447 lines
64 KiB
XML
447 lines
64 KiB
XML
<document id="8C7C968508E9540D55CFA5C8F33D70E1" ID-CLB-Dataset="6632" ID-DOI="10.5281/zenodo.4648807" ID-GBIF-Dataset="7e925546-ee72-463d-a9bf-09ba55ffa2bf" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648807" ID-ZooBank="6B0DBB7A-D677-4655-A3B7-9928010145F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617136465751" checkinUser="carolina" docAuthor="Kovařík, František, Lowe, Graeme & Šťáhlavský, František" docDate="2020" docId="03DA92459574FFDDFEC1F907FBBA6025" docLanguage="en" docName="Euscorpius.321.1-13.pdf" docOrigin="Euscorpius 321" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Reddyanus justi Kovařík & Lowe & Šťáhlavský 2020, sp. n." docType="treatment" docUuid="3FB403D2-AF00-426F-9C4B-109D98750FD0" docUuidSource="ZooBank" docVersion="10" lastPageNumber="10" masterDocId="FFE3EA3D9570FFD6FFF5FF9FFFCA6466" masterDocTitle="Reddyanus justi sp n from Laos (Scorpiones Buthidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1698935501017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5AA69BD423C6EE0CC488E90ED307A231">Reddyanus justi sp n from Laos (Scorpiones Buthidae)</mods:title>
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<mods:namePart id="23D059581818AAB1502F8C1D0C889F68">Lowe, Graeme</mods:namePart>
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<mods:namePart id="9EADC8E9E0D841A4A20793616509E0B8">Šťáhlavský, František</mods:namePart>
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<mods:date id="76CDF6736B2ACC2EB55D1A1E9A02B565">2020</mods:date>
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<taxonomicName id="4C7358D09574FFD2FEC1F907FE3F62D5" ID-CoL="8LYYZ" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[308,501,1688,1715]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">Reddyanus justi</taxonomicName>
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.
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<figureCitation id="13483FD69574FFD2FECEF923FE1362B1" box="[315,473,1724,1751]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–9" captionStart-2="Figures 10–15" captionStart-3="Figures 16–23" captionStart-4="Figures 24–43" captionStart-5="Figures 44–48" captionStart-6="Figures 49–50" captionStart-7="Figure 51" captionStartId-0="3.[113,190,2009,2031]" captionStartId-1="5.[115,192,1977,1999]" captionStartId-2="6.[117,194,1986,2008]" captionStartId-3="7.[116,193,1950,1972]" captionStartId-4="8.[114,191,1742,1764]" captionStartId-5="9.[116,193,1211,1233]" captionStartId-6="9.[113,191,1907,1929]" captionStartId-7="10.[113,181,2009,2031]" captionTargetBox-0="[113,1474,234,1991]" captionTargetBox-1="[127,1461,227,1963]" captionTargetBox-2="[129,1472,251,1940]" captionTargetBox-3="[113,1474,244,1934]" captionTargetBox-4="[137,1450,270,1706]" captionTargetBox-5="[118,1467,290,1170]" captionTargetBox-6="[115,1469,1360,1855]" captionTargetBox-7="[113,1474,234,1990]" captionTargetId-0="figure-32@3.[113,1474,234,1994]" captionTargetId-1="figure-73@5.[113,1474,217,1974]" captionTargetId-2="figure-67@6.[113,1474,234,1958]" captionTargetId-3="figure-66@7.[113,1474,244,1934]" captionTargetId-4="figure-181@8.[115,1476,234,1726]" captionTargetId-5="figure-59@9.[113,1474,266,1175]" captionTargetId-6="figure-60@9.[113,1474,1330,1865]" captionTargetId-7="figure-30@10.[113,1474,234,1992]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="Figures 1–2. Reddyanus justi sp. n., male holotype (1) and male holotype (right) with female paratype (2) in vivo habitus." captionText-1="Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6)." captionText-2="Figures 10–15: Reddyanus justi sp. n., metasoma and telson. Figures 10–12. Holotype male, lateral (10), ventral (11), and dorsal (12) views. Figures 13–15. Paratype female, lateral (13), ventral (14), and dorsal (15) views. Scale bars: 10 mm (10–12, 13–15)." captionText-3="Figures 16–23: Reddyanus justi sp. n. Figures 16–17, 20–23. Male holotype, carapace and tergites I–III (16), coXosternal area and sternites (17), and left legs I–IV, retrolateral aspect (20–23). Figures 18–19. Female paratype, carapace and tergites I–III (18) and coXosternal area and sternites (19)." captionText-4="Figures 24–43: Reddyanus justi sp. n. Figures 24–34. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition. Figures 35–43.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31." captionText-5="Figures 44–48: Reddyanus justi sp. n., left hemispermatophore. Figures 44–45. Capsule and flagellum, conveX (44) and posterior-conveX (45) views. Figures 46–48. Whole hemispermatophore, conveX (46), posterior-conveX (47) and concave (48) views. Scale bars: 500 μm (44–45), 1 mm (46–48)." captionText-6="Figures 49–50. Reddyanus justi sp. n., male holotype. Postpachytene (49) and ideogram (50) (y aXis - % of the diploid chromosome length, dark grey marks chromosomes in multivalent association). Arrowhead indicates a slightly longer pair of chromosomes. Scale bar: 5 μm (49)." captionText-7="Figure 51. Map showing confirmed distribution of Reddyanus spp. in Southeast Asia." figureDoi-0="http://doi.org/10.5281/zenodo.4648810" figureDoi-1="http://doi.org/10.5281/zenodo.4648812" figureDoi-2="http://doi.org/10.5281/zenodo.4648814" figureDoi-3="http://doi.org/10.5281/zenodo.4648816" figureDoi-4="http://doi.org/10.5281/zenodo.4648819" figureDoi-5="http://doi.org/10.5281/zenodo.4648821" figureDoi-6="http://doi.org/10.5281/zenodo.4648823" figureDoi-7="http://doi.org/10.5281/zenodo.4648825" httpUri-0="https://zenodo.org/record/4648810/files/figure.png" httpUri-1="https://zenodo.org/record/4648812/files/figure.png" httpUri-2="https://zenodo.org/record/4648814/files/figure.png" httpUri-3="https://zenodo.org/record/4648816/files/figure.png" httpUri-4="https://zenodo.org/record/4648819/files/figure.png" httpUri-5="https://zenodo.org/record/4648821/files/figure.png" httpUri-6="https://zenodo.org/record/4648823/files/figure.png" httpUri-7="https://zenodo.org/record/4648825/files/figure.png" pageId="4" pageNumber="3">Figures 1–51</figureCitation>
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,
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<tableCitation id="C6F116E89574FFD2FE10F923FDF062B0" box="[485,570,1724,1750]" captionStart="Table 1" captionStartId="4.[115,171,994,1015]" captionTargetId="graphics-481@4.[112,1482,234,972]" captionTargetPageId="4" captionText="Table 1. Comparative measurements of adults of Reddyanus justi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/DF0C73DB9574FFD2FF86FC7DFEB66073" pageId="4" pageNumber="3" tableUuid="DF0C73DB9574FFD2FF86FC7DFEB66073">Table 1</tableCitation>
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)
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http://zoobank.org/urn:lsid:zoobank.org:act:3FB403D2- AF00-426F-9C4B-109
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FD0
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|
||
<typeStatus id="54C89DF19574FFD2FF34F87EFEE7639F" box="[193,301,2017,2041]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
), leg.
|
||
<collectorName id="268646859574FFD2FE98F87DFE3F639A" box="[365,501,2018,2044]" pageId="4" pageNumber="3">M. Černička</collectorName>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="ED62BB969574FFD2FE0AF87EFD88639F" box="[511,578,2017,2041]" pageId="4" pageNumber="3">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36970D89574FFD2FCDAFBA7FBAC60D6" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8BCC23539574FFD2FCDAFBA7FBAC60D6" blockId="4.[815,1472,1079,1200]" pageId="4" pageNumber="3">ETYMOLOGY. The specific epithet is a patronym honoring Pavel Just, an arachnologist who has assisted us greatly with our studies on scorpions in the laboratory as well as in the field during the past several years.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36970D89574FFD2FCDAFB47FB2B6229" pageId="4" pageNumber="3" type="diagnosis">
|
||
<paragraph id="8BCC23539574FFD2FCDAFB47FB2B6229" blockId="4.[815,1473,1239,1616]" pageId="4" pageNumber="3">
|
||
DIAGNOSIS (
|
||
<specimenCount id="9D75E8DA9574FFD2FC47FB47FC036096" box="[946,969,1240,1264]" pageId="4" pageNumber="3" type="male">♂</specimenCount>
|
||
<specimenCount id="9D75E8DA9574FFD2FC3CFB47FC136096" box="[969,985,1240,1264]" pageId="4" pageNumber="3" type="female">♀</specimenCount>
|
||
). Adult length up to
|
||
<quantity id="4C8B8EB69574FFD2FB35FB47FAC66089" box="[1216,1292,1240,1264]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="3" unit="mm" value="33.0">33 mm</quantity>
|
||
(female) and
|
||
<quantity id="4C8B8EB69574FFD2FA53FB47FC926169" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="3" unit="mm" value="47.0">47 mm</quantity>
|
||
(male). Males with more elongate metasomal segments and telson compared to females. Pedipalps and legs yellow to orange, with reduced brown spots. Manus of pedipalps yellow, fingers reddish black. Metasomal segment I with 10 carinae, segments II–IV with 8 carinae. Metasoma V with 5 carinae, which are reduced in males. Posterior terminal tubercle on dorsolateral carinae of metasoma II–III of both seXes very slightly enlarged. Subaculear tooth broad, in females with terminus rounded, in males beveled and irregular quadrilateral in shape, dorsally with 5–7 granules in 3 rows. Pectinal tooth count
|
||
<quantity id="4C8B8EB69574FFD2FC84F9A7FC056229" box="[881,975,1591,1616]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="4" pageNumber="3" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">13–14 in</quantity>
|
||
males,
|
||
<quantity id="4C8B8EB69574FFD2FBD4F9A7FB4A6229" box="[1057,1152,1591,1616]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="4" pageNumber="3" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
|
||
females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36970D89574FFDDFCDAF9E7FBBA6025" lastPageId="11" lastPageNumber="10" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="8BCC23539574FFD2FCDAF9E7FAF56369" blockId="4.[815,1473,1655,2032]" pageId="4" pageNumber="3">
|
||
DESCRIPTION.The adults are
|
||
<quantity id="4C8B8EB69574FFD2FBAEF9E8FB0662E9" box="[1115,1228,1655,1679]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="4" pageNumber="3" unit="mm" value="46.0" valueMax="47.0" valueMin="45.0">45–47 mm</quantity>
|
||
(males) and
|
||
<quantity id="4C8B8EB69574FFD2FAA5F9E7FA0A62E9" box="[1360,1472,1656,1680]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="4" pageNumber="3" unit="mm" value="31.5" valueMax="33.0" valueMin="30.0">30–33 mm</quantity>
|
||
(females) long. Habitus as shown in
|
||
<figureCitation id="13483FD69574FFD2FB33F908FAE562C9" box="[1222,1327,1687,1711]" captionStart="Figures 3–9" captionStartId="5.[115,192,1977,1999]" captionTargetBox="[127,1461,227,1963]" captionTargetId="figure-73@5.[113,1474,217,1974]" captionTargetPageId="5" captionText="Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6)." figureDoi="http://doi.org/10.5281/zenodo.4648812" httpUri="https://zenodo.org/record/4648812/files/figure.png" pageId="4" pageNumber="3">Figs. 3–6</figureCitation>
|
||
. For position and distribution of trichobothria of pedipalps see
|
||
<figureCitation id="13483FD69574FFD2FAB7F928FA0B62B6" box="[1346,1473,1719,1744]" captionStart="Figures 24–43" captionStartId="8.[114,191,1742,1764]" captionTargetBox="[137,1450,270,1706]" captionTargetId="figure-181@8.[115,1476,234,1726]" captionTargetPageId="8" captionText="Figures 24–43: Reddyanus justi sp. n. Figures 24–34. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition. Figures 35–43.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31." figureDoi="http://doi.org/10.5281/zenodo.4648819" httpUri="https://zenodo.org/record/4648819/files/figure.png" pageId="4" pageNumber="3">Figs. 24–28</figureCitation>
|
||
and
|
||
<figureCitation id="13483FD69574FFD2FCABF948FC2A6296" box="[862,992,1751,1776]" captionStart="Figures 24–43" captionStartId="8.[114,191,1742,1764]" captionTargetBox="[137,1450,270,1706]" captionTargetId="figure-181@8.[115,1476,234,1726]" captionTargetPageId="8" captionText="Figures 24–43: Reddyanus justi sp. n. Figures 24–34. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition. Figures 35–43.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31." figureDoi="http://doi.org/10.5281/zenodo.4648819" httpUri="https://zenodo.org/record/4648819/files/figure.png" pageId="4" pageNumber="3">Figs. 30–31</figureCitation>
|
||
. SeXual dimorphism: adult male with longer metasomal segments and longer, narrower telson.
|
||
</paragraph>
|
||
<paragraph id="8BCC23539574FFDEFCDAF888FE836397" blockId="4.[815,1473,1655,2032]" lastBlockId="8.[111,769,1913,2033]" lastPageId="8" lastPageNumber="7" pageId="4" pageNumber="3">
|
||
<emphasis id="B907FF419574FFD2FCDAF888FC606349" bold="true" box="[815,938,1815,1839]" pageId="4" pageNumber="3">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="13483FD69574FFD2FC4DF888FBD76349" box="[952,1053,1815,1839]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–9" captionStartId-0="3.[113,190,2009,2031]" captionStartId-1="5.[115,192,1977,1999]" captionTargetBox-0="[113,1474,234,1991]" captionTargetBox-1="[127,1461,227,1963]" captionTargetId-0="figure-32@3.[113,1474,234,1994]" captionTargetId-1="figure-73@5.[113,1474,217,1974]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–2. Reddyanus justi sp. n., male holotype (1) and male holotype (right) with female paratype (2) in vivo habitus." captionText-1="Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6)." figureDoi-0="http://doi.org/10.5281/zenodo.4648810" figureDoi-1="http://doi.org/10.5281/zenodo.4648812" httpUri-0="https://zenodo.org/record/4648810/files/figure.png" httpUri-1="https://zenodo.org/record/4648812/files/figure.png" pageId="4" pageNumber="3">Figs. 1–6</figureCitation>
|
||
). Base color yellow to orange with dark spots. Chelicerae are yellow, strongly reticulated. Fingers of chelicerae are darkly marked. Carapace and mesosoma patterned with 3 dark longitudinal strips. Ventral side of mesosoma yellow to yellowish brown, with several fuscous markings on sternite VII. Pedipalp femur and patella yellow or orange with small reduced dark maculae which are almost missing on femur. Legs also yellow or orange with black spots. Metasoma is yellowish or reddish brown with black spots, metasoma V darker than metasoma I–IV, telson yellow, red to reddish black.
|
||
</paragraph>
|
||
<caption id="DF0C73DB9575FFD3FF86F826FF136C6F" ID-DOI="http://doi.org/10.5281/zenodo.4648812" ID-Zenodo-Dep="4648812" httpUri="https://zenodo.org/record/4648812/files/figure.png" pageId="5" pageNumber="4" startId="5.[115,192,1977,1999]" targetBox="[127,1461,227,1963]" targetPageId="5">
|
||
<paragraph id="8BCC23539575FFD3FF86F826FF136C6F" blockId="5.[115,1473,1977,2057]" pageId="5" pageNumber="4">
|
||
<emphasis id="B907FF419575FFD3FF86F826FF2363A8" bold="true" box="[115,233,1977,1999]" pageId="5" pageNumber="4">Figures 3–9</emphasis>
|
||
:
|
||
<taxonomicName id="4C7358D09575FFD3FF03F826FE4463A9" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[246,398,1977,1999]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF419575FFD3FF03F826FE4463A9" box="[246,398,1977,1999]" italics="true" pageId="5" pageNumber="4">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A9575FFD3FE61F826FE0363A8" box="[404,457,1977,1998]" pageId="5" pageNumber="4" rank="species">
|
||
<emphasis id="B907FF419575FFD3FE61F826FE6163A8" bold="true" box="[404,427,1977,1998]" pageId="5" pageNumber="4">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF419575FFD3FE43F826FE0963A8" bold="true" box="[438,451,1977,1998]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B907FF419575FFD3FE3AF826FD8C63A8" bold="true" box="[463,582,1977,1999]" pageId="5" pageNumber="4">Figures 3–4</emphasis>
|
||
,
|
||
<emphasis id="B907FF419575FFD3FDA4F826FDBF63A8" bold="true" box="[593,629,1977,1998]" pageId="5" pageNumber="4">8–9</emphasis>
|
||
. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9).
|
||
<emphasis id="B907FF419575FFD3FE4CF849FDFA638D" bold="true" box="[441,560,2006,2028]" pageId="5" pageNumber="4">Figures 5–7</emphasis>
|
||
. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C73DB9576FFD0FF80F85DFB126393" ID-DOI="http://doi.org/10.5281/zenodo.4648814" ID-Zenodo-Dep="4648814" httpUri="https://zenodo.org/record/4648814/files/figure.png" pageId="6" pageNumber="5" startId="6.[117,194,1986,2008]" targetBox="[129,1472,251,1940]" targetPageId="6">
|
||
<paragraph id="8BCC23539576FFD0FF80F85DFB126393" blockId="6.[117,1476,1986,2037]" pageId="6" pageNumber="5">
|
||
<emphasis id="B907FF419576FFD0FF80F85DFECE63B1" bold="true" box="[117,260,1986,2008]" pageId="6" pageNumber="5">Figures 10–15</emphasis>
|
||
:
|
||
<taxonomicName id="4C7358D09576FFD0FEE4F85DFE6363BE" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[273,425,1986,2008]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF419576FFD0FEE4F85DFE6363BE" box="[273,425,1986,2008]" italics="true" pageId="6" pageNumber="5">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A9576FFD0FE45F85CFE2F63BE" box="[432,485,1987,2008]" pageId="6" pageNumber="5" rank="species">
|
||
<emphasis id="B907FF419576FFD0FE45F85CFE0D63BE" bold="true" box="[432,455,1987,2008]" pageId="6" pageNumber="5">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF419576FFD0FE27F85CFE1563BE" bold="true" box="[466,479,1987,2008]" pageId="6" pageNumber="5">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, metasoma and telson.
|
||
<emphasis id="B907FF419576FFD0FD32F85DFC9C63B1" bold="true" box="[711,854,1986,2008]" pageId="6" pageNumber="5">Figures 10–12</emphasis>
|
||
. Holotype male, lateral (10), ventral (11), and dorsal (12) views.
|
||
<emphasis id="B907FF419576FFD0FF80F840FECE6392" bold="true" box="[117,260,2015,2037]" pageId="6" pageNumber="5">Figures 13–15</emphasis>
|
||
. Paratype female, lateral (13), ventral (14), and dorsal (15) views. Scale bars: 10 mm (10–12, 13–15).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C73DB9577FFD1FF81F801FF326388" ID-DOI="http://doi.org/10.5281/zenodo.4648816" ID-Zenodo-Dep="4648816" httpUri="https://zenodo.org/record/4648816/files/figure.png" pageId="7" pageNumber="6" startId="7.[116,193,1950,1972]" targetBox="[113,1474,244,1934]" targetPageId="7">
|
||
<paragraph id="8BCC23539577FFD1FF81F801FF326388" blockId="7.[116,1474,1950,2030]" pageId="7" pageNumber="6">
|
||
<emphasis id="B907FF419577FFD1FF81F801FEC963D5" bold="true" box="[116,259,1950,1972]" pageId="7" pageNumber="6">Figures 16–23</emphasis>
|
||
:
|
||
<taxonomicName id="4C7358D09577FFD1FEE5F800FE6363D2" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[272,425,1951,1972]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF419577FFD1FEE5F800FE6363D2" box="[272,425,1951,1972]" italics="true" pageId="7" pageNumber="6">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A9577FFD1FE45F800FE2C63D2" box="[432,486,1951,1972]" pageId="7" pageNumber="6" rank="species">
|
||
<emphasis id="B907FF419577FFD1FE45F800FE0D63D2" bold="true" box="[432,455,1951,1972]" pageId="7" pageNumber="6">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF419577FFD1FE26F800FE2A63D2" bold="true" box="[467,480,1951,1972]" pageId="7" pageNumber="6">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B907FF419577FFD1FE19F801FDB663D5" bold="true" box="[492,636,1950,1972]" pageId="7" pageNumber="6">Figures 16–17</emphasis>
|
||
,
|
||
<emphasis id="B907FF419577FFD1FD7DF801FD0E63D5" bold="true" box="[648,708,1950,1972]" pageId="7" pageNumber="6">20–23</emphasis>
|
||
. Male holotype, carapace and tergites I–III (16), coXosternal area and sternites (17), and left legs I–IV, retrolateral aspect (20–23).
|
||
<emphasis id="B907FF419577FFD1FD94F824FD3A63B6" bold="true" box="[609,752,1979,2001]" pageId="7" pageNumber="6">Figures 18–19</emphasis>
|
||
. Female paratype, carapace and tergites I–III (18) and coXosternal area and sternites (19).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C73DB9578FFDEFF87F951FE2E6331" ID-DOI="http://doi.org/10.5281/zenodo.4648819" ID-Zenodo-Dep="4648819" httpUri="https://zenodo.org/record/4648819/files/figure.png" pageId="8" pageNumber="7" startId="8.[114,191,1742,1764]" targetBox="[137,1450,270,1706]" targetPageId="8">
|
||
<paragraph id="8BCC23539578FFDEFF87F951FE2E6331" blockId="8.[113,1472,1742,1880]" pageId="8" pageNumber="7">
|
||
<emphasis id="B907FF419578FFDEFF87F951FECA6282" bold="true" box="[114,256,1742,1764]" pageId="8" pageNumber="7">Figures 24–43</emphasis>
|
||
:
|
||
<taxonomicName id="4C7358D09578FFDEFEF9F950FE6F6282" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[268,421,1743,1764]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF419578FFDEFEF9F950FE6F6282" box="[268,421,1743,1764]" italics="true" pageId="8" pageNumber="7">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A9578FFDEFE5EF950FE2A6282" box="[427,480,1743,1764]" pageId="8" pageNumber="7" rank="species">
|
||
<emphasis id="B907FF419578FFDEFE5EF950FE086282" bold="true" box="[427,450,1743,1764]" pageId="8" pageNumber="7">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF419578FFDEFE38F950FE106282" bold="true" box="[461,474,1743,1764]" pageId="8" pageNumber="7">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B907FF419578FFDEFE13F951FDBF6282" bold="true" box="[486,629,1742,1764]" pageId="8" pageNumber="7">Figures 24–34</emphasis>
|
||
. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition.
|
||
<emphasis id="B907FF419578FFDEFE6EF897FDE3637B" bold="true" box="[411,553,1800,1822]" pageId="8" pageNumber="7">Figures 35–43</emphasis>
|
||
.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC23539578FFDDFCDBF8E6FF046645" blockId="8.[814,1471,1913,2033]" lastBlockId="11.[113,778,235,2019]" lastPageId="11" lastPageNumber="10" pageId="8" pageNumber="7">
|
||
<emphasis id="B907FF419578FFDEFCDBF8E6FB9363F7" bold="true" box="[814,1113,1913,1937]" pageId="8" pageNumber="7">Carapace and mesosoma</emphasis>
|
||
(
|
||
<figureCitation id="13483FD69578FFDEFB9BF8E6FB3363F7" box="[1134,1273,1913,1937]" captionStart="Figures 16–23" captionStartId="7.[116,193,1950,1972]" captionTargetBox="[113,1474,244,1934]" captionTargetId="figure-66@7.[113,1474,244,1934]" captionTargetPageId="7" captionText="Figures 16–23: Reddyanus justi sp. n. Figures 16–17, 20–23. Male holotype, carapace and tergites I–III (16), coXosternal area and sternites (17), and left legs I–IV, retrolateral aspect (20–23). Figures 18–19. Female paratype, carapace and tergites I–III (18) and coXosternal area and sternites (19)." figureDoi="http://doi.org/10.5281/zenodo.4648816" httpUri="https://zenodo.org/record/4648816/files/figure.png" pageId="8" pageNumber="7">Figs. 16–19</figureCitation>
|
||
). Entire carapace covered with large granules; carinae absent. Anterior margin of carapace is medially emarginate. Tergites I–VI with 1 median carina, tergite VII pentacarinate. All tegites strongly granulated. Pectinal tooth count
|
||
<quantity id="4C8B8EB6957BFFDDFE14FF74FD8F6565" box="[481,581,235,259]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="11" pageNumber="10" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">13–14 in</quantity>
|
||
males,
|
||
<quantity id="4C8B8EB6957BFFDDFD55FF74FCC96565" box="[672,771,235,259]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="11" pageNumber="10" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
|
||
females. Pectine marginal tips eXtending to posterior end of sternite III in males, to 3/4 of sternite III in females. Pectines with 3 marginal lamellae, 6–7 middle lamellae. Lamellae and fulcra bear numerous pale microsetae. Sternites smooth with 2 parallel furrows, eXcept sternite VII which bears 4 carinae and is granulated. Granulation also present on lateral margins of sternite VI. Posterior medial area of sternite V with a glabrous zone well developed in males, reduced in females.
|
||
</paragraph>
|
||
<caption id="DF0C73DB9579FFDFFF81FB24FECE616D" ID-DOI="http://doi.org/10.5281/zenodo.4648821" ID-Zenodo-Dep="4648821" httpUri="https://zenodo.org/record/4648821/files/figure.png" pageId="9" pageNumber="8" startId="9.[116,193,1211,1233]" targetBox="[118,1467,290,1170]" targetPageId="9">
|
||
<paragraph id="8BCC23539579FFDFFF81FB24FECE616D" blockId="9.[116,1474,1211,1291]" pageId="9" pageNumber="8">
|
||
<emphasis id="B907FF419579FFDFFF81FB24FEC860B7" bold="true" box="[116,258,1211,1233]" pageId="9" pageNumber="8">Figures 44–48</emphasis>
|
||
:
|
||
<taxonomicName id="4C7358D09579FFDFFEFBFB23FE6F60B7" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[270,421,1212,1233]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF419579FFDFFEFBFB23FE6F60B7" box="[270,421,1212,1233]" italics="true" pageId="9" pageNumber="8">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A9579FFDFFE5EFB23FE1560B7" box="[427,479,1212,1233]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B907FF419579FFDFFE5EFB23FE0860B7" bold="true" box="[427,450,1212,1233]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF419579FFDFFE38FB23FE1060B7" bold="true" box="[461,474,1212,1233]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, left hemispermatophore.
|
||
<emphasis id="B907FF419579FFDFFD2FFB24FCA260B7" bold="true" box="[730,872,1211,1233]" pageId="9" pageNumber="8">Figures 44–45</emphasis>
|
||
. Capsule and flagellum, conveX (44) and posterior-conveX (45) views.
|
||
<emphasis id="B907FF419579FFDFFF4CFB47FE82608B" bold="true" box="[185,328,1240,1262]" pageId="9" pageNumber="8">Figures 46–48</emphasis>
|
||
. Whole hemispermatophore, conveX (46), posterior-conveX (47) and concave (48) views. Scale bars: 500 μm (44–45), 1 mm (46–48).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C73DB9579FFDFFF84F8ECFF6B63A5" ID-DOI="http://doi.org/10.5281/zenodo.4648823" ID-Zenodo-Dep="4648823" httpUri="https://zenodo.org/record/4648823/files/figure.png" pageId="9" pageNumber="8" startId="9.[113,191,1907,1929]" targetBox="[115,1469,1360,1855]" targetPageId="9">
|
||
<paragraph id="8BCC23539579FFDFFF84F8ECFF6B63A5" blockId="9.[113,1472,1907,1987]" pageId="9" pageNumber="8">
|
||
<emphasis id="B907FF419579FFDFFF84F8ECFEC863EE" bold="true" box="[113,258,1907,1929]" pageId="9" pageNumber="8">Figures 49–50</emphasis>
|
||
.
|
||
<taxonomicName id="4C7358D09579FFDFFEFBF8ECFE6363EF" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[270,425,1907,1929]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF419579FFDFFEFBF8ECFE6363EF" box="[270,425,1907,1929]" italics="true" pageId="9" pageNumber="8">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A9579FFDFFE5AF8ECFE2E63EF" box="[431,484,1907,1929]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="B907FF419579FFDFFE5AF8ECFE0C63EE" bold="true" box="[431,454,1907,1928]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF419579FFDFFE27F8ECFE1563EE" bold="true" box="[466,479,1907,1928]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male holotype. Postpachytene (49) and ideogram (50) (y aXis - % of the diploid chromosome length, dark grey marks chromosomes in multivalent association). Arrowhead indicates a slightly longer pair of chromosomes. Scale bar: 5 μm (49).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C73DB957AFFDCFF84F846FC636389" ID-DOI="http://doi.org/10.5281/zenodo.4648825" ID-Zenodo-Dep="4648825" box="[113,937,2009,2031]" httpUri="https://zenodo.org/record/4648825/files/figure.png" pageId="10" pageNumber="9" startId="10.[113,181,2009,2031]" targetBox="[113,1474,234,1990]" targetPageId="10">
|
||
<paragraph id="8BCC2353957AFFDCFF84F846FC636389" blockId="10.[113,937,2009,2031]" box="[113,937,2009,2031]" pageId="10" pageNumber="9">
|
||
<emphasis id="B907FF41957AFFDCFF84F846FF196388" bold="true" box="[113,211,2009,2031]" pageId="10" pageNumber="9">Figure 51</emphasis>
|
||
. Map showing confirmed distribution of
|
||
<taxonomicName id="4C7358D0957AFFDCFDABF846FD0D6388" box="[606,711,2009,2030]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907FF41957AFFDCFDABF846FD0D6388" box="[606,711,2009,2030]" italics="true" pageId="10" pageNumber="9">Reddyanus</emphasis>
|
||
</taxonomicName>
|
||
spp. in Southeast Asia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC2353957BFFDDFF84FDB4FF2367A4" blockId="11.[113,778,235,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B907FF41957BFFDDFF84FDB4FEA46625" bold="true" box="[113,366,555,579]" pageId="11" pageNumber="10">Metasoma and telson</emphasis>
|
||
(
|
||
<figureCitation id="13483FD6957BFFDDFE74FDB4FE366624" box="[385,508,554,579]" captionStart-0="Figures 3–9" captionStart-1="Figures 10–15" captionStartId-0="5.[115,192,1977,1999]" captionStartId-1="6.[117,194,1986,2008]" captionTargetBox-0="[127,1461,227,1963]" captionTargetBox-1="[129,1472,251,1940]" captionTargetId-0="figure-73@5.[113,1474,217,1974]" captionTargetId-1="figure-67@6.[113,1474,234,1958]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6)." captionText-1="Figures 10–15: Reddyanus justi sp. n., metasoma and telson. Figures 10–12. Holotype male, lateral (10), ventral (11), and dorsal (12) views. Figures 13–15. Paratype female, lateral (13), ventral (14), and dorsal (15) views. Scale bars: 10 mm (10–12, 13–15)." figureDoi-0="http://doi.org/10.5281/zenodo.4648812" figureDoi-1="http://doi.org/10.5281/zenodo.4648814" httpUri-0="https://zenodo.org/record/4648812/files/figure.png" httpUri-1="https://zenodo.org/record/4648814/files/figure.png" pageId="11" pageNumber="10">Figs. 7–15</figureCitation>
|
||
). Metasomal segment I with 10 carinae, segments II–IV with eight carinae. Metasoma V with 5 carinae which are reduced in males. Vesicle of telson in males with a ventral carina, in females with ventral and lateral carinae. Metasoma with intercarinal surfaces sparsely granulated, with the eXception of metasoma V of males which is smooth. Posterior terminal tubercle on dorsolateral carinae of metasoma II–III of both seXes very slightly enlarged. Telson longer and narrower in males than females. Subaculear tubercle broad in lateral profile, with terminus rounded in females, beveled in males which have a tubercle with irregular quadrilateral shape. Tubercle dorsally with 5–7 granules in three rows.
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFF84FC55FDFA60A5" blockId="11.[113,778,235,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B907FF41957BFFDDFF84FC55FF2A6784" bold="true" box="[113,224,970,994]" pageId="11" pageNumber="10">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="13483FD6957BFFDDFF02FC54FE486785" box="[247,386,971,995]" captionStart="Figures 24–43" captionStartId="8.[114,191,1742,1764]" captionTargetBox="[137,1450,270,1706]" captionTargetId="figure-181@8.[115,1476,234,1726]" captionTargetPageId="8" captionText="Figures 24–43: Reddyanus justi sp. n. Figures 24–34. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition. Figures 35–43.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31." figureDoi="http://doi.org/10.5281/zenodo.4648819" httpUri="https://zenodo.org/record/4648819/files/figure.png" pageId="11" pageNumber="10">Figs. 24–43</figureCitation>
|
||
). Pedipalps very sparsely hirsute. Femur with 5 granulated carinae, patella with 7 granulated carinae. Chela with 4–5 carinae which are granulated in both seXes and more strongly developed in females. Chela sparsely granulated, patella and femur are rather densely granulated eXcept ventrally. Chela movable finger with 6 rows of granules, fiXed finger with 7 rows; fingers bearing 6 eXternal accessory granules, 5–6 internal accessory granules.
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFF84FB54FD236105" blockId="11.[113,778,235,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B907FF41957BFFDDFF84FB54FF6C6085" bold="true" box="[113,166,1227,1251]" pageId="11" pageNumber="10">Legs</emphasis>
|
||
(
|
||
<figureCitation id="13483FD6957BFFDDFF43FB54FEF36085" box="[182,313,1227,1251]" captionStart="Figures 16–23" captionStartId="7.[116,193,1950,1972]" captionTargetBox="[113,1474,244,1934]" captionTargetId="figure-66@7.[113,1474,244,1934]" captionTargetPageId="7" captionText="Figures 16–23: Reddyanus justi sp. n. Figures 16–17, 20–23. Male holotype, carapace and tergites I–III (16), coXosternal area and sternites (17), and left legs I–IV, retrolateral aspect (20–23). Figures 18–19. Female paratype, carapace and tergites I–III (18) and coXosternal area and sternites (19)." figureDoi="http://doi.org/10.5281/zenodo.4648816" httpUri="https://zenodo.org/record/4648816/files/figure.png" pageId="11" pageNumber="10">Figs. 20–23</figureCitation>
|
||
). Femora with 3–4 carinae, patellae with 5 carinae, both granulated. Legs without tibial spurs. Tibia and tarsomeres of legs with macrosetae not arranged serially as bristle combs on dorsal surfaces. Tarsomere II of leg IV with two sparse rows of <20 spiniform setae on ventral surface.
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFF84FAF5FF686204" blockId="11.[113,778,235,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B907FF41957BFFDDFF84FAF5FE9661E4" bold="true" box="[113,348,1386,1410]" pageId="11" pageNumber="10">Hemispermatophore</emphasis>
|
||
(
|
||
<figureCitation id="13483FD6957BFFDDFE8EFAF4FDC561E5" box="[379,527,1387,1411]" captionStart="Figures 44–48" captionStartId="9.[116,193,1211,1233]" captionTargetBox="[118,1467,290,1170]" captionTargetId="figure-59@9.[113,1474,266,1175]" captionTargetPageId="9" captionText="Figures 44–48: Reddyanus justi sp. n., left hemispermatophore. Figures 44–45. Capsule and flagellum, conveX (44) and posterior-conveX (45) views. Figures 46–48. Whole hemispermatophore, conveX (46), posterior-conveX (47) and concave (48) views. Scale bars: 500 μm (44–45), 1 mm (46–48)." figureDoi="http://doi.org/10.5281/zenodo.4648821" httpUri="https://zenodo.org/record/4648821/files/figure.png" pageId="11" pageNumber="10">Figs. 44–48</figureCitation>
|
||
). Flagelliform, with flagellum long, narrow and coiled. Capsule relatively large, ca. half as long as trunk. Sperm hemiduct simple, undivided, long, with one prominent carina; posteriorly fused to base of flagellum. Basal lobe large, triangular in profile with sharp verteX. Trunk short, broad. Measurements: trunk length
|
||
<quantity id="4C8B8EB6957BFFDDFD17F995FF576224" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="10" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, capsule length
|
||
<quantity id="4C8B8EB6957BFFDDFEBEF9B4FE646224" box="[331,430,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="11" pageNumber="10" unit="mm" value="1.12">1.12 mm</quantity>
|
||
, uncoiled flagellum length
|
||
<quantity id="4C8B8EB6957BFFDDFD21F9B4FF546204" metricMagnitude="-3" metricUnit="m" metricValue="3.89" pageId="11" pageNumber="10" unit="mm" value="3.89">3.89 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFF84F9F4FD8963A5" blockId="11.[113,778,235,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B907FF41957BFFDDFF84F9F4FF2062E5" bold="true" box="[113,234,1643,1667]" pageId="11" pageNumber="10">Karyotype</emphasis>
|
||
(
|
||
<figureCitation id="13483FD6957BFFDDFF0CF9F4FEB162E4" box="[249,379,1642,1667]" captionStart="Figures 49–50" captionStartId="9.[113,191,1907,1929]" captionTargetBox="[115,1469,1360,1855]" captionTargetId="figure-60@9.[113,1474,1330,1865]" captionTargetPageId="9" captionText="Figures 49–50. Reddyanus justi sp. n., male holotype. Postpachytene (49) and ideogram (50) (y aXis - % of the diploid chromosome length, dark grey marks chromosomes in multivalent association). Arrowhead indicates a slightly longer pair of chromosomes. Scale bar: 5 μm (49)." figureDoi="http://doi.org/10.5281/zenodo.4648823" httpUri="https://zenodo.org/record/4648823/files/figure.png" pageId="11" pageNumber="10">Figs. 49–50</figureCitation>
|
||
). Diploid number of these specimens was 12 chromosomes. The first pair of chromosomes was slightly longer (each chromosome from this pair forms 10.10 % of the diploid set), whereas the remaining chromosomes gradually decreased in length from 8.68 % to 7.32 % of the diploid set. Only bivalents without chiasmata were observed during male meiosis, and no centromeres on the chromosomes. These characteristics are typical for scorpions in the family
|
||
<taxonomicName id="4C7358D0957BFFDDFF84F8F4FF1B63E5" authorityName="C. L. Koch" authorityYear="1837" box="[113,209,1899,1923]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
(e. g.
|
||
<bibRefCitation id="EFE25EA2957BFFDDFEEFF8F4FE2D63E5" author="MATTOS, V. F." box="[282,487,1899,1923]" pageId="11" pageNumber="10" pagination="121 - 136" refId="ref3675" refString="MATTOS, V. F., D. M. CELLA, L. S. CARVALHO, D. M. CANDIDO & M. C. SCHNEIDER. 2013. High chromosome variability and the presence of multivalent associations in buthid scorpions. Chromosome Research, 21: 121 - 136." type="journal article" year="2013">Mattos et al., 2013</bibRefCitation>
|
||
) and the lower number of chromosomes appears to be typical of the genus
|
||
<taxonomicName id="4C7358D0957BFFDDFD7BF814FCC963C5" box="[654,771,1931,1955]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907FF41957BFFDDFD7BF814FCC963C5" box="[654,771,1931,1955]" italics="true" pageId="11" pageNumber="10">Reddyanus</emphasis>
|
||
</taxonomicName>
|
||
(2n=11–17) (KovařÍk & ŠťÁhlavský, 2019).
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFF84F854FE626385" blockId="11.[113,778,235,2019]" box="[113,424,1995,2019]" pageId="11" pageNumber="10">
|
||
<emphasis id="B907FF41957BFFDDFF84F854FED26385" bold="true" box="[113,280,1995,2019]" pageId="11" pageNumber="10">Measurements</emphasis>
|
||
. See
|
||
<tableCitation id="C6F116E8957BFFDDFEA7F854FE686385" box="[338,418,1995,2019]" captionStart="Table 1" captionStartId="4.[115,171,994,1015]" captionTargetId="graphics-481@4.[112,1482,234,972]" captionTargetPageId="4" captionText="Table 1. Comparative measurements of adults of Reddyanus justi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/DF0C73DB9574FFD2FF86FC7DFEB66073" pageId="11" pageNumber="10" tableUuid="DF0C73DB9574FFD2FF86FC7DFEB66073">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFCC5FF74FBFA66E5" blockId="11.[816,1474,235,645]" pageId="11" pageNumber="10">
|
||
AFFINITIES. The described features distinguish
|
||
<taxonomicName id="4C7358D0957BFFDDFABCFF74FCAA6544" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF41957BFFDDFABCFF74FCAA6544" italics="true" pageId="11" pageNumber="10">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A957BFFDDFC9EFE94FC646544" box="[875,942,267,291]" pageId="11" pageNumber="10" rank="species">
|
||
<emphasis id="B907FF41957BFFDDFC9EFE94FC4F6545" bold="true" box="[875,901,267,291]" pageId="11" pageNumber="10">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF41957BFFDDFC62FE94FC6C6545" bold="true" box="[919,934,267,291]" pageId="11" pageNumber="10">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from all other species of the genus. The new species is characterized by the irregular quadrilateral shape in lateral profile of the subaculear tubercle in males, which is a unique character within the genus (
|
||
<figureCitation id="13483FD6957BFFDDFAC7FEF4FAB965E5" box="[1330,1395,363,387]" captionStart="Figures 3–9" captionStartId="5.[115,192,1977,1999]" captionTargetBox="[127,1461,227,1963]" captionTargetId="figure-73@5.[113,1474,217,1974]" captionTargetPageId="5" captionText="Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6)." figureDoi="http://doi.org/10.5281/zenodo.4648812" httpUri="https://zenodo.org/record/4648812/files/figure.png" pageId="11" pageNumber="10">Fig. 9</figureCitation>
|
||
versus figs.
|
||
<quantity id="4C8B8EB6957BFFDDFC99FE14FC2465C5" box="[876,1006,395,419]" metricMagnitude="0" metricUnit="m" metricValue="5.3721" metricValueMax="5.4356" metricValueMin="5.3086" pageId="11" pageNumber="10" unit="in" value="211.5" valueMax="214.0" valueMin="209.0">209–214 in</quantity>
|
||
KovařÍk & ŠťÁhlavský, 2019: 40). The morphologically closest species is
|
||
<taxonomicName id="4C7358D0957BFFDDFB4BFE34FCBB6585" authority="(KovarIk, 2003)" baseAuthorityName="KovarIk" baseAuthorityYear="2003" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="petrzelkai">
|
||
<emphasis id="B907FF41957BFFDDFB4BFE34FA8765A5" box="[1214,1357,427,451]" italics="true" pageId="11" pageNumber="10">R. petrzelkai</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE25EA2957BFFDDFAABFE34FCA36585" author="KOVARIK" pageId="11" pageNumber="10" pagination="1 - 19" refId="ref3351" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">KovařÍk, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(see also a key in KovařÍk & ŠťÁhlavský, 2019: 43) from
|
||
<collectingCountry id="F36463C3957BFFDDFC98FE74FC006665" box="[877,970,491,515]" name="Vietnam" pageId="11" pageNumber="10">Vietnam</collectingCountry>
|
||
which has a large subaculear tubercle that is rounded (
|
||
<figureCitation id="13483FD6957BFFDDFC68FD94FC2B6645" box="[925,993,523,547]" captionStart="Figures 3–9" captionStartId="5.[115,192,1977,1999]" captionTargetBox="[127,1461,227,1963]" captionTargetId="figure-73@5.[113,1474,217,1974]" captionTargetPageId="5" captionText="Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6)." figureDoi="http://doi.org/10.5281/zenodo.4648812" httpUri="https://zenodo.org/record/4648812/files/figure.png" pageId="11" pageNumber="10">Fig. 9</figureCitation>
|
||
versus fig.
|
||
<quantity id="4C8B8EB6957BFFDDFB9CFD94FB7A6645" box="[1129,1200,523,547]" metricMagnitude="0" metricUnit="m" metricValue="5.334" pageId="11" pageNumber="10" unit="in" value="210.0">210 in</quantity>
|
||
KovařÍk & ŠťÁhlavský, 2019: 40) and stronger dark maculation on its pedipalp segments (
|
||
<figureCitation id="13483FD6957BFFDDFC44FDD4FB8A6605" box="[945,1088,587,611]" captionStart="Figures 24–43" captionStartId="8.[114,191,1742,1764]" captionTargetBox="[137,1450,270,1706]" captionTargetId="figure-181@8.[115,1476,234,1726]" captionTargetPageId="8" captionText="Figures 24–43: Reddyanus justi sp. n. Figures 24–34. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition. Figures 35–43.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31." figureDoi="http://doi.org/10.5281/zenodo.4648819" httpUri="https://zenodo.org/record/4648819/files/figure.png" pageId="11" pageNumber="10">Figs. 24–30</figureCitation>
|
||
versus fig.
|
||
<quantity id="4C8B8EB6957BFFDDFB2AFDD4FAE56605" box="[1247,1327,587,611]" metricMagnitude="0" metricUnit="m" metricValue="5.0546" pageId="11" pageNumber="10" unit="in" value="199.0">199 in</quantity>
|
||
KovařÍk & ŠťÁhlavský, 2019: 39).
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFCC5FD34FC6F6065" blockId="11.[816,1474,683,1027]" pageId="11" pageNumber="10">
|
||
REMARKS. The capsule structures and overall dimensions of the hemispermatophore of
|
||
<taxonomicName id="4C7358D0957BFFDDFBBEFD54FB396684" authority="Kovařík & Lowe & Šťáhlavský, 2020" authorityName="Kovařík & Lowe & Šťáhlavský" authorityYear="2020" box="[1099,1267,715,739]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="justi" status="sp. nov.">
|
||
<emphasis id="B907FF41957BFFDDFBBEFD54FB396684" box="[1099,1267,715,739]" italics="true" pageId="11" pageNumber="10">Reddyanus justi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234423A957BFFDDFB0DFD54FAF96684" box="[1272,1331,715,739]" pageId="11" pageNumber="10" rank="species">
|
||
<emphasis id="B907FF41957BFFDDFB0DFD54FADB6685" bold="true" box="[1272,1297,715,739]" pageId="11" pageNumber="10">sp</emphasis>
|
||
.
|
||
<emphasis id="B907FF41957BFFDDFAE8FD54FAE66685" bold="true" box="[1309,1324,715,739]" pageId="11" pageNumber="10">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
are similar to those recorded for other species of the genus (i. e.,
|
||
<taxonomicName id="4C7358D0957BFFDDFAB2FD74FC1D6745" authority="(Karsch, 1879)" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="basilicus">
|
||
<emphasis id="B907FF41957BFFDDFAB2FD74FA9D6764" box="[1351,1367,747,770]" italics="true" pageId="11" pageNumber="10">R</emphasis>
|
||
.
|
||
<emphasis id="B907FF41957BFFDDFA96FD74FA086765" box="[1379,1474,747,771]" italics="true" pageId="11" pageNumber="10">basilicus</emphasis>
|
||
(Karsch, 1879)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7358D0957BFFDDFC13FC94FAA46745" authority="KovarIk et al., 2016" authorityName="KovarIk" authorityYear="2016" box="[998,1390,779,805]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ceylonensis">
|
||
<emphasis id="B907FF41957BFFDDFC13FC94FC3C6744" box="[998,1014,779,802]" italics="true" pageId="11" pageNumber="10">R</emphasis>
|
||
.
|
||
<emphasis id="B907FF41957BFFDDFBFDFC94FB496745" box="[1032,1155,779,803]" italics="true" pageId="11" pageNumber="10">ceylonensis</emphasis>
|
||
KovařÍk et al., 2016
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7358D0957BFFDDFA5EFC94FBD66725" authority="(Vachon, 1982)" baseAuthorityName="Vachon" baseAuthorityYear="1982" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B907FF41957BFFDDFA5EFC94FA716744" box="[1451,1467,779,802]" italics="true" pageId="11" pageNumber="10">R</emphasis>
|
||
.
|
||
<emphasis id="B907FF41957BFFDDFCC5FCB4FCA76725" box="[816,877,811,835]" italics="true" pageId="11" pageNumber="10">loebli</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE25EA2957BFFDDFC8EFCB4FBDE6725" author="VACHON" box="[891,1044,811,835]" pageId="11" pageNumber="10" pagination="77 - 114" refId="ref4061" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." type="journal article" year="1982">Vachon, 1982</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; cf. KovařÍk et al, 2016). These species also eXhibit a simple, monocarinate sperm hemiduct, large triangular basal lobe and long, coiled flagellum. This provides additional support for our hypothesis that these features may be diagnostic for the genus (KovařÍk et al, 2016), although the hemispermatophores of other
|
||
<taxonomicName id="4C7358D0957BFFDDFB83FC54FB216785" box="[1142,1259,971,995]" class="Arachnida" family="Buthidae" genus="Reddyanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907FF41957BFFDDFB83FC54FB216785" box="[1142,1259,971,995]" italics="true" pageId="11" pageNumber="10">Reddyanus</emphasis>
|
||
</taxonomicName>
|
||
species have yet to be studied.
|
||
</paragraph>
|
||
<paragraph id="8BCC2353957BFFDDFCC5FBB4FBBA6025" blockId="11.[816,1136,1067,1091]" box="[816,1136,1067,1091]" pageId="11" pageNumber="10">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F36463C3957BFFDDFC24FBB4FBCF6024" box="[977,1029,1067,1090]" name="Laos" pageId="11" pageNumber="10">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="13483FD6957BFFDDFBE1FBB4FBA96025" box="[1044,1123,1067,1091]" captionStart="Figure 51" captionStartId="10.[113,181,2009,2031]" captionTargetBox="[113,1474,234,1990]" captionTargetId="figure-30@10.[113,1474,234,1992]" captionTargetPageId="10" captionText="Figure 51. Map showing confirmed distribution of Reddyanus spp. in Southeast Asia." figureDoi="http://doi.org/10.5281/zenodo.4648825" httpUri="https://zenodo.org/record/4648825/files/figure.png" pageId="11" pageNumber="10">Fig. 51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |