF5928FF27FE7CFEB1FBBB" box="[224,325,1087,1113]" pageId="5" pageNumber="344" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3516516FFEF5928FF00FE21FC19FB9F" box="[199,1005,1122,1149]" pageId="5" pageNumber="344" type="distribution">
<paragraph id="8BF4369DFFEF5928FF00FE21FC19FB9F" blockId="5.[151,1437,151,1149]" box="[199,1005,1122,1149]" pageId="5" pageNumber="344">
<emphasis id="B93FEA8FFFEF5928FF00FE21FE85FB9F" bold="true" box="[199,369,1122,1149]" pageId="5" pageNumber="344">DISTRIBUTION.</emphasis>
The species is known only from
<collectingCountry id="F35C760DFFEF5928FD22FE20FCDDFB9F" box="[741,809,1123,1149]" name="Brazil" pageId="5" pageNumber="344">Brazil</collectingCountry>
(
<bibRefCitation id="EFDA4B6CFFEF5928FCFFFE20FC15FB9F" author="Metcalf" box="[824,993,1123,1149]" firstAuthor="Metcalf" pageId="5" pageNumber="344" pagination="1 - 492" refId="ref13979" refString="Metcalf, Z. P. (1963 c) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 2. Tibicinidae. North Carolina State College Contribution, 1564, i - vi + 1 - 492." type="journal article" year="1963" yearSuffix="c">Metcalf 1963c</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>